Phuket mar. biol. Cent. Res. Bull. 70: 1–6 (2011)
ON THE IDENTITY OF THE MANGROVE CRAB, PARACLEISTOSTOMA ERIOPHORUM
NOBILI, 1903 (CRUSTACEA: BRACHYURA: CAMPTANDRIIDAE)
Peter K. L. Ng 1, 2, Cheryl G. S. Tan 2 and Rueangrit Promdam 3
1
Tropical Marine Science Institute and Raffles Museum of Biodiversity Research, National University of
Singapore, 14, Science Drive 4, Singapore 117543, Republic of Singapore.
2
Department of Biological Sciences, National University of Singapore, 14, Science Drive 4, Singapore
119260, Republic of Singapore.
3
Reference Collection, Phuket Marine Biological Center, Phuket Marine Biological Center, P.O. Box 60,
Phuket, 83000, Thailand. (r_promdam@yahoo.com)
Corresponding author: P. K. L. Ng, e-mail: dbsngkl@nus.edu.sg
ABSTRACT: The identity of the poorly known camptandriid mangrove crab Paracleistostoma
eriophorum Nobili, 1903, is clarified. It is shown to be a senior synonym of Paracleistostoma
tweediei Tan & Humpherys, 1995, and the taxonomy of the species is discussed, with the range
of the species extended to Thailand. Notes on its ecology are also provided.
INTRODUCTION
The camptandriid genus Paracleistostoma
De Man, 1895, is currently represented by eight
species (Rahayu & Ng, 2003; Ng et al., 2008).
Ng et al. (2008: 233, 234) commented that the
poorly known Paracleistostoma eriophorum
Nobili, 1903, was actually a senior synonym of
Paracleistostoma tweediei Tan & Humpherys,
1995, but did not elaborate.
We here redescribe and figure the type of
Paracleistostoma eriophorum Nobili, 1903, and
comment on its taxonomic relationships.
MATERIALS AND METHODS
The terminology follows that in Tan &
Ng (1999). Measurements provided, in millimetres,
are of the carapace length and width respectively.
Specimens are deposited in the Museo Regionale
di Scienze Naturali (MZUT), Sezione di Zoologia,
Turin, Italy; Zoological Reference Collection (ZRC)
of the Raffles Museum of Biodiversity Research,
National University of Singapore, Singapore;
Phuket Marine Biological Centre (PMBC), Phuket,
Thailand; and Queensland Museum (QM),
Brisbane, Australia.
Paracleistostoma eriophorum Nobili, 1903
(Figs. 1, 2)
Paracleistostoma eriophorum Nobili, 1903: 23.
- Manning & Holthuis, 1981: 209 (list) - Ng
et al., 2008: 233, 234.
Paracleistostoma wardi - Yang, 1979: 39 (list).
- Harminto, 1988: 88 (nec P. wardi Rathbun,
1926). - Tan & Ng, 1994: 83 (list).
Paracleistostoma tweediei Tan & Humpherys,
1995: 251, figs. 1–3.
Paracleistostoma tweediei Tan & Ng, 1995: 608.
Material examined. - Holotype male of P.
eriophorum (7.5 x 10.0 mm) (MZUT Cr1200, ex
1531), Buntal, Sarawak, East Malaysia, don. R.
Shelford, 1902. Others - SINGAPORE: 1 male
(holotype of P. tweediei) (7.9 x 10.2 mm) (ZRC
1987.57), 8 males (paratypes of P. tweediei) (5.3
x 6.7 mm, 6.4 x 8.3 mm, 6.6 x 8.4 mm (ZRC
1987.60–62), 7.4 x 9.4 mm, 7.5 x 9.7 mm, 7.5 x
10.0 mm, 8.3 x 10.7 mm, 8.8 x 11.5 mm (ZRC
1987.57–59), Lim Chu Kang mangrove, coll. P.
K. L. Ng & S. Harminto, 13 September 1986; 1
male (7.2 x 9.3 mm) (ZRC 1987.1382),
Woodlands, Singapore, coll. S, Harminto, 2 June
1987. PENINSULAR MALAYSIA: 18 males (ZRC
1998.1007), Matang mangroves, Perak, coll. C.
G. S. Tan, 8 June 1995; 4 males (5.0–6.9 x 6.1–
8.8 mm) (ZRC 1998.998), Matang mangroves,
Perak, coll. A. Ramli, M. Faizal, C. G. S. Tan, 10
June 1995. BRUNEI: 1 male (9.3 x 11.9 mm) (QM
W19967), Batu Marang, coll. N. A. Mackenzie,
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Phuket mar. biol. Cent. Res. Bull.
Figure 1. Paracleistostoma eriophorum Nobili, 1903. A, holotype male (7.5 x 10.0 mm) (MZUT
Cr1200, ex 1531), Sarawak; B, C, male (7.6 x 9.6 mm) (ZRC), Thailand. A, B, overall views; C, left
G1 in situ.
March 1993. THAILAND: 1 male (7.6 x 9.6 mm),
1 female (6.5 x 8.3 mm) (ZRC 2010.0071), 1 male
(6.9 x 8.5 mm) (ZRC 2010.0072); 3 males (10.9
x 8.3 mm, 10.4 x 8.0 mm, 10.0 x 7.7 mm), 3
females (12.0 x 9.0 mm, 9.0 x 7.0 mm, 8.8 x 6.8
mm) (PMBC 23738), mangroves, Khanom District,
Nakhon Srithammarat, Gulf of Thailand, coll. R.
Promdam & A, Kaenphet, 23 June 2007.
Type locality. - Buntal, Sarawak.
Diagnosis (holotype male of P. eriophorum). Carapace surface smooth, slightly convex, broadly
quadrangular, broader than long (1.2–1.3 times);
regions indistinct, epigastric lobes moderately low;
metagastric groove shallow. Front almost straight
from dorsal view, upper surface slightly concave
3
Identity of the mangrove crab
Figure 2. Paracleistostoma eriophorum Nobili, 1903. A, G–I, holotype male (7.5 x 10.0 mm) (MZUT
Cr1200, ex 1531), Sarawak; B–F, male (7.9 x 10.2 mm) (ZRC 1987.57), holotype of P. tweediei Tan
& Humpherys, 1995, Singapore. A, B, male abdomen; C, frontal view showing epistome; D, anterior
part of male thoracic sternum; E, outer view of left chela; F, G, dorsal view of right carpus of cheliped;
H, I, left G1 (setae not drawn). Scales: A, G = 0.5 mm; H, I = 1.0 mm.
medially, anterolateral angles of front rounded;
frontomedial projection triangular, apex rounded;
orbital margins setose; supraorbital margin weakly
granular, sinuous; infraorbital margin straight, finely
granular along central third, inner infraorbital tooth
well developed, distinct from rest of infraorbital
margin; suborbital ridge defined by line of granules;
posterolateral border and surfaces of carapace with
dense setae; posterolateral margin lined with small
granules, completely obscured by pubescence;
posterior carapace margin straight. Anterior and
posterior halves of epistome separated by median
transverse ridge; postero medial-tooth broadly
triangular, ending in small, pointed extremity,
posterior margin on either side almost straight;
sulcus separating epistome from anterior
4
Phuket mar. biol. Cent. Res. Bull.
pterygostomian margin indistinct. Third maxilliped
ischium broadly rectangular, outer distal margin
with row of long setae along outer half; merus
with well developed outer distal angle which is
bilobed, outer surface uneven, inverted V shaped
sulci on outer surface indistinct. Chelipeds
subequal, enlarged; carpus subglobular, outer
surface almost smooth, inner surface sparsely
granular, inner dorsal edge finely beaded with
granules, appearing subcarinate, inner ventral edge
with large triangular granules, with short row of
long setae. Ambulatory legs relatively long, densely
setose; first and last pairs of ambulatory legs
almost equal in length, first pair with setae only on
dorsal surface of merus. Abdominal somites 2–5
completely fused, sutures not discernible, suture
between somites 2 and 3 visible only as short,
shallow grooves on either side of abdomen; somite
1 broader than somite 2, with transverse mdian
ridge; somite 6 shorter than telson, lateral margin
sinuous. G1 recurved, apex large, globular, with
short but distinct submedian spine-like distal
appendage.
Remarks. - Nobili’s (1903) description of
Paracleistostoma eriophorum was relatively
detailed but he did not provide a figure. He
distinguished P. eriophorum from P. depressum and
P. cristatum by the unusually tomentose
ambulatory legs and the presence of two carinae
of long granules on the inferior and superior
margins of the outer surface of the cheliped palm.
Nobili (1903) also mentioned that the posterior
carapace, including the median branchial region
and the area around the intestinal region, bears a
thick, short felt like tomentum, and the ambulatory
legs were covered with long, woolly setae with
the dactyli bearing sulci and setae. From his
comparisons, Serène (1974:64) noted “... it is
possible that eriophorum is a Leipocten or Baruna”.
After examining the type, there is no doubt
that P. eriophorum is a species of Paracleistostoma.
As noted by Ng et al. (2008), P. eriophorum Nobili,
1903, is a senior synonym of P. tweediei Tan &
Humpherys (1995). The type specimens of both
species are of similar size, and agree closely in all
characters. The holotype male of P. eriophorum
is not in a good condition (Fig. 1A) with most of
the appendages detached. Most of the setae on
the carapace have also been lost, although one can
discern where the main setose areas were (Fig.
1A). The distal part of the G1 of the holotype male
is somewhat less swollen (Fig. 2H, I) compared
to the recent material (Fig. 1C), but this is almost
certainly because of its age and some dehydration.
Paracleistostoma eriophorum appears to
be close to P. depressum and is distinguished from
all its congeners by the unique structure of the
G1, which has the distal appendage modified to
form a spine like process. It is one of the two
Paracleistostoma species having the inner ventral
edge of the carpus lined with sharp granules, the
only other Paracleistostoma species exhibiting this
condition being P. longimanum. Also, in P.
eriophorum, the ridge below the infraorbital margin
is formed by a raised line of granules. This
condition is also found in P. wardi.
Neither Nobili (1903) nor Tan &
Humpherys (1991) described the epistome and the
anterior sternal structure, and these are included
in the present description (Fig. 2C, D). Only one
female specimen of this species (from Thailand)
is known. It closely resembles the male in all nonsexual features except that the chelae are even
more slender, and the patches of setae on the
posterolateral regions are less clearly demarcated,
partially because there are also short setae on the
dorsal parts of the carapace which are normally
glabrous in males. The female abdomen is
subcircular, covers the thoracic sternum, and all
the somites and telson are mobile.
Harminto (1988) recorded this species as
P. wardi (Rathbun, 1926). His specimens were
examined and compared with Australian specimens
of P. wardi in the ZRC and were found to differ
significantly from the latter species in terms of
the structures of the G1, epistome and male
cheliped carpus. Tan & Ng (1994) had incorrectly
recorded this species as P. wardi in their list of
mangrove Brachyura.
The figure of the frontal region and
abdomen provided by Tan & Humpherys (1995)
is somewhat inaccurate in that the inner infraorbital
tooth is not clearly shown (Tan & Humpherys,
1995: Fig. 2A) and the sutures between abdominal
segments 3–5 are shown to be present as notches
5
Identity of the mangrove crab
on either side of the abdomen (Tan & Humpherys,
1995: Fig. 2G). In fact, only the suture between
somites 3 and 4 is visible as a notch on either side
of the abdomen. The above features are refigured
here (Fig. 2A, B).
As discussed by Ng et al. (2008),
there is a nomenclatural problem with the names
Paracleistostoma tweediei Tan & Humpherys,
1995, and Paracleistostoma tweediei Tan & Ng,
1995. Ng et al. (2008: 234) noted that “Both are
objective synonyms as they are based on the same
holotype. The problem arose because C. G. S. Tan
and P. K. L. Ng originally intended to publish the
new species, and the paper was prepared for a
regional symposium in 1994 (Tan & Ng, 1995).
At the same time, P. J. F. Davie and A. Humpherys
had independently also discovered the species. As
it was clear that the two taxa in question were
conspecific, P. J. F. Davie and P. K. L. Ng left it
to C. G. S. Tan and A. Humpherys to finish the
new species description on their own (Tan &
Humpherys, 1995). In early 1995, P. K. L. Ng
asked the editors of the symposium volume to
have the paragraph on the new Paracleistostoma
in their paper deleted, but although they agreed,
the changes were not made, no proofs were sent,
and the volume was published in 1995 with the
problem paragraph still intact. This mistake
nevertheless validates P. tweediei C. G. S. Tan &
Ng, 1995. In any case, the symposium article
came out later than the paper in the Raffles Bulletin
of Zoology. Since the symposium volume did not
have a publication date, under the Code, it should
be regarded as published on 31 December 2005.
This gives P. tweediei C. G. S. Tan & Humpherys,
1995, priority over P. tweediei C. G. S. Tan &
Ng, 1995.”
Ecology. - In Perak, Malaysia, Paracleistostoma
eriophorum was collected from the Matang
mangroves along with specimens of P. depressum
and the varunid Metaplax elegans De Man, 1888.
These were all found in burrows in soft, brown
mud, close to the edge of the river. In the Khanom
District, Thailand, P. eriophorum were found in
mangrove habitats near rivers, with burrows
typically occurring in the areas between nearby
mangrove trees. Sympatric species there, include
the sesarmids Episesarma palawanense (Rathbun,
1914), Parasesarma ungulatum (H. Milne
Edwards, 1853), Perisesarma eumolpe (De Man,
1895) and P. indiarum (Tweedie, 1940).
Distribution. - Buntal, Sarawak, East Malaysia;
Brunei, Peninsular Malaysia, Singapore, and now
for the first time from the eastern coast of Thailand.
ACKNOWLEDGEMENTS
The authors thank Giovanni Balma of the
University of Turin for his help in obtaining the
type of P. eriophorum. Part of the study was
supported by research grants to the first author
from the National University of Singapore. The
third author was supported by the TOTAL
Corporate Foundation, TOTAL E&P Thailand and
the TRF/BIOTEC Special Program for Biodiversity
Research and Training Grant BRT R_149023 and
he would like to thank Arwut Kaenphet, staff of
Walailak University, who worked closely with him
during the field trips to the mangroves of Khanom.
REFERENCES
Harminto, S. 1988. Systematics of the family Ocypodidae Rafinesque, 1815, s. lat. (Crustacea: Decapoda:
Brachyura) of Southeast Asia. Unpublished M.Sc. thesis, Department of Zoology, National
University of Singapore.
Manning, R.B. & L.B. Holthuis. 1981. West African Brachyuran Crabs (Crustacea: Decapoda). Smithson.
Contr. Zool. 306: 1–379.
Ng, P.K.L., D. Guinot & P.J.F. Davie. 2008. Systema Brachyurorum: Part I. An annotated checklist of
extant brachyuran crabs of the world. Raffles Bull. Zool., Suppl. 17: 1–286.
Nobili, G. 1903. Contributo alla fauna carcinologica di Borneo. Boll. Mus. Zool. Anat. comp. Torino.
18(447): 1–32, 3 figs.
6
Phuket mar. biol. Cent. Res. Bull.
Rahayu, D.L. & P.K.L. Ng, 2003. The Camptandriidae of Irian Jaya, Indonesia, including the descriptions
of two new species (Decapoda: Brachyura). J. Crust. Biol. 23(4): 951–962.
Rathbun, M.J. 1926. Brachyuran crabs from Australia and New Guinea. Rec. Aust. Mus. 15(2): 177–
182, pls. 14–16.
Serène, R. 1974. Note on the genera and species of the Camptandriinae Stimpson, 1858 (Decapoda,
Brachyura: Ocypodidae). Treubia. 28(3): 59–117.
Tan, C.G.S. & A. Humpherys. 1995. Paracleistostoma tweediei, a new species of camptandriine crab,
from Singapore and Brunei mangroves (Crustacea: Decapoda: Brachyura: Ocypodidae). Raffles
Bull. Zool. 43(1): 251–256.
Tan, C.G.S. & P.K.L. Ng. 1995. The Camptandriinae of Singapore and Malaysia with description of
one new species of Paracleistostoma De Man, 1895 (Crustacea: Decapoda: Brachyura:
Ocypodidae). Proc 3rd ASEAN-Australia Symp. Living Coastal Resources Vol. 2: research
papers. C. R. Wilkinson, S. Sudara & L. M. Chou (Eds.), Bangkok, pp. 607–611.
Tan, C.G.S. & P.K.L. Ng. 1994. An annotated checklist of mangrove brachyuran crabs from Malaysia
and Singapore. Hydrobiologia. 285: 75–84.
Tan, C.G.S. & P.K.L. Ng. 1999. A revision of the genus Camptandrium Stimpson, 1858 (Crustacea:
Decapoda: Brachyura: Camptandriidae). Raffles Bull. Zool. 47(1): 193–219.
Yang, C. M. 1979. A list of Brachyura in the Zoological Reference Collection of the Department of
Zoology. Unpublished checklist, Department of Zoology, University of Singapore, 60 pp.
(mimeographed).
Manuscript received: 20 October 2009
Accepted: 9 April 2010
Phuket mar. biol. Cent. Res. Bull. 70: 7–14 (2011)
NEW RECORDS OF SPIDER CRABS OF THE GENERA CYRTOMAIA MIERS, 1886,
AND PLATYMAIA MIERS, 1886 (DECAPODA: MAJOIDEA: INACHIDAE)
FROM THE ANDAMAN SEA, THAILAND
Rueangrit Promdam
Department of Biology, Faculty of Science, Prince of Songkla University, Hatyai, Songkhla 90112,
Thailand
Email: r_promdam@yahoo.com
ABSTRACT Two species of deep-water spider crabs, Cyrtomaia suhmii Miers, 1886, and
Platymaia alcocki Miers, 1886, based on the materials of the BIOSHELF Project (1996–2000)
are recorded for the first time from Thailand. It is also the first record of these two genera from
the Andaman Sea coast of Thailand. Adult and juvenile forms of C. suhmii collected from the
same locality confirm that C. curviceros Bouvier, 1915, is a junior synonym, the characters
previously used to separate them explained by size and age.
Key words: new records, spider crabs, Decapoda, Majoidea, Inachidae, Cyrtomaia, Platymaia,
Thailand
INTRODUCTION
TAXONOMY
Two rare spider crabs, Cyrtomaia suhmii
Miers, 1886, and Platymaia alcocki Miers, 1886,
were collected by the R.V. Chakratong Tongyai
from the Andaman Sea during the BIOSHELF
project surveys of the west coast of Thailand in
1996–2000. Based on Ng and Davie (2002), and
Naiyanetr (2007), the species as well as the genera
had not been previously reported from Thai
waters. The present report serves to formally
record these species, as well as to discuss variation
in adult and juvenile specimens.
Family Inachidae MacLeay, 1838
Genus Cyrtomaia Miers, 1886
Cyrtomaia suhmii Miers, 1886
(Figs. 1, 2A–F)
MATERIALS AND METHODS
Specimens examined are deposited in the
Reference Collection of Phuket Marine Biological
Center (PMBC). All measurements provided are
pre-rostral lengths and carapace widths
respectively. For details on all BIOSHELF stations,
see Aungtonya et al. (2000). The classification
used here follows Ng et al. (2008).
Cyrtomaia suhmii Miers, 1886: 16, pl. 3, fig. 2,
2a–2c; Rathbun, 1918: 6; Richer de
Forges and Guinot, 1990: 525; Richer de
Forges and Ng, 2007: 56 (list); Ng et al.,
2008: 111 (list).
Cyrtomaia suhmi - Rathbun, 1893: 230; Griffin,
1974: 9; Guinot and Richer de Forges,
1982a: 1096; 1982b: 21, fig. 10, 11A–B,
23B; Griffin and Tranter, 1986b: 352, figs.
1, 2; Richer de Forges and Guinot, 1988:
42–43 fig. 2B, C, pl. 2F–G; Poore, 2004:
360; 361 fig. 108.
Cyrtomaia Suhmi var. curviceros Bouvier, 1915b:
9–15, pls. 1.
Cyrtomaja suhmi typica Serène and Lohavanijya,
1973: 46 (in key).
8
Phuket mar. biol. Cent. Res. Bull.
Figure 1. Cyrtomaia suhmii, adult male (60.2 × 67.1 mm) (PMBC 19999), dorsal view of whole
animal. Scale = 20 mm.
Cyrtomaja suhmi curvicornis Serène and
Lohavanijya, 1973: 45, 46 (in key).
Cyrtomaia curviceros Sakai, 1976: 181; Guinot and
Richer de Forges, 1982a: 1096; 1982b:
24, fig. 12A–D; Richer de Forges and
Guinot, 1988: 42–43, fig. 2A; Ng et al.,
2001: 13; 81 fig. 3h; Richer de Forges
and Ng, 2007: 56 (list).
not Cyrtomaia suhmi - Griffin and Brown. 1976:
252, fig. 6; Griffin and Tranter, 1986a:
30, fig. 91g [= C. griffini Richer de
Forges and Guinot, 1990].
not Cyrtomaia suhmi typica Doflein, 1904 : 54–
55, pl. 19: figs. 1, 2 [= C. gaillardi Guinot
and Richer de Forges, 1982].
Material examined
PMBC 19999, 1 male (60.2 × 67.1 mm),
BIOSHELF St. E8, 0832´N, 09604´E, Otter trawl,
488 m, coll. S. Bussarawit and C. Aungtonya,
06.02.1999; PMBC 20000, 2 juv. (11.8 × 11.5 mm,
8.6 × 8.2 mm), BIOSHELF St. J10, 0715´N,
09715´E, Agassiz trawl, 695–683 m, coll. C.
Aungtonya & V. Vongpanich, 19.02.2000; PMBC
19929, 1 juv. (7.7 × 7.4 mm), 1 female (56.3 ×
61.2 mm), BIOSHELF St. L10, 0645´N, 09723´E,
Agassiz trawl, 707–651 m, coll. C. Aungtonya &
V. Vongpanich, 21.02.2000
Diagnosis
Carapace smooth, not granular or
spinulate, with 2 long, subparallel protogastric
spines. Pseudorostral spine divergent, arched.
Basal antennal article cylindrical, armed with 3–5
spines (include internal spine). Orbital border
smooth, intercalated orbital granule present (in
young) or absent, without preocular spines (in
adults). Ocular spine blunt.
9
New records on spider crabs
Figure 2. Cyrtomaia suhmii: A–C, juvenile (11.8 × 11.5 mm) (PMBC 20000); D–F, adult male (60.2 ×
67.1 mm) (PMBC 19999); A, D, dorsal view; B, E, anterior part of carapace showing rostrum; C, F,
lateral view of carapace. Scale = 10 mm.
Distribution
Indo-West Pacific: Japan, Philippines,
Indonesia, Australia, Thailand (Andaman Sea) and
India.
Remarks
Cyrtomaia suhmii was described by Miers
(1886) on the basis of an imperfect crushed young
male specimen collected from Talaud Islands (Tulur
10
Phuket mar. biol. Cent. Res. Bull.
Islands) (see Guinot & Richer de Forges 1982b:
22, fig. 10), Indonesia, by the Challenger
Expedition. Subsequently, Cyrtomaia suhmii
curviceros Bouvier, 1915, was described from Japan
and recognized as a distinct species by Sakai
(1976). The status of C. curviceros has long been
uncertain. Griffin and Tranter (1986) treated both
names as synonyms, while Richer de Forges and
Guinot (1982a, 1988) argued that it may be a good
species. Ng and Huang (1997) treated Taiwanese
material of the species as C. curviceros (see also
Ng et al., 2001), and until recently, was still
regarded as a distinct species (see Richer de Forges
and Ng, 2007). In their annotated checklist of extant
brachyuran crabs of the world, Ng et al. (2008)
treated C. suhmii curviceros as a synonym of C.
suhmii but without comments. Examination of
more material of other allied species like C. griffini
Richer de Forges & Ng, 1990, has shown that the
characters used to separate C. curviceros from C.
suhmii are not valid at the species level (P. K. L.
Ng, pers. comm., see also Richer de Forges and
Ng, 2008).
Specimens of various sizes were collected
from the same Thai locality. The young specimens
are similar to the original description and figure of
C. suhmii by Guinot and Richer de Forges (1982b:
22, 23, fig. 10, 11A, B), especially with regards to
the protogastric spines which are divergent (Miers,
1886: 16, pl. 3, fig. 2). On the other hand, large
male specimens resemble C. curviceros (Fig. 1,
2D–F), although the number of the spines on the
basal antennal article is slightly different. This
difference, however, is not significant and can
easily be explained by variation. The present series
of specimens confirm that the differences which
have been observed in the past between these two
species are due to size and age, and that C. suhnii
curviceros Bouvier, 1915, clearly is a junior
synonym of C. suhmii Miers, 1886.
Genus Platymaia Miers, 1886
Platymaia alcocki Rathbun, 1916
(Figs. 3, 4A–F)
Platymaia wyville-thomsoni. - Wood-Mason and
Alcock, 1891: 258; Alcock, 1895: 181–
182; Doflein, 1904: 59–69, 160–165, figs.
2–6; pls. 2, 20–23, 39, 43: figs. 5, 6; pl.
Figure 3. Platymaia alcocki, adult male (71.0 × 77.5 mm) (PMBC 20367), dorsal view of whole
animal. Scale = 20 mm.
11
New records on spider crabs
Figure 4. Platymaia alcocki: A–C, adult male (71.0 × 77.5 mm); D–F, young male (25.9 × 30.4 mm)
(PMBC 20367); A, D, dorsal view; B, E, anterior part of carapace showing rostrum; C, F, lateral view
of carapace. Scale = 10 mm.
50: figs. 2, 5, 6 [not Platymaia wyvillethomsoni Miers, 1886].
Platymaia alcocki Rathbun, 1916: 530; 1918: 8;
Serène and Lohavanijya, 1973: 48 (in key);
Griffin, 1974: 27; Griffin and Tranter,
1986a: 48 (Pl. 4b; figs. 10a, b; 11c);
1986b: 357; Takeda and Webber, 2006:
194.
Not Platymaia alcocki - Takeda and Miyake, 1969:
498–500, figs. 10a, b, 11a–c; Sakai, 1976:
12
Phuket mar. biol. Cent. Res. Bull.
176–177, fig. 94a; Dai and Yang, 1991:
120 [=Platymaia wyville-thomsoni Miers,
1886].
Material examined
PMBC 20367, 4 male (71.0 × 77.5 mm, 25.9 ×
30.4 mm, 20.6 × 22.7 mm, 20.2 × 22.8 mm), 5
female (19.4 × 21.4 mm, 19.0 × 21.4 mm, 19.0 ×
22.1 mm, 18.7 × 21.2 mm, 18.4 × 20.3 mm),
BIOSHELF St. J8, 0715´N, 09730´E, Agassiz
trawl, 490–479 m, coll. S. C. Aungtonya & V.
Vongpanich, 18.02.2000
Diagnosis
Carapace tranversly subcircular, surface
nearly smooth in adults, rostral spine subequal to
interantennular spine, preocular spine absent;
protogastric ridge with 2 tubercles, lacking spines
(in adults); groove on each side of mesogastric
region deep, clearly separating adjacent branchial
regions; palm of male chelipeds more than twice
as long as high, distally increasing in width;
propodus of ambulatory legs flattened, 7.5 times
width of fourth ambulatory leg.
Remarks
Platymaia alcocki can sometimes be
misidentified with P. wywillethomsoni because
young and juveniles of P. alcocki also have
prominent spines on all the carapace regions
(Alcock, 1895). Although all the spines on the
dorsal carapace surface of adult P. alcocki are
reduced to only tubercles (Fig. 4A–C), adult P.
wywillethomsoni are still armed with one or two
spines on each side of the protogastric regions
even as adults (Griffin and Tranter, 1986a: 307,
fig. 5b). The main diagnostic feature of P. alcocki
is the deep groove on each side of mesogastric
region, which separates the two adjacent branchial
regions (Rathbun, 1916) (Fig. 4A, D). This
character is useful even for young and adult
specimens of P. alcocki and is clearly evident on
the series of Thai specimens here.
ACKNOWLEDGEMENTS
The author would like to thanks Peter K.
L. Ng of the Raffles Museum of Biodiversity
Research, National University of Singapore, for
reviewing the manuscript.
Distribution
Indian Ocean: Andaman Sea to eastern
Africa
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Decapoda Brachyura). Ann. Inst. Oceanogr., Paris. 58(1) 1982: 5–87.
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In: Report on the Scientific Results of the Voyage of HMS Challenger during the years 1873–
76, Zool. 17(49): i–xii, 1–362, pls 1–29.
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Phuket Mar. Biol. Cent., Spec. Publ. 23(2): 369–384.
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extant Brachyuran crabs of the world. Raffles Bull. Zool., Suppl. 17: 1–286.
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Taiwan (Crustacea: Decapoda). Natl. Taiwan Mus. spec. Publ. Ser. 11: 1–86, 8 col. pls.
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CSIRO Publishing. 574 pp.
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Endeavour. 5(1): 1–29, pls. 1–15
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Hist, nat., Paris. 1988. 10(4)A(1): 39–55.
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Decapoda: Brachyura). Mem. Qld. Mus. 28: 523–530.
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of a new species. Raffles Bull. Zool., Suppl. 16: 55–65.
Richer de Forges, B. and P. K. L. Ng, 2008. New records of deep-sea spider crabs of the genus
Cyrtomaia Miers, 1886, from the Pacific Ocean, with description of a new species (Crustacea:
Decapoda: Brachyura: Majidae). Zootaxa, 1861: 17–28.
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379, maps 1–3, pp. 1–16, pls. 1–251.
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Expedition, including a review of the Homolidae. Naga Rep. 4(4): 1–186, pls. 1–21.
Takeda, M. & S. Miyake. 1969. Crabs from the East China Sea. III Brachygnatha Oxyrhyncha. J. Fac.
Agric. Kyushu Univ. 15(4): 469–521, figs. 1–12, pls. 17–18.
Takeda, M. & W.R., Webber. 2006. Crabs from the Kermadec Islands. In: Tomida, Y. et al. Proceedings
of the 7th and 8th Symposia on Collection Building and Natural History Studies in Asia and the
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Manuscript received: 9 February 2011
Accepted: 1 August 2011
Phuket mar. biol. Cent. Res. Bull. 70: 15–27 (2011)
NEW RECORDS OF THE BOBTAIL SQUID, EUPRYMNA HYLLEBERGI
NATEEWATHANA, 1997 WITH DESIGNATION OF A NEOTYPE
Charatsee Aungtonya1, Anuwat Nateewathana2, Ole Secher Tendal3 and Jaruwat Nabhitabhata4
1
Phuket Marine Biological Center, P.O. Box 60, Phuket 83000, Thailand
2
Department of Marine and Coastal Resources, Bangkok, Thailand
3
Zoological Museum, SNM, University of Copenhagen, Universitetsparken 15, DK-2100
Copenhagen, Denmark
4
Centre for Biodiversity of Peninsular Thailand (CBIPT), Faculty of Science,
Prince of Songkla University, Hatyai, Songkhla 90112, Thailand
Corresponding author: Charatsee Aungtonya
Email: charatsee@gmail.com
ABSTRACT: A total of 265 specimens, 118 males and 147 females, of the bobtail squid, Euprymna
hyllebergi, are registered in the Reference Collection, Phuket Marine Biological Center, Thailand.
Nearly all specimens were collected from Thai waters, the Andaman Sea and the Gulf of Thailand.
The species is here reported as a new record for Myanmar waters and the distribution in the
Indian Ocean has been extended to the west of the Andaman Islands. A male neotype is designated
on a specimen collected from the original locality in order to replace the lost male holotype
specimen. The neotype is deposited in the Reference Collection of Phuket Marine Biological
Center, Thailand, as is the remaining female paratype, whereas selected specimens are deposited
in National Science Museum Tokyo, Japan, U.S. National Museum, Washington D.C., USA and
the Zoological Museum, SNM, University of Copenhagen, Denmark. Measurements of the
neotype, paratype, and selected specimens are presented and discussed.
Key word: bobtail squid, Euprymna hyllebergi, Thai waters, neotype
INTRODUCTION
Bobtail squids are small benthic
cephalopods, belonging to the family Sepiolidae
Leach, 1817, which contains about 14 genera and
over 50 species. Sepiolid squids are distributed
worldwide, ranging from tropical to temperate and
sub-polar waters of all oceans (Nateewathana,
1997a).
According to the most recent revision by
Norman & Lu (1997) and a later paper on the genus
by Nateewathana (1997b), 13 species have been
referred to Euprymna Steenstrup, 1887. Six
species, E. berryi Sasaki, 1929, E. hoylei Adam,
1986, E. hyllebergi Nateewathana, 1997, E. morsei
(Verrill, 1881), E. scolopes Berry, 1913, and E.
tasmanica (Peffer, 1884) are currently accepted
in the genus. Two species, E. pusilla (Peffer,
1884) and E. schneehageni (Peffer, 1884), were
placed as invalid while four species, E. albatrossae
Voss, 1963, E. bursa (Pfeffer, 1884), E. phenax
Voss, 1963, and E. stenodactyla (Grant, 1833),
were treated as unresolved species (Norman &
Lu, 1997). E. albatrossae, E. bursa, and E. phenax
were known only from the types. The original type
material of E. stenodactyla is lost, no further
material has been found in Mauritius yet and the
description of the species is poor (Norman & Lu,
1997). Furthermore, a nominal species, E. similis
Sasaki, 1913, was considered a junior synonym
of E. morsei (Norman & Lu, 1997).
The species E. morsei, E. stenodactyla,
E. berryi and E. hyllebergi, have been reported
from Thai waters, the Andaman Sea and the Gulf
of Thailand, by Chotiyaputta et al. (1992),
Chotiyaputta (1993), Supongpan (1995),
Nateewathana (1997b) and Nateewathana et al.
(2001).
16
Phuket mar. biol. Cent. Res. Bull.
According to Nateewathana et al. (2001)
the occurrence of E. morsei, E. stenodactyla, and
E. berryi in Thai waters was still uncertain since
the voucher material could not be located.
However, E. morsei and E. stenodactyla from the
Andaman Sea and Myanmar waters have been
registered at the PMBC Reference Collection in
2007. Future taxonomic study on those specimens
is pending.
All specimens of Euprymna hyllebergi
deposited in the Reference Collection of Phuket
Marine Biological Center, were checked and
measured, but none were in accordance with the
holotype. The first and the fourth authors
contacted the cephalopod taxonomists who had
previously studied the type material, but they all
denied any knowledge about the fate of those types.
We conclude that the holotype is lost.
Euprymna hyllebergi is close to E. hoylei,
but differs in the arrangement of suckers on the
hectocotylized arm, the size and distribution of
enlarged suckers on arms II–IV, and in the length
of the tentacles (Nateewathana, 1997b: Fig. 3).
E. hoylei is known from the tropical
Western Pacific to North-West Australia (Norman
& Lu, 1997). It is characterized by lacking enlarged
suckers on arm II in mature males, having 3–4
enlarged suckers in dorsal and ventral rows of arm
pair IV and ventral row only of arm pair III
(Norman & Lu, 1997, Nesis, 1987). The suckers
on arm II–IV of E. hyllebergi do not have any
significantly or abruptly enlarged suckers
(Nateewathana, 1997b). E. hyllebergi has a high
number of stout papillae with a slitlike aperture in
the distal half of the hectocotylus. The papillae of
E. hyllebergi are crowded in rows of 4–6 suckers
in the middle part, and reduced to two rows at the
tip of the hectocotylus.
The male characters are the most
important features distinguishing the two species,
E. hoylei and E. hyllebergi. Furthermore, the
arrangement of the sessile suckers of the male is
the only diagnostic feature to characterize the
species of the genus, as well as other genera in
the Family Sepiolidae. Hence it is necessary to
select a male neotype of E. hyllebergi.
The present report lists all specimens of
this species in the Reference Collection, designates
the male neotype of Euprymna hyllebergi in order
to replace the lost male holotype and paratypes
previously listed in Aungtonya et al. (2006, 2007),
and describes specimens deposited in other
museums. This study aims to facilitate future
revision and taxonomic work on this genus and to
update the known distribution of the species.
MATERIALS AND METHODS
The bobtail squids were obtained from
fishing boats, research vessels, fish markets, and
fish landings along the Andaman Sea coast of
Thailand and the Gulf of Thailand as described by
Nateewathana (1997b) and Nateewathana et al.
(2001). Additional material for the present study
was primarily obtained from the R.V. Chulaphorn
of the Deep Sea Fishery Technology Research and
Development Institute operating off the Andaman
Sea coast of Thailand, and from fishing boats, fish
markets and fish landings along the Andaman Sea
coast of Thailand and the Gulf of Thailand.
Taxonomic studies are based on Nateewathana
(1997b).
The specimen designated as the neotype
and five lots of specimens were selected from the
samples in the Reference Collection of Phuket
Marine Biological Center. The neotype was, like
the holotype, collected from small scale fishing
areas in Trang Province, at a depth of less than 40
m off the Andaman Sea Coast of Thailand.
Measurements follow those of
Nateewathana (1997b). Definitions of
measurements and indices are summarized in Table
1 and in Figure 1. The difference of the means of
all the measurements and indices is compared with
those listed by Nateewathana (1997b). Differences
were considered significant when
Specimens of the species, the newly
designated neotype, and selected materials are
deposited in the Reference Collection of Phuket
Marine Biological Center, Thailand (PMBC),
National Science Museum Tokyo, Japan (NSMT),
U.S. National Museum, Washington, D.C., USA
(USNM), and the Zoological Museum, SNM,
University of Copenhagen, Denmark (ZMUC).
17
New records of the bobtail squid
Table 1. Definition of counts, measurements and indices (modified after Nateewathana, 1997b).
ML
Mantle Length
MWI
Mantle Width Index
FLI
FWI
Fin Length Index
Fin Width Index
FBI
HLI
Fin Base Index
Head Length Index
HWI
Head Width Index
ALI
Arm Length Index
ASI
Arm Sucker Index
TtLI
Tentacle Length Index
ClLI
HcLI
Club Length Index
Hectocotylized Length Index
EDI
Eye Diameter Index
LnDI
Lens Diameter Index
dorsal mantle length measured from the anterior-most
point of mantle to posterior tip, along dorsal mid-line.
greatest straight-line (dorsal) width of mantle as a
percentage of mantle length.
greatest length of fins as a percentage of mantle length.
greatest width (dorsally) across both fins as a percentage
of mantle length.
length of fin base as a percentage of mantle length.
dorsal length of head measured from point of fusion of
dorsal arms to anterior tip of nuchal locking cartilage as a
percentage of mantle length.
width of head across midpoint of eyes as a percentage of
mantle length.
length of arm measured from first basal (proximal-most)
sucker to tip of arm as a percentage of mantle length.
diameter of largest normal arm sucker on each designated
arm as a percentage of mantle length.
total length of tentacular stalk and club as a percentage of
mantle length.
length of club as a percentage of mantle length.
length of modified portion of arm measured from
proximal-most modified sucker to tip of arm as a
percentage of total length of hectocotylized arm
(left arm I).
diameter of eye across bulbus as a percentage of mantle
length.
diameter of eyelens as a percentage of mantle length.
RESULTS
A total of 265 specimens, 118 males and
147 females, were registered in the Reference
Collection (see Table 2). Nearly all of the specimens
were collection from Thai waters, the Andaman
Sea and the Gulf of Thailand, except for PMBC
11781 and PMBC 11775 which were collected
from the Indian Ocean, west of the Andaman
Islands and Myanmar waters, respectively. The
latter specimens are the first records of E.
hyllebergi from the Indian Ocean, west of the
Andaman Islands and Myanmar waters,
respectively.
The neotype was selected from PMBC
11742 and registered as PMBC 21339 (Fig. 2).
Three lots of NSMT-Mo 76424, USNM 1150474,
and ZMUC CEP-25 have been selected from PMBC
11742, PMBC 11748 and PMBC 11747,
respectively. Two females of PMBC 11734 have
been selected and registered as NSMT-Mo 76425
and ZMUC CEP-51. The male neotype, PMBC
accession number 21339, is deposited in the
Reference Collection of Phuket Marine Biological
Center, as is the remaining female paratype, PMBC
11731.
Selected measurements and indices of the
type materials and selected specimens of E.
hyllebergi are presented in Table 3. Mean of indices
(in percent), standard deviation of five males and
four females are presented in Table 4. The t-test
comparing the means of all the measurements and
indices of the present material and those given by
Nateewathana (1997b) are presented in Table 5.
18
Phuket mar. biol. Cent. Res. Bull.
Figure 1. Diagrammatic illustration of measurements in sepiolids (modified after Nateewathana, 1997b).
AL = arm length, C1L = club length, ED = eye diameter, FB = fin base, FL = fin length, FW = fin width,
HL = head length, HW = head width, LD = lens diameter, ML = mantle length, MW = mantle width, TtL
= tentacle length.
Euprymna hyllebergi Nateewathana, 1997
(Figs. 2A–B, 3A–B, 4A–C, Table 2–5)
Euprymna hyllebergi – Nateewathana, 1997b: 466–
474, figs. 2–5, Table 2.
Neotype. PMBC 21339, 1 male, Kantang fish
landing, Trang province, 14 Feb 1996 (Figs. 2–
3).
Paratype. PMBC 11731, 1 female, Tha Lane,
Phangnga Bay, push net, 23 July 1989
Selected specimens. NSMT-Mo 76424, 1 male,
Kantang fish landing, Trang Province, 14 Feb
1996; NSMT-Mo 76425, 1 female, Chong Pra,
Phangnga Bay, push net, 18 June 1988; USNM
1150474, 1 male and 1 female, Phuket fish market,
Phuket Province, 23 Aug 1995; ZMUC CEP-25, 2
males, Krabi fish market, Krabi province, 17 Dec
1994; ZMUC CEP-51, 1 female, Chong Pra,
Phangnga Bay, push net, 18 June 1988.
Material examined. 265 specimens have been
examined and registered (see Table 2).
Description. Colour in alcohol yellowish;
numerous large and conspicuous purplish brown
chromatophores cover mantle, head and arms,
more dense in middle part of dorsal and ventral
sides. Mantle dome-shaped, thick, broad
19
New records of the bobtail squid
anteriorly and rounded posteriorly, slightly longer
than wide (Figs. 2–3, 4A–B); antero-dorsal margin
fused with head (Figs. 2, 4A). Large portion of
the tubular funnel visible, median ventral margin
slightly concave (Figs. 2B, 4B–C). Fins rounded,
almost circular in outline, separate and attached at
middle of mantle; anterior margin to posterior
margin nearly straight. Head broad, dorsoventrally
flattened and compact, width slightly less than
mantle width; eyes large, oval with almost circular
pupil. Funnel long, tubular, free for most of its
length, reaching to gap between ventral arms (Fig.
4B–C). Funnel valve triangular to rounded flaplike.
Funnel organs with a roughly triangular dorsal pad
and two oval ventral pads. Anterior end of dorsal
pad with a low crest projected in midline. Funnel
locking cartilage ovate, rounded at both ends with
deep groove (Fig. 4C); mantle cartilage straight,
ridge-like, two times longer than funnel cartilage.
Arms long, subequal, rounded in cross section,
tapering distally. Web C (between arms II and III)
developed and D distinct (between arms III and
IV). Web D extends up about one-fourth of arm
length. Suckers almost equal-sized, subglobular,
with a small, smooth aperture; pedicel thin. In
females, arm suckers quadriserial but biserial at
base and distally. In male, arm suckers globular
A
with smooth horny ring, rounded aperture on
normal sucker, but more ovate on enlarged one.
Left arm I of male hectocotylized, stout, shorter
than other arms and curved outwards
(Nateewathana, 1997b: figs. 3a, 4a–b). The size
of arm suckers on different arms varied. Tentacles
round, flattened on oral surface, edges of flat area
angled, dorsal margin elevated into a low keel
towards club. Club short, curled, and rounded with
numerous rows of very small suckers. Gladius
absent. Buccal membrane smooth without suckers.
Type locality. Small scale fishing areas in Trang
Province, in the depth less than 40 m, the Andaman
Sea coast of Thailand.
Distribution. Indian Ocean (Andaman Sea and
Myanmar waters with range extension to the west
of the Andaman Islands) and Gulf of Thailand.
Remarks. The characters of all specimens match
well with those of E. hyllebergi, as described and
figured by Nateewathana (1997b). Description of
beak, radula, ink sac, and spermatophores of the
species are not repeated herein.
The right tentacle in NSMT-Mo 76424 and
the left tentacle in ZMUC CEP-51 are missing.
B
Figure 2: Euprymna hyllebergi Nateewathana, 1997 (PMBC 21339). Neotype, male: (A) dorsal and
(B) ventral.
20
Phuket mar. biol. Cent. Res. Bull.
A
B
Figure 3: Euprymna hyllebergi Nateewathana, 1997 (PMBC 11731). Paratype, female: (A) dorsal and
(B) ventral.
A
B
C
Figure 4. Euprymna hyllebergi Nateewathana, 1997 (PMBC 21339). Neotype, male: (A) dorsal view;
(B) ventral view; (C) funnel and funnel locking-cartilages. Scales = 2 cm (A–B) and 0.5 cm (C).
Table 2. Details of specimens of Euprymna hyllebergi remaining in the Reference Collection of Phuket Marine Biological Center.
No. PMBC No.
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
11727
11728
11729
11730
11731
11732
11733
11734
11735
11736
11737
11738
11739
11740
11741
11742
11744
11745
11746
11747
11748
11749
11750
11751
11752
11753
11755
11756
11757
11758
11759
11760
11761
5
1
HOLOTYPE specimen lost
6
11
PARATYPE
1
1
6
3
2
3
PARATYPE specimen lost
5
PARATYPE specimens lost
9
9
32
12
1
1
5
12
2
1
1
4
3
13
4
1
1
1
3
1
3
Female
Locality
Province
Identifier
2
–
Phuket
A. Nateewathana
Krabi
Trang
Trang
Ranong
Phangnga
Trang
Krabi
Phangnga
Phangnga
Phangnga
Trang
Andaman Sea
Krabi
Phangnga
Phangnga
Trang
Krabi
Krabi
Ranong
Krabi
Phuket
Phuket
Phangnga
Phangnga
Phuket
Krabi
Satul
Phangnga
Ranong
Andaman Sea
Phangnga
Phuket
Krabi
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
1
2
5
3
1
1
4
6
1
1
3
2
1
3
5
2
3
15
6
4
5
1
2
3
6
1
1
2
7
6
17
6
1
1
1
7
1
1
1
2
7
3
1
1
1
1
Krabi fish market
Kantang fish landing
Kantang fish landing
Kapur
Tha Lane, Phangnga Bay
Rok Island
Krabi fish market
Chong Pra, Phangnga Bay
Raya Ring Island
Tha Lane, Phangnga Bay
Kantang fish landing
8°02´N, 98°30´E
Southwest of Lanta Island
Taeng Cape, Phangnga Bay
Phangnga Bay
Kantang fish landing
Krabi fish market
Krabi fish market
–
Krabi fish market
Phuket fish market
Tukkae Cape
Raya Ring Island, Phangnga Bay
Mak Island, Phangnga Bay
Phuket fish market
Southwest of Lanta Island
Tamalang fish landing
–
Jam Island
7°48´N, 98°30´E
Nakae Island
Sapum Bay
Dam Hok Island
Remarks
original recorded as
Euprymna stenodactyla
(Grant, 1833) by C.F.E.
Roper in 1979
21
1607
No. of spec. Male
New records of the bobtail squid
1
Type
No. PMBC No.
22
Table 2. (continued)
No. of spec.
NEOTYPE
1
2
2
1
1
1
2
1
3
1
1
1
18
1
1
1
3
16
26
1
10
1
1
Male
Female
58
23937
4
3
1
–
–
A. Nateewathana
59
25004
5
3
2
–
–
A. Nateewathana
2
5
2
10
17
1
6
4
1
1
Trang
Phangnga
Krabi
–
Krabi
Andaman Sea
Andaman Sea
Phuket
Krabi
Ranong
Trang
Myanmar waters
Pattani
Trang
Samut Prakan
Indian Ocean
Andaman Sea
Trang
Trang
Andaman Sea
Andaman Sea
Andaman Sea
Trang
Identifier
11762
11763
11764
11765
11766
11767
11768
11770
11772
11773
11774
11775
11776
11777
11780
11781
11782
11785
11787
14420
19903
19904
21339
1
1
2
1
1
1
1
1
13
1
1
1
1
6
9
Trang fish market
off Yao Yai Island
East of Lanta Island
–
–
6°58´N, 98°48´E
6°57´N, 98°47´E
Rang Yai Island
–
–
Kantang fish landing
13°27´N, 97°00´E
Pattani fish landing
Kantang fish landing
Samut Prakan fish landing
7°17´N, 90°00´E
7°04´N, 98°53´E
Kantang fish landing
Kantang fish landing
9°34´N, 97°49´E
7°51´N, 98°38´E
9°00´N, 97°54´E
Kantang fish landing
Province
35
36
37
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
1
1
1
2
1
Locality
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
A. Nateewathana
T. Kubodera
T. Kubodera
A. Nateewathana
Remarks
specimen was selected
from PMBC 11742
specimens found in the
bottle of PMBC
11737, but not
identical to original
paratype
specimens found in the
bottle of PMBC
11739, but not
identical to original
paratype specimens
Remarks: the number of specimens in some PMBC accession numbers is changed, compared to that published in Appendix 1 of Nateewathana (1997b), and some
PMBC accession numbers were not included in that appendix.
Phuket mar. biol. Cent. Res. Bull.
Type
Table 3. Measurements (mm) and indices (%) of neotype, paratype, and selected specimens of Euprymna hyllebergi from the Andaman Sea,
Thailand.
Paratype
Registration
No.
PMBC
21339
PMBC
11731
NSMT-Mo
76424
NSMT-Mo
76425
Index\Sex
Male
Female
Male
Female
Male
ML (mm)
MWI
FLI
FWI
FBI
HLI
HWI
ALII
ALIII
ALIIII
ALIVI
ASII
ASIII
ASIIII
ASIVI
TtLI
CILI
HcLI
EDI
LnDI
30.98
84.83
44.09
129.50
26.44
54.71
63.52
106.751
143.54
120.27
116.17
3.23
4.03
5.23
6.17
327.73
42.32
65.73
22.60
12.33
33.63
77.19
48.41
119.92
28.49
52.81
61.11
97.71
114.93
98.331
110.70
2.97
4.16
4.46
4.46
234.672
46.65
59.99
19.80
13.56
35.22
74.82
51.02
135.15
33.19
50.03
59.71
59.00
74.761
69.96
67.181
2.13
2.30
2.98
2.73
84.44
31.63
29.02
90.45
54.51
146.76
35.70
53.62
64.13
91.94
124.911
104.311
105.10
3.96
3.79
5.34
5.00
290.18
31.741
62.93
27.08
13.20
22.27
91.02
58.96
152.09
46.74
46.43
73.15
69.33
86.30
77.191
73.37
2.92
3.37
2.92
2.25
119.98
34.98
25.82
15.67
Selected specimens
24.11
12.12
USNM
1150474
Female
26.79
88.28
50.09
159.69
39.94
64.05
67.19
91.75
99.22
93.77
96.90
2.24
2.61
2.43
2.43
246.62
62.64
24.15
15.57
ZMUC CEP
25
ZMUC CEP
51
Male
Female
27.21
83.83
52.11
132.82
32.97
53.55
68.83
88.06
110.66
101.14
98.46
2.94
2.94
4.41
4.59
285.34
42.08
58.01
30.10
15.40
28.65
81.85
48.94
134.90
37.63
58.71
64.96
99.34
124.15
108.34
101.22
4.01
3.66
5.34
4.54
229.60
41.64
60.94
26.35
14.52
28.42
76.18
57.74
135.50
34.24
57.35
57.74
84.87
102.39
91.87
88.53
2.53
2.43
3.20
2.81
169.11
39.94
0.00
25.97
13.62
New records of the bobtail squid
Neotype
Remarks: 1 = tip of the right arm broken; 2 = left tentacle due to the right one lost. The values in bold text are out of the range of the
measurements and indices reported by Nateewathana (1997b).
23
24
Phuket mar. biol. Cent. Res. Bull.
Table 4. Mean (X) of indices (%) and standard deviation of type materials and selected specimens
of Euprymna hyllebergi from the Andaman Sea, Thailand.
Male
Female
Index
ML (mm)
MWI
FLI
FWI
FBI
HLI
HWI
ALII
ALIII
ALIIII
ALIVI
ASII
ASIII
ASIIII
ASIVI
TtLI
CILI
HcLI
EDI
LnDI
n
Mean
SD
5
5
5
5
5
5
5
5
5
5
5
5
5
5
5
5
5
5
5
5
29.90
83.63
49.61
132.78
32.24
54.68
64.51
96.76
123.64
106.48
106.33
3.42
3.72
4.96
4.95
273.50
40.89
61.52
25.19
13.80
2.48
4.81
3.96
9.70
4.72
2.35
2.81
7.18
12.67
8.56
7.17
0.53
0.48
0.48
0.71
41.21
5.50
2.95
4.02
1.19
Range
n
Mean
SD
27.21–33.63
77.19–90.45
44.09–54.51
119.92–146.76
26.44–37.63
52.81–58.71
61.11–68.83
88.06–106.75
110.66–143.54
98.33–120.27
98.46–116.17
2.94–4.01
2.94–4.16
4.41–5.34
4.46–6.17
229.60–327.73
31.74–46.65
58.01–65.73
19.98–30.10
12.33–15.40
4
4
4
4
4
4
4
4
4
4
4
4
4
4
4
4
4
4
4
4
28.18
82.57
54.45
145.61
38.53
54.47
64.45
76.24
90.67
83.20
81.50
2.46
2.68
2.88
2.55
155.04
42.30
5.37
8.27
4.54
12.27
6.23
7.84
7.09
14.83
12.68
11.52
13.64
0.35
0.48
0.33
0.26
70.23
13.98
25.01
14.24
1.02
1.70
Range
22.27–35.22
74.82–91.02
50.09–58.96
135.15–159.69
33.19–46.74
46.43–64.05
57.74–73.15
59.00–91.75
74.76–102.39
69.96–93.77
67.18–96.90
2.13–2.92
2.30–3.37
2.43–3.20
2.25–2.81
84.44–246.62
31.63–62.64
24.11–25.97
12.12–15.67
Table 5. t-test of the mean of all measurements and indices by Nateewathana (1997b) and this study.
Nateewathana (1997b)
Male
present study
Index
n
Mean
SD
n
Mean
ML (mm)
MWI
FLI
FWI
FBI
HLI
HWI
ALII
ALIII
ALIIII
ALIVI
ASII
ASIII
ASIIII
ASIVI
TtLI
CILI
HcLI
EDI
LnDI
20
20
20
20
20
20
20
20
20
20
20
20
20
20
20
20
20
20
20
20
27.20
81.20
49.10
131.10
31.50
52.80
65.20
103.70
141.00
118.20
118.60
3.50
4.40
5.10
5.30
282.80
48.40
63.40
23.30
14.40
4.20
11.30
7.10
13.80
4.00
8.50
7.10
20.00
15.70
14.10
18.00
0.50
0.60
0.70
0.60
47.10
7.10
8.30
5.60
2.80
5
5
5
5
5
5
5
5
5
5
5
5
5
5
5
5
5
5
5
5
29.90
83.63
49.61
132.78
32.24
54.68
64.51
96.76
123.64
106.48
106.33
3.42
3.72
4.96
4.95
273.50
40.89
61.52
25.19
13.80
SD
T-test
2.48
4.81
3.96
9.70
4.72
2.35
2.81
7.18
12.67
8.56
7.17
0.53
0.48
0.48
0.71
41.21
5.50
2.95
4.02
1.19
No diff
No diff
No diff
No diff
No diff
No diff
No diff
No diff
Sig diff
No diff
No diff
No diff
Sig diff
No diff
No diff
No diff
Sig diff
No diff
No diff
No diff
25
New records of the bobtail squid
Table 5. (continued)
Nateewathana (1997b)
Female
Index
n
ML (mm)
MWI
FLI
FWI
FBI
HLI
HWI
ALII
ALIII
ALIIII
ALIVI
ASII
ASIII
ASIIII
ASIVI
TtLI
CILI
EDI
LnDI
20
20
20
20
20
20
20
20
20
20
20
20
20
20
20
19
19
20
20
Mean
24.90
80.70
52.50
142.40
35.80
55.30
67.70
85.40
110.60
93.20
89.70
2.70
2.80
3.00
2.80
266.60
53.30
23.50
14.90
present study
SD
n
Mean
SD
T-test
5.50
8.90
7.80
12.80
5.00
6.20
7.90
11.40
12.30
8.80
9.20
0.40
0.50
0.40
0.40
60.70
10.90
3.80
2.10
4
4
4
4
4
4
4
4
4
4
4
4
4
4
4
4
4
4
4
28.18
82.57
54.45
145.61
38.53
54.47
64.45
76.24
90.67
83.20
81.50
2.46
2.68
2.88
2.55
155.04
42.30
25.01
14.24
5.37
8.27
4.54
12.27
6.23
7.84
7.09
14.83
12.68
11.52
13.64
0.35
0.48
0.33
0.26
70.23
13.98
1.02
1.70
No diff
No diff
No diff
No diff
No diff
No diff
No diff
No diff
Sig diff
No diff
No diff
No diff
No diff
No diff
No diff
Sig diff
No diff
No diff
No diff
DISCUSSION
The Euprymna hyllebergi arm length
formula is II > III > IV > I, but occasionally II >
III = IV > I (Nateewathana, 1997b). The arm length
formulas of the present material of males and
females (Table 4, II > III > IV > I), matches those
of Nateewathana (1997b) as well as those in the
neotype and paratype (Table 3). Differences are,
however, found in some of the selected specimens;
four specimens show II > III > IV > I but three
other specimens show II > IV > III > I (Table 3).
These differences occur because the tips of the
right arms were broken.
All measurements and indices of the
neotype are in the ranges reported by
Nateewathana (1997b). Some measurements and
indices in the paratype and selected specimens are,
however, out of these ranges, i.e., Mantle Length
(ML), Arm Length Index (ALI), Arm Sucker Index
(ASI), Tentacle Length Index (TtLI), and Club
Length Index (ClLI) (Tables 3–4). Furthermore,
the t-test of the mean of all measurements and
indices by Nateewathana (1997b) and in this study
were considered significant for ALII in both male
and female, for ASIII and ClLI in male only, and
for TtLI in female (Table 5).
These differences may occur because the
suckers on arms easily fall off and are lost in
preserved material and the tip of the first right arm
in male specimen was broken. Therefore some
errors occurred in estimation of the length of the
first right arm, which was measured from the first
basal (proxima-most) sucker to tip of arm as a
percentage of mantle length.
Furthermore, according to C.C. Lu (pers.
comm., in Nateewathana, 1997b), the length of
the tentacles in cephalopods may not be a usable
taxonomic character since it varies greatly
depending on how the specimen is fixed and
preserved. Live fixed animals have short tentacles
because of contraction and, on the other hand,
specimens thawed from frozen condition have long
tentacles. In order for this character to be of some
26
Phuket mar. biol. Cent. Res. Bull.
use, specimens should be fixed under identical
conditions. Specimens from the same batch may
vary greatly (C.C. Lu, pers. comm., in
Nateewathana, 1977b).
The mantle length, the first arm length
index, the tentacle length index, and club length
index are the main characters/indices used for
distinguishing the species from other members of
the genus. Although the tentacle length is
particularly important in recognizing this species,
it is not necessarily an autopomorphy for the
species. It is the combination of the arrangement
of suckers on the hectocotylized arm, the size and
distribution of enlarged suckers on arms II–IV,
and in the length of the tentacles that distinguishes
Euprymna hyllebergi.
ACKNOWLEDGEMENTS
We express our appreciation to Dr.
Kongkiat Kittiwattanawong, Phuket Marine
Biological Center (PMBC) for his kind assistance
in t-test Analysis, and Dr. Janek von Byern,
University of Vienna, Austria, for providing valuable
comments on the draft of this manuscript. Sincere
thanks are given to Ms. Kanjana Bussabong, the
Reference Collection, Phuket Marine Biological
Center, and Ms. Thitima Boonkaew, training
student, PSU, for their kind assistance. Our sincere
thanks also go to Mr. Rueangrit Promdam, (former
staff of Reference Collection), and Mr. Patirat
Singdum, the PMBC artist, for photographing and
drawings.
REFERENCES
Adam, W. 1986. Contribution à la connaissance du genre Euprymna Steenstrup, 1887 (Mollusca:
Cephalopoda). Bulletin de l’Institut Royal des Science Naturelles de Belgique, Biologie. 56:
131–136.
Aungtonya, C., K. Bussabong, K. Wongkamhaeng, I. Eedkerd, and S. Ubonsuwan. 2006. Type specimens
in the Reference Collection of Phuket Marine Biological Center. Reference Collection and
Aquarium Contributions no. 4: 45 p. (in Thai).
Aungtonya, C., S. Sungjeen, W. Tripanichkul, K. Bussabong, and P. Kongthaworn. 2007. Recent
records of the Class Cephalopoda (Phylum Mollusca) in the Reference Collection of Phuket
Marine Biological Center. Reference Collection and Aquarium Contributions no. 5: 69 p. (in
Thai).
Berry, S. S. 1913. Some new Hawaiian cephalopods. Proc. U. S. natn. Mus. 45: 563–566.
Chotiyaputta, C. 1993. Cephalopod resources of Thailand. Pp. 71–80. In: Okutani,T., R.K. O’Dor &
T. Kubodera (eds.). Recent Advances in Fisheries Biology. Tokai University Press, Tokyo.
Chotiyaputta, C., T. Okutani, and S. Chaitiamvong. 1992. Systematic study of cephalopods in Thailand.
A Research Report submitted to the National Research Council of Thailand under JSPS-NRCT
Cooperation Project. 100 pp. (In Thai with English abstract).
Grant, R. E. 1883. On a new species of Sepiola (Sepiola stenodactyla) from the Mauritius, presented
by C. Telfair, Esq. Proceedings of the Zoological Society of London. 1: 42–43.
Nateewathana, A. 1997a. Systematics of the Cephalopoda of the Andaman Sea, Thailand. Ph.D. Thesis.
Institute of Biological Sciences, Faculty of Natural Sciences, University of Aarhus. 343 pp.
Nateewathana, A. 1997b. The Sepiolidae (Cephalopoda) of the Andaman Sea, Thailand, with description
of Euprymna hyllebergi sp. nov. Phuket Marine Biological Center Special Publication. 17:
465–481.
Nateewathana, A., J. Nabhitabhata, and P. Nilaphat. 2001. A new record of bobtail squid, Euprymna
hyllebergi Nateewathana, 1997 in the Gulf of Thailand. Phuket Marine Biological Center Special
Publication. 25(2): 501–506.
Nesis, K.N. 1987. Cephalopods of the world. T.F.H. Publications, Inc. Ltd., New Jersey. 351 pp.
27
New records of the bobtail squid
Norman, M.D. & C.C. Lu. 1997. Redescription of the southern dumpling squid Euprymna tasmanica
and a revision of the genus Euprymna (Cephalopoda: Sepiolidae). J. Mar. Biol. Ass. U.K. 77:
1109–1137.
Pfeffer, G.J. 1884. Die Cephalopoden des Hamburger Naturhistorischen Museums. Abhandlungen des
Naturwissenschaftlichen Vereins in Hamburg. 8(2): 63–90.
Sasaki, M. 1913. Decapod cephalopods found in Japan: Sepiolidae. Zool. Mag., Tokyo. 25: 247–252.
Steenstrup, J.J. 1887. Notae teuthologicae, no. 6. Species generic Sepiolae Maris Mediterranei. Oversi.
K. Danske Vidensk Selsk Forh. Copenhagen. 1887: 47–66.
Supongpan, M. 1995. Cephalopod resources in the Gulf of Thailand. pp. 191–203. In: Nabhitabhata, J.
(ed). Biology and Culture of Cephalopods. Rayong Coastal Aquaculture Station, Contribution
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Verrill, A.E. 1881. The cephalopods of the north-eastern coast of America: part 2. The smaller
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Manuscript received: 17 August 2009
Accepted: 12 June 2011
28
Phuket mar. biol. Cent. Res. Bull.
Phuket mar. biol. Cent. Res. Bull. 70: 29–77 (2011)
THE FISHES OF SOUTHWESTERN THAILAND, THE ANDAMAN SEA – A REVIEW OF
RESEARCH AND A PROVISIONAL CHECKLIST OF SPECIES
Ukkrit Satapoomin
Phuket Marine Biological Center, P.O. Box 60, Phuket 83000, Thailand
Email: ukkrit@yahoo.com
ABSTRACT: A brief review of recent ichthyological research and compiled results on the records
of fishes from the Andaman Sea coast of Thailand are presented. A total of 1,746 species in 198
families of fishes are currently known from the area. The 10 most speciose families are the
Gobiidae (227 species), Labridae (78), Pomacentridae (71), Serranidae (61), Apogonidae (60),
Blenniidae (52), Carangidae (52), Scorpaenidae (49), Lutjanidae (39) and Chaetodontidae (37),
and these together account for 42% of the total fish fauna. The ichthyofauna is dominated by
reef-associated fishes (983 species) and pelagic/benthic fishes inhabiting offshore habitats (971
species). A zoogeographic analysis reveals a peculiarity of the fauna of this marginal-sea region,
viz., being one of the areas of sympatry of Indian and Pacific Ocean fishes, as well as harbouring
regional endemics. The PMBC Reference Collection currently contains examples of only about
63% of the fishes known from the area, indicating a need for further local sampling campaigns
and/or international collaborative exploratory research programs in the Andaman Sea to build up
the Reference Collection.
Keywords: Thailand, Andaman Sea, fishes, checklist, diversity.
INTRODUCTION
The Andaman Sea is an approximately
1,200 km long x 650 km wide body of water to
the southeast of the Bay of Bengal. It is bounded
by the Gulf of Mataban in the north, the Malay
Peninsula in the east, the Andaman and Nicobar
Islands in the west and the northern tip of Sumatra
in the south. At its southeastern corner, the
Andaman Sea narrows to form the Strait of
Malacca, which separates the Malay Peninsula from
the island of Sumatra. Along an approximate 1,000
km length of the Andaman Sea coast of Thailand,
a variety of marine and coastal habitats are present,
viz., mangroves, mud-flats, seagrass beds, rocky
shores, intertidal sand-flats, coral reefs, open
pelagic zones, and complex seabed systems on the
continental shelf. The integrity of different peculiar
environmental settings and/or characteristics
among those habitat types provides suitable refuges
for a vast variety of aquatic biota, including fishes.
Although, biogeographically, it belongs to the
eastern Indian Ocean, it is strongly influenced by
the western Pacific Ocean biota via the
northwesterly through-flow in the Strait of
Malacca. It is thus an area of distributional
confluence of the biotas of both the Indian Ocean
and the Pacific Ocean (Satapoomin, 2002a, 2007;
Kimura and Satapoomin, 2009). Being a marginal
sea separated from the main Indian Ocean by the
Andaman-Nicobar island arcs, it has been subjected
to a profound effect of sea-level lowering, especially
during major glaciations when it became a
separated, enclosed basin. Such isolation processes
are believed to have promoted speciation and/or
endemism of the fish fauna in the Andaman Sea
(e.g., McManus, 1985; Randall, 1998; Randall and
Satapoomin, 1999). The area is thus characterized
by its high diversity and the unique faunal
composition of the fishes. However, knowledge
and information on the overall diversity of marine
fishes in the area is still limited, because very few
attempts had been made to compile a list of all
known fish species from the area.
The purposes of the present paper are to
provide a brief review of recent ichthyological
research, and to establish a comprehensive
checklist of all known marine and brackish-water
30
Phuket mar. biol. Cent. Res. Bull.
fishes from the Andaman Sea coast of Thailand.
The checklist was compiled from various sources
of information, including published literature,
unpublished data made available under several
sampling programs/projects, and the fish collection
data repository of the Phuket Marine Biological
Center (PMBC). It should be noted that the overall
list of species presented herein is only provisional
as a number of species records and identifications
in previously published literature (particularly those
of inventory, non-taxonomic works), cannot be
verified. Nonetheless, this provisional checklist will
serve as a significant baseline for further
establishment of a conventional, annotated
checklist or taxonomic review of the Andaman Sea
fishes.
METHODS
The compilation of the species list was
based largely on accumulating information from
the earlier literature, including inventory works and/
or faunistic surveys, field guides, and taxonomic
works. Some unpublished data, including recent
unpublished survey results made by the author,
collection data of the Royal Ontario Museum’s
project on gobioid fishes of southwestern Thailand
in 1993 (under auspices of Dr. Richard
Winterbottom), and the PMBC fish collection data
repository were also incorporated into the current
checklist. Selected taxonomic papers pertaining to
records of fishes from the Andaman Sea coast of
Thailand used to compile the current checklist are
listed in Appendix B.
The systematic arrangement of families
follows Nelson (2006). Species in each family are
arranged alphabetically. The scientific names of
fishes used in the cited literature have been updated
where possible, following recent usage, and hence
junior synonyms were omitted. Since the
compilation was also based, in part, on literature
that provided only a list of names, the author has
only been able to check or correct
misidentifications to a limited extent. In this regard,
only correct names were retained, synonyms were
allocated according to the latest usage, while
doubtful species records were excluded.
In addition to scientific names and
classification, the checklist provides basic
information on habitats and zoogeographic
distribution of each species. The availability of
voucher specimens lodged at the Reference
Collection of the PMBC was also emphasized in
order to draw the attention of ichthyologists who
may wish to examine these specimens. The
designation of habitat type(s) for each species was
based largely on either quoted information from
the compiled literature or from the author’s
unpublished survey results and observations. Four
principal habitat categories adopted herein included
mangrove (Mg), coral reef (Cr), seagrass bed (Sg),
and other offshore habitats (OShr). These
categories can be considered as good
representations of the major marine and coastal
features along the southwestern Thailand coastline.
Each of these habitat types, however, was more
or less loosely defined in order to accommodate a
range of associated minor biotopes of each habitat.
The offshore category, in particular, was the most
inclusive term as it encompassed a range of
benthic- and pelagic-related habitats. This
designation includes both near-shore and faroffshore fishes considered as either pelagic (i.e.,
coastal pelagic, offshore pelagic, epipelagic, and
mesopelagic) or demersal (i.e., coastal demersal,
benthopelagic, and deep demersal) species. The
mangrove habitat was considered to include certain
inter-connected zones like mud-flats and the upper
reaches of the estuarine areas. Coral reef was
defined to include several nearby and/or associated
areas, namely intertidal rocky shore/rock platform,
intertidal sand-flat, and subtidal bare-seafloor in
front of the reef (so called as forereef).
Most of the well defined zoogeographic
distribution categories (see, e.g., Myers, 1991)
were generally adopted or redefined with additional
explanation, while some new categories were added
with their definitions. The definitions for those
distribution categories and their acronyms are as
follows: Indo-Pacific (IP), widespread across the
Indian Ocean and western portion of the Pacific
with an eastern limit east of non-marginal areas of
the Pacific Plate; Indo-west Pacific (IWP), a subset
of the Indo-Pacific with northeastern limit on the
Pacific Plate, marginally in the Carolines but not
reaching the Marshalls; West Pacific (WP), limited
to the area bound by the Andaman Island and
Cocos-Keeling Island in the west to the margin of
31
The fishes of southwestern Thailand
the Pacific Plate in the east, north to southern
Japan, south to western and eastern Australia; West
central Pacific (WCP), west Pacific and Pacific
Plate distributions and may include the East Pacific;
Indo-Australian Archipelago (IA), a subset of the
West Pacific, north to southern Taiwan, east to
the Solomons, south to the Great Barrier Reef;
Indo-Malayan Archipelago (IM), a subset of IndoAustralian which is limited to the areas within
bounds of Malay Peninsula, Sumatra, Java, New
Guinea and Philippines; Indian Ocean (IO),
widespread throughout the Indian Ocean; Northern
Indian Ocean (NIO), the extended range of eastern
Indian Ocean to encompass the northwestern
limits which include the west coast of India,
Pakistan, Oman and Persian Gulf; Eastern Indian
Ocean (EIO), the eastern section of the Indian
Ocean westwards to Laccadive, Maldive and
Chagos archipelagoes; the eastern extremity of the
Indian Ocean (EEIO), a narrow stripe or transition
area along the joint margin of Indian-Australian
and Asian lithospheric plates, including the
Andaman Sea, Sumatra, Java, Bali, Flores, CocosKeeling/Christmas Islands and northwestern
Australian Shelf; Andaman Sea province (AS), the
area assigned to accommodate species having
restricted distribution only to the Andaman Sea or
ranging southwards only to western Sumatra;
Circumtropical (CIR), tropical seas worldwide; and
Cosmopolitan (COS), tropical and temperate seas
worldwide. The boundary of some distribution
categories is presented (Fig. 1) for clarification of
the definitions of those areas.
RESULTS AND DISCUSSION
1. Brief review of ichthyological research
The fish fauna of the Andaman Sea coast
of Thailand was poorly documented until relatively
recent times. There were several well-known
western ichthyologists/naturalists (e.g., Cuvier,
Valenciennes, Bleeker, Günther, Fowler, Regan,
Boulenger, etc.) who reported, listed or described
Thai fishes between 1800 and the 1950s, but their
works were restricted only to either the Gulf of
Thailand (marine) or inland (freshwater) fishes
(see Monkolprasit et al. (1997) for detailed
historical review). The first marine fish of the
Andaman Sea coast of Thailand described by a
westerner was a reef-associated goby, Pipidonia
quinquecincta Smith, 1931 [valid as Gobiopsis
quinquecincta (Smith)]. During his term of service
(1923–1935) as the Adviser in Fisheries to the
Department of Fisheries of Thailand (formerly
Siamese Bureau of Fisheries), Smith published a
number of articles pertaining to freshwater fishes
and fishes of the Gulf of Thailand. Later, in 1945,
he published a monograph entitled “The Freshwater Fishes of Siam, or Thailand” (Smith, 1954).
Suvatti (1950) was the first Thai ichthyologist to
publish a list of Thai fishes. Several subsequent
studies on marine fishes in Thai waters by the
Thais were confined to groups of particular
interest, for instance Pleuronectiformes (by
Punpoka, 1964), Lutjanidae (Banasopit, 1968),
Serranidae (Banasopit, 1969; Monkolprasit, 1983),
Chaetodontidae (Banasopit, 1971), Leiognathidae
(Monkolprasit,
1973),
Elasmobranchii
(Monkolprasit, 1984), and Labridae (Monkolprasit,
1989). The foremost comprehensive compiled
checklist of the fishes of Thailand, which included
both freshwater and marine fishes, was published
by Monkolprasit et al. in 1997. It listed 171 fish
families with a total of 1,741 species, about 30%
(516 species) of which can be considered as
freshwater fishes. The records of marine fishes
in the checklist were compiled from the literature
published before 1990, except for two papers (one
on mangrove fishes by Monkolprasit (1994) and
one on ammodytid fishes by Ida et al. (1994)).
That checklist recorded 710 species representing
129 families of fishes from the Andaman Sea.
A few collections of reef fishes from the
Similan Islands were made during the R/V Te Vega
Cruise 2, eastern Indian Ocean cruise (from
Singapore to Colombo, Ceylon), in late 1963 (http:/
/tevega.stanford.edu/cruises.htm).
These
collections were deposited at the U.S. National
Museum of Natural History, and some were
designated as types of new fish species described
subsequently (i.e., Melichthys indicus Randall &
Klausewitz, 1973, Gymnothorax pseudoherrei
Böhlke, 2000, and Helcogramma lacuna Williams
& Howe, 2003). In February 1979, Randall
(Bishop Museum, Hawaii) and Allen (Western
Australian Museum) made a few collections and
photographed reef fishes around Phuket and the
Similan Islands (see Allen et al., 2005). Although
32
Phuket mar. biol. Cent. Res. Bull.
Figure 1. Approximate range of some zoogeographic distribution categories. A: Indo-Pacific (IP),
Indo-west Pacific (IWP), West central Pacific (WCP), and West Pacific (WP); B: Indo-Australian
archipelago (IA) and Indo-Malayan Archipelago (IM); C: Northern Indian Ocean (NIO), Eastern Indian
Ocean (EIO), and Eastern extremity of Indian Ocean (EEIO).
no comprehensive list of fishes resulted from that
effort, the collected materials had been used in
describing several new species (i.e., Leptojulis
chrysotaenia Randall & Ferraris, 1981, Chromis
flavipectoralis Randall, 1988, Pomacentrus adelus
Allen, 1991, P. azuremaculatus Allen, 1991, P.
polyspinus Allen, 1991, P. similis Allen, 1991,
Neopomacentrus sororius Randall & Allen, 2005,
Plesiops auritus Mooi, 1995, Plectorhinchus
macrospilus Satapoomin & Randall, 2000,
Ctenochaetus truncatus Randall & Clements, 2001,
Scorpaenopsis possi Randall & Eschmeyer, 2001,
Helcogramma lacuna Williams & Howe, 2003, and
Opistognathus cyanospilotus Smith-Vaniz, 2009).
The most important collections of reef fishes for
the area were made in November 1993 under the
collaborative project between the Royal Ontario
Museum (ROM), Canada and the Phuket Marine
Biological Center (PMBC). From well over 35
sampling stations using the ichthyocide rotenone
33
The fishes of southwestern Thailand
around Phuket and adjacent islands, nearly eighteen
thousand specimens representing about 570 species
in 75 families of fishes were collected and
documented. It was also the first time that
significant information on small cryptobenthic reef
fishes for the area was obtained. The collected
materials have been used in the descriptions of 13
new species to date (including 7 species in Gobiidae
and 1 in each of the following families: Apogonidae,
Serranidae, Pomacentridae, Opistognathidae,
Tripterygiidae, and Soleidae), with several other
descriptions underway. Satapoomin (1993)
established a first comprehensive checklist of coral
reef fishes of the west coast of Thailand by
compiling information from both available literature
(dated back to 1950) and the author’s survey
results carried out during 1990–1992. The list
revealed a total of 597 species belonging to 66
families of reef fishes in the area. Allen et al. (2005)
provided an extensive list, consisting of 764 species
in 70 families, of coral reef fishes being recorded
and/or known for the area. The most recent list of
coral reef fishes, with 888 species in 85 families,
was published by Satapoomin (2007).
There are a limited number of studies on
ichthyofauna of either mangroves or seagrass beds
in the Andaman Sea coast of Thailand, most of
which were confined to particular sampling sites
(e.g., Boonruang et al., 1994; Monkolprasit, 1994;
Janekitkarn and Monkolprasit, 1994). Satapoomin
and Poovachiranon (1997) provided the first
comprehensive list of the fish fauna from several
major mangrove and seagrass bed sites along the
coast of southwestern Thailand. That study
revealed a total of 280 species representing 75
families of fishes, which included 232 species (69
families) and 149 species (51 families) found in
mangroves and seagrass beds, respectively. In
2002, a study by Fish Team of the Trang Project
(2002) provided a list, together with illustrations,
of 134 species in 49 families of fishes inhabiting a
mangrove estuary at Sikao, southwestern Thailand.
Under the JSPS (Japan Society for the Promotion
of Science) Multilateral Project on Coastal Marine
Science, Matsuura and Kimura (2005) published
a guide book with taxonomic accounts and
illustrations of 128 species in 46 families of fishes
inhabiting mangroves, seagrass beds and
surrounding area of Libong Island, Trang Province.
The most recent list of the fish fauna in the seagrass
beds of the Andaman Sea coast of Thailand, with
230 species in 62 families, was published by
Satapoomin (2006).
The first publication on demersal and/or
pelagic fishes of the Andaman Sea coast of
Thailand was published by Lohakarn (1968). This
included taxonomic accounts of 109 species in 57
families. Fishes collected by bottom trawls during
the 5th Thai-Danish Expedition to the Andaman
Sea in 1966 (the first Thai-Danish marine
expedition) were reported by Boonyanate and
Hylleberg (1993) with a total records of 80 species
in 41 families. Other early studies on demersal
fishes in the area were mostly focused on fisheries
aspects and, in most cases, faunal composition
was reported mainly as higher taxonomic ranks
rather than individual species (e.g., Pokapunt and
Sinoda, 1978; Boonragsa, 1988; Chantawong,
1996). Some studies on experimental operations
of certain types of fishing gears reported not more
than 50 fish species each (e.g., Wongratana,
1982b, 1983; Ruengpan, 1988). A more
comprehensive list of demersal fishes collected by
otterboard trawl nets and deep sea shrimp traps
along the Andaman Sea coast of Thailand at depths
of about 30–420 m was published by Wongratana
(1982a), with 256 species in 91 families. Also,
Pokapunt and Tantivala (1987) reported 182
species of demersal fishes (in 66 families) in the
area from surveys using otterboard trawls at water
depths less than 90 m. Deep-sea fishes of the
Andaman Sea were poorly known. The results of
surveys during 1975 and 1976 on fishes inhabiting
the continental shelf within the depth range of 200–
550 m was reported by Pokapunt et al. (1983),
with taxonomic accounts of 31 species in 20
families. Under the Scientific Cooperation
Programme between Denmark and Thailand during
1996–2000, the Biodiversity of the Andaman Sea
Shelf (BIOSHELF) Project was undertaken by the
staff of the PMBC together with their Danish
counterparts. The benthic fauna, including fishes,
was sampled at depths of 40–900 m. Preliminary
results revealed a total of 55 families in 20 orders
of fishes, most of which were deep-water fishes
that have not yet been identified (Aungtonya et
34
Phuket mar. biol. Cent. Res. Bull.
al., 2000). Under the JSPS project, a
comprehensive guide book entitled “Fishes of
Andaman Sea, West Coast of Southern Thailand”
covered 778 species in 106 families of fishes
(Kimura et al., 2009).
The Reference Collection of the PMBC is
one of the major sources of information on marine
biodiversity of the Andaman Sea coast of Thailand.
The collection of fishes, in particular, was largely
established as a result of the FAO/DANIDA
Seminar on Fish Taxonomy in Southeast Asia held
at the PMBC in 1972. Apart from the Andaman
Sea fishes, a number of specimens obtained from
the Gulf of Thailand, as well as from neighboring
countries (i.e., India, Malaysia, Singapore,
Indonesia, Philippines, and Hong Kong) were
brought in during the seminar and subsequently
deposited in the PMBC Reference Collection. Some
deposited material of fishes from Australia was
donated by the Australian Museum under a past
exchange programme. One lot of fishes collected
from Myanmar during the first joint Myanmar-Thai
Fishery Exploratory Survey in 1990 was also
deposited in the Reference Collection
(Nateewathana et al., 1983). A revised checklist
of fishes in the PMBC Reference Collection was
published by Nateewathana et al. (1983),
comprising a total of 629 species in 119 families.
The collection has gradually increased following a
number of subsequent researches (e.g.,
Sirimontraporn et al., 1993; Satapoomin and
Poovachiranon, 1997; Satapoomin, 1999, 2002b)
and intensive sampling programmes (e.g., a
collaborative project between ROM and PMBC in
1993 and the PMBC’s past and ongoing projects,
namely Coral Reef Management Project (1995–
1999), Seagrass Resources Management and
Dugong Conservation Project (1997–2000), and
Assessment of Coastal Marine Resources Project
(since 2004)). According to the recent PMBC data
repository
(http://pmbc.go.th/pmbc-rc/
Taxonomy.php/), the most reliable records of fishes
(excluding
doubtful
records
and/or
misidentifications) deposited in the Reference
Collection total at least 1,200 species belonging to
170 families. The Andaman Sea fishes, in particular,
are well represented with a total number of 1,093
species, comprising 1, 39, and 1,053 species in
Myxini, Chondrichthyes, and Actinopterygii,
respectively.
2. Faunal composition and zoogeography
The present checklist contains 1,746
species, representing 696 genera, 198 families, 40
orders, and 3 classes, of marine and coastal fishes
from the Andaman Sea coast of Thailand (Appendix
A). The figure is relatively high as compared to
those reported from several other localities in the
Indian Ocean. For example, 533 species were
reported from the Cocos (Keeling) Islands (Allen
and Smith-Vaniz, 1994), 603 from the Laccadive
Archipelago (Jones and Kumaran, 1980), 622 from
Christmas Island (Hobbs et al., 2010), 784 from
the Chagos Archipelago (Winterbottom and
Anderson, 1999), 880 from Seychelles and
adjacent areas (Smith and Smith, 1969), 885 from
the Reunion Island (Letourneur et al., 2004), 930
species from Oman (Randall, 1995), 1,078 from
the Red Sea (Golani and Bogorodsky, 2010), 1,090
from the Maldives (Anderson et al., 1998), and
1,800 (as an estimate) from southern Africa
(Heemstra and Heemstra, 2004). There may be
the real differences in fish diversity among those
localities, but it should be noted that the number
of species recorded probably reflect biases or
differences in any or all of the following aspects:
sampling effort, habitat/depth representation, and
geographical coverage.
The 10 most speciose families (Gobiidae,
Labridae, Pomacentridae, Serranidae, Apogonidae,
Blenniidae, Carangidae, Scorpaenidae, Lutjanidae
and Chaetodontidae) collectively account for about
42% of the total fish fauna. The cumulative
percentage contribution when the top 20 families
are taken into account (Table 1) is over half the
total (56%), suggesting that the fauna is not
predominated by only a few families but by a set
of several families. This seems to reflect the
diversity of habitats encompassed by the current
faunal list. Each habitat usually contains a unique
composition of the predominant fish families,
except for the Gobiidae, which is the most diverse
group in nearly all coastal habitats (e.g., coral reefs,
seagrass beds and mangroves – Satapoomin, 2006,
2007; Fish Team of the Trang Project, 2002). The
current checklist, which includes species from
35
The fishes of southwestern Thailand
Table 1. Twenty most speciose families of fishes of southwestern Thailand, the Andaman Sea.
Family
Number of genera
Number of species
% of total fish fauna
Gobiidae
Labridae
Pomacentridae
Serranidae
Apogonidae
Blenniidae
Carangidae
Scorpaenidae
Lutjanidae
Chaetodontidae
Acanthuridae
Muraenidae
Syngnathidae
Carcharhinidae
Nemipteridae
Engraulidae
Lethrinidae
Leiognathidae
Tetraodontidae
Scaridae
62
30
18
17
14
21
20
23
10
7
5
7
11
7
3
5
5
8
8
6
227
78
71
61
60
52
52
49
39
37
32
32
26
25
25
24
22
21
21
20
13.0
4.5
4.1
3.5
3.4
3.0
3.0
2.8
2.2
2.1
1.8
1.8
1.5
1.4
1.4
1.4
1.3
1.2
1.2
1.1
Subtotal
287
974
55.8
Other 176 families
409
772
44.2
Total
696
1,746
100.0
various habitats, thus evens out the significance
contribution of certain predominant families in the
analysis. The relative abundance of these families
is similar, although with variable ranking, to those
in studies of inshore fishes at many other IndoPacific localities, for example, Reunion Island
(Letourneur et al., 2004), Maldive Islands (Randall
and Anderson, 1993), Chagos Archipelago
(Winterbottom and Anderson, 1999), northwestern
Australia (Allen, 1993), Calamianes Islands,
Palawan Province (Allen, 2000), southern Taiwan
(Shao et al., 2008), Yaku-shima Island, southern
Japan (Motomura et al., 2010), and Mariana
Islands (Myers and Donaldson, 2003). The
Gobiidae, in particular, was usually the leading
family when it was adequately sampled. Other less
diverse families include 25 families with a range
of species numbers of 10–19 species, 27 families
with 5–9 species, 59 families with 2–4 species,
and the remaining 66 families each with single
species.
With respect to habitat association, the fish
fauna consists overwhelmingly of reef fishes,
which comprise about 56% (983 species) of the
total species. The coastal species from mangroves
and seagrass beds represent about 17% (296
species) and 14% (246 species) of the total fauna,
respectively. The other major component, which
is loosely categorized to accommodate fishes
inhabiting a variety of offshore habitats (spanning
pelagic to benthic zones at various depths),
comprises about 55% (971 species) of the total
36
Phuket mar. biol. Cent. Res. Bull.
fauna. The figures accounting for the proportion
of species shared between habitats vary
considerably and are relatively low, i.e., 42%
between mangroves and seagrass beds, 20.5%
between offshore habitats and coral reefs, 16.8%
between offshore habitats and seagrass beds,
15.6% between offshore habitats and mangroves,
9.4% between coral reefs and seagrass beds, and
5.3% between coral reefs and mangroves. The
species shared among the 4 biotopes account for
only 2.3% of the total species recorded. Species
shared between or among habitats, however,
comprise a number of species which are strongly
dependent on more than a single habitat type, and
often exhibit a segregated habitat-use pattern with
respect to their different life-history stages (e.g.,
Mumby et al., 2004; Shibuno et al., 2008). This
seems to suggest an inter-system connection in
fish populations. Because unique assemblages of
species are restricted to each particular habitat
type, while others depend upon the existence of
multiple habitat-types through connectivity, special
attention must be paid to holistic approaches of
multi-ecosystems management and conservation
if the overall diversity of the fishes is to be
maintained. At present, there is growing concern
over the science of connectivity because
understanding of the linkage among ecosystems
is considered as a key to the success of
conservation and management of marine and
coastal areas, particularly when emphasizing the
design of marine protected areas (MPAs) and MPA
network (Botsford et al., 2009; Sale et al., 2010).
The zoogeographic analysis of species
with adequately known distribution ranges is
presented in Table 2. The analysis reveals that the
fish fauna consists primarily of species which are
widely distributed either in the Indo-Pacific or Indowest Pacific (50.7%). Approximately 17.1% of the
fauna are found within portions of the western
Pacific and northeastern Indian Ocean (i.e., NIO–
WCP, NIO–WP, NIO–IA, NIO–IM, EIO–WCP,
EIO–WP, EIO–IA, and EIO–IM). Similarly 15.5%
of species have a distribution range in the western
Pacific or portions of this region (i.e., WCP, WP,
IA, and IM). This clearly confirms that eastern
most boundary of the Indian Ocean including the
Andaman Sea overlaps the biogeographic region
of the West Pacific. About 12% of the fauna has a
primarily Indian Ocean range, with lesser values
(Table 3) for either the fauna confined to the eastern
extremity of the Indian Ocean (about 1%) or the
fauna restricted only to the Andaman Sea province
(1.7%). The remaining species (4.9%) have panoceanic ranges, including circumtropical and
cosmopolitan distributions. The presence of fauna
having both an Indian Ocean distribution range and
also a western Pacific distribution range in the area
can be considered as compelling evidence of a
distribution confluence of both Indian Ocean and
Pacific Ocean faunas in the region. The boundary
where sympatry of the eastern (= Pacific Ocean)
and western (= Indian Ocean) populations is most
evident seems to encompass Christmas Island and
the Java–Bali region (see, Randall, 1998; Allen and
Adrim, 2003; Hobbs et al., 2010). The strong
influence of the Indian Ocean fauna in the area
can be further highlighted by considering the
number of sibling species that involve Indian and
Pacific Ocean species pairs, and some geographic
variants that contain different Indian and Pacific
Ocean colour morphs. While inconsistent
considerations among ichthyologists over
taxonomic status (i.e., variants, subspecies, or full
species) of the latter exist, concrete morphological
and/or genetic evidence is required to ascertain
their taxonomic status (Randall, 1998; Hastings
and Springer, 2009). As exemplified in Table 4,
which includes 68 sibling species-pairs (in 17
families) and 18 species (in 10 families) possessing
geographic variation in colour morphs, the
Andaman Sea fauna is predominantly represented
by the Indian Ocean forms.
Since establishment of the PMBC
Reference Collection about 25 years ago, a great
deal of knowledge has accumulated and well over
190 new species (see also Aungtonya et al., 2006)
have been described from the Andaman Sea. At
present, the Reference Collection maintains type
materials of 21 fish species, including 9 holotypes
and 27 lots of paratypes (Table 5). Reliable records
of Andaman Sea fishes currently deposited at the
Reference Collection account for about 63% of
the total fish fauna. The fish collection thus still
does not include a comprehensive representation
of the Andaman Sea fish fauna. In order to maintain
its significant role in supporting biodiversity
research of the ichthyofauna of this particular
37
The fishes of southwestern Thailand
region, the fish collection needs to be upgraded
by additional sampling so that the missing 27% of
species can be made available as voucher or
reference materials for future researchers. Also
worth mentioning here are the figures reflecting
biotope representation of the fishes catalogued in
the Reference Collection, which vary considerably,
i.e., 64% of total offshore fishes and 65% of the
reef-associated fishes, but 81% of the mangrove
fishes and 88% of the seagrass fishes. These
figures can form a basis for targeting future
sampling programs for the Reference Collection.
Small and cryptic reef fishes and particularly those
inhabiting the deep-reef habitats are in need of
extensive sampling. For fishes dwelling among
various offshore habitats, deep demersal,
benthopelagic, and mesopelagic fishes are among
the groups that have not been adequately sampled.
Deep sea exploratory research programs are
generally costly and particularly require specific
techniques, equipment as well as taxonomic
expertise. The establishment of such programs is
likely to require international collaborative efforts
and funding.
Although the present checklist provides
the most extensive species records for the area, it
should be considered as a provisional rather than
a complete faunal list. Also, there are a number of
identifications that still need verification through
examination of voucher specimens. Furthermore,
the sampling efforts for several fish groups and/
or in certain habitats remain deficient. The present
provisional checklist, however, can at least help
to draw the attention of ichthyologists who are
interested in fish fauna of the Andaman Sea to
where there are knowledge gaps. Also, it can serve
as a baseline for a comprehensive taxonomic
review and/or annotated checklist of the Andaman
Sea fishes. It is hoped that, joint research or
collaborative efforts will be possible in the near
future to accomplish such goals. While the current
limited effort and interest, especially with respect
to taxonomy, has slowed down the progress in
this research area, a concomitant obstacle
regarding decreasing personnel trained in
ichthyology in Thailand seems to be more critical
and needs urgent action.
Table 2. Zoogeographic analysis of the fish fauna of southwestern Thailand, the Andaman Sea.
Distribution
Indo-Pacific (IP)
Indo-West Pacific (IWP)
West central Pacific (WCP)
West Pacific (WP)
Indo-Australian Archipelago (IA)
Indo-Malayan Archipelago (IM)
Indian Ocean (IO)
Northern Indian Ocean (NIO)
Eastern Indian Ocean (EIO)
Northern Indian Ocean to West central Pacific (NIO–WCP)
Northern Indian Ocean to West Pacific (NIO–WP)
Northern Indian Ocean to Indo-Australian Archipelago (NIO–IA)
Northern Indian Ocean to Indo-Malayan Archipelago (NIO–IM)
Eastern Indian Ocean to West central Pacific (EIO–WCP)
Eastern Indian Ocean to West Pacific (EIO–WP)
Eastern Indian Ocean to Indo-Australian Archipelago (EIO–IA)
Eastern Indian Ocean Indo-Malayan Archipelago (EIO–IM)
Eastern extremity of Indian Ocean (EEIO)
Andaman Sea (AS)
Circumtropical (CIR)
Cosmopolitan (COS)
No. species
%
417
448
27
137
48
53
99
10
46
2
37
21
29
12
112
28
50
15
28
38
46
24.5
26.3
1.6
8.0
2.8
3.1
5.8
0.6
2.7
0.1
2.2
1.2
1.7
0.7
6.6
1.6
3.0
0.9
1.7
2.2
2.7
38
Phuket mar. biol. Cent. Res. Bull.
Table 3. List of fishes currently known to have restricted distribution in the Andaman Sea and adjacent
areas: A) Andaman Sea endemics, including the species ranges southward to western Sumatra; B) The
species found in Andaman Sea and some other localities along eastern edge of the eastern Indian Ocean
(= eastern extremity of Indian Ocean (EEIO))
Family: Species
A)Andaman Sea endemics
Myxinidae:
Eptatretus indrambaryai Wongratana, 1983
Proscylliidae:
Proscyllium magnificum Last & Vongpanich, 2004
Rajidae:
Anacanthobatis sp.1
Acropomatidae:
Acropoma argentistigma Okamoto & Ida, 2002
Aploactinidae:
Cocotropus echinatus (Cantor, 1850)
Apogonidae:
Apogon sp.1
Archamia ataenia Randall & Satapoomin, 1999
Blenniidae:
Blenniella leopardus (Fowler, 1904)
Meiacanthus urostigma Smith-Vaniz, Satapoomin & Allen, 2001
Bythitidae:
Alionematichthys phuketensis Møller & Schwarzhans, 2008
Gobiidae:
Callogobius andamanensis Menon & Chatterjee, 1974
Scartelaos cantoris (Day, 1871)
Trimma fucatum Winterbottom & Southcott, 2007
Trimma sanguinellus Winterbottom & Southcott, 2007
Haemulidae:
Hapalogenys merguiensis Iwatsuki, Satapoomin & Amaoka, 2000
Plectorhinchus macrospilus Satapoomin & Randall, 2000
Pomadasys andamanensis McKay & Satapoomin, 1994
Labridae:
Cirrhilabrus joanallenae Allen, 2000
Halichoeres kallochroma (Bleeker, 1853)
Monacanthidae:
Paraluteres sp.1
Ophichthidae:
Allips concolor McCosker, 1972
Phallostethidae:
Phenacostethus posthon Roberts, 1971
Plesiopidae:
Plesiops thysanopterus Mooi, 1995
Pomacentridae:
Chromis xouthos Allen & Erdmann, 2005
Pomacentrus polyspinus Allen, 1991
Serranidae:
Cephalopholis polyspila Randall & Satapoomin, 2000
Meganthias filiferus Randall & Heemstra, 2007
Distribution
Andaman Sea
Andaman Sea
Andaman Sea
Andaman Sea
Andaman Sea
West coast of Thailand
Andaman Sea–W. Sumatra
Andaman Sea–W. Sumatra
Andaman Sea–W. Sumatra
West coast of Thailand
Andaman Sea
Andaman Sea
West coast of Thailand
Andaman Sea–W. Sumatra
Andaman Sea
West coast of Thailand
West coast of Thailand
Andaman Sea–W. Sumatra
Andaman Sea–W. Sumatra
Andaman Sea
Andaman Sea
Andaman Sea
N. Sumatra (Andaman Sea)
Andaman Sea–W. Sumatra
West coast of Thailand
Andaman Sea–W. Sumatra
Andaman Sea
39
The fishes of southwestern Thailand
Table 3. (continued)
Family: Species
Soleidae:
Aseraggodes satapoomini Randall & Desoutter-Meniger, 2007
Sternoptychidae:
Polyipnus asper Harold, 1994
Tripterygiidae:
Helcogramma lacuna Williams & Howe, 2003
B) Limited to eastern extremity of Indian Ocean (EEIO)
Urolophidae:
Urolophus javanicus (Martens, 1864)
Blenniidae:
Ecsenius lubbocki Springer, 1988
Ecsenius paroculus Springer, 1988
Praealticus triangulus (Chapman, 1951)
Labridae:
Cirrhilabrus cyanopleura (Bleeker, 1851)
Malacanthidae:
Hoplolatilus luteus Allen & Kuiter, 1989
Opistognathidae:
Opistognathus cyanospilotus Smith-Vaniz, 2009
Polynemidae:
Filimanus perplexa Feltes, 1991
Pomacentridae:
Pomacentrus alleni Burgess, 1981
Pomacentrus azuremaculatus Allen, 1991
Pomacentrus xanthosternus Allen, 1991
Pseudochromidae:
Pseudochromis andamanensis Lubbock, 1980
Scaridae:
Chlorurus troschelii (Bleeker, 1853)
Scarus maculipinna Westneat, Satapoomin & Randall, 2007
Siganidae:
Siganus magnificus (Burgess, 1977)
Distribution
West coast of Thailand
Andaman Sea
West coast of Thailand
Andaman Sea, Java Sea (W. Java)
Andaman Sea, Bali
Andaman Sea., W. Sumatra, Java
Sea (Seribu Is.& N. Java)
Andaman Sea, Java Sea (N. Java)
Andaman Sea, W. Sumatra,
Christmas Is., Java Sea (Pulau Putri
& N. Java), Bali
Andaman Sea, Bali, Flores
Andaman Sea, Bali
Andaman Sea, W. Sumatra,
S. Java, Bali
Andaman Sea, Christmas Is.,
Java Sea (Sunda Strait &
Pulau Putri)
Andaman Sea, Java Sea (Seribu Is.)
Andaman Sea, W. Sumatra,
Java Sea (Seribu Is.), Bali
Andaman Sea, W. Sumatra,
Bali, Timor Sea (Ashmore Reef &
Cartier Is., N. Australia)
Andaman Sea, Java Sea (Seribu Is.)
Andaman Sea, W. Sumatra, Java
Sea (Pulau Putri)
Andaman Sea, W. Sumatra, Java
Sea (W. Java)
40
Phuket mar. biol. Cent. Res. Bull.
Table 4. Examples of Indian and Pacific Ocean species pairs and geographic variants that the Andaman
Sea fishes represented by the Indian Ocean forms.
A) Sibling species: Andaman Sea fish (vs. its Pacific sibling)
Acanthuridae
Acanthurus tristis (A. pyroferus)
Ctenochaetus truncatus (C. strigosus)
Naso elegans (N. lituratus)
Zebrasoma desjardinii (Z. veliferum)
Apogonidae
Archamia ataenia (A. zosterophora)
Blenniidae
Aspidontus tractus (A. taeniatus)
Atrosalarias fuscus (A. holomelas)
Blenniella cyanostigma (B. caudolineata)
Blenniella leopardus (B. interrupta)
Meiacanthus urostigma (M. ditrema)
Plagiotremus phenax (P. laudandus)
Chaetodontidae
Chaetodon andamanensis (C. plebeius)
Chaetodon triangulum (C. baronessa)
Chaetodon falcula (C. ulietensis)
Chaetodon gardineri (C. selene)
Chaetodon interruptus (C. unimaculatus)
Chaetodon madagaskariensis (C. mertensii)
Chaetodon melannotus (C. ocellicaudus)
Chaetodon trifasciatus (C. lunulatus)
Hemitaurichthys zoster (H. polylepis)
Heniochus pleurotaenia (H. varius)
Gobiidae
Amblygobius semicinctus (A. phalaena)
Trimma flammeum (T. macrophthalma)
Labridae
Anampses lineatus (A. melanurus)
Cirrhilabrus cyanopleura (C. lyukyuensis)
Coris cuvieri (C. gaimard)
Bodianus diana (B. dictynna)
Gomphosus caeruleus (G. varius)
Halichoeres cosmetus (H. claudia)
Halichoeres kallochroma (H. leucurus)
Halichoeres leucoxanthus (H. chrysus)
Halichoeres vrolokii (H. melanurus)
Halichoeres zeylonicus (H. hartzfeldii)
Paracheilinus mccoskeri (P. carpenteri)
Pseudojuloides kaleidos (P. cerasinus)
Mullidae
Parupeneus trifasciatus (P. crassilabris)
Pinguipedidae
Parapercis maculata (P. pulchella)
Pomacanthidae
Centropyge eibli (C. vroliki)
Genicanthus caudovittatus (G. melanospilos)
Pomacentridae
Amphiprion akallopisos (A. sandarcinos)
Amphiprion ephippium (A. melanopus)
Amphiprion sebae (A. polymnus)
Chromis dimidiata (C. iomelas)
Chromis opercularis (C. xanthura)
Dascyllus carneus (D. reticulatus)
Neopomacentrus sororius (N. azysron)
Pomacentrus similis (P. coelestis)
Pomacentrus xanthosternus (P. simsiang)
Ptereleotridae
Ptereleotris arabica (P. hanae)
Scaridae
Chlorurus capistratoides (C. japanensis)
Chlorurus rhakoura (C. oedema)
Chlorurus sordidus (C. spilurus)
Chlorurus strongylocephalus (C. microrhinos)
Chlorurus troschelii (C. bleekeri)
Hipposcarus harid (H. longiceps)
Scarus maculipinna (S. flavipectoralis)
Scarus russelii (S. schlegeli)
Scarus scaber (S. oviceps)
Scarus viridifucatus (S. spinus)
Scorpaenidae
Pterois miles (P. volitans)
Serranidae
Cephalopholis polyspila (C. microprion)
Pogonoperca ocellata (P. punctata)
Pseudanthias ignitus (P. dispar)
Rabualichthys stigmaticus (R. altipinnis)
Siganidae
Siganus puelloides (S. puellus)
Siganus stellatus (S. punctatus)
Syngnathidae
Doryramphus multiannulatus (D. pessuliferus)
Tetraodontidae
Canthigaster smithae (C. epilampra)
41
The fishes of southwestern Thailand
Table 4. (continued)
B) Geographic variants: Andaman Sea fish possessing Indian Ocean colour morph
Dasyatidae
*(4)
Dasyatis kuhlii
Acanthuridae
*(7)
Paracanthurus hepatus
Balistidae
*(5, 7)
Balistapus undulatus
Caesionidae
Caesio cuning
Labridae
*(9)
Epibulus insidiator
*(5, 7)
Halichoeres hortulanus
*(5)
Labroides dimidiatus
*(7)
Thalassoma jansenii
Lutjanidae
*(1, 7)
Lutjanus russelli
Nemipteridae
*(6, 7)
Scolopsis bilineatus
Scolopsis vosmeri
Pomacanthidae
*(7, 8)
Pomacanthus imperator
*(5, 7)
Pygoplites diacanthus
Pomacentridae
*(2)
Chrysiptera rollandi
*(7)
Neoglyphidodon nigroris
*(6)
Pomacentrus chrysurus
*(2, 8)
Pomacentrus pavo
Serranidae
*(3)
Cephalopholis urodeta
*Note: Recognition of allopatric colour morphs in certain species can be referred to (1) Allen and
Talbot, 1985; (2) Allen, 1991; (3) Randall and Heemstra, 1991; (4) Randall, 1995; (5) Randall, 1998;
(6) Satapoomin, 2000; (7) Kuiter and Tonozuka, 2001; (8) Lieske and Myers, 2001; and (9) Carlson et
al., 2008.
Table 5. List of fishes for which their type materials are deposited at the Reference Collection of the
Phuket Marine Biological Center. Number in bracket denotes the number of catalogued lots for paratypes.
Family: Species
Apogonidae:
Archamia ataenia Randall & Satapoomin, 1999
Blenniidae:
Meiacanthus urostigma Smith-Vaniz, Satapoomin & Allen, 2001
Gobiidae:
Cryptocentrus crocatus Wongratana, 1975
[valid as Myersina crocatus (Wongratana, 1975)]
Gladiogobius rex Shibukawa & Allen, 2007
Myersina adonis Shibukawa & Satapoomin, 2006
Periophthalmus walailakae Darumas & Tantichodok, 2002
Trimma fucatum Winterbottom & Southcott, 2007
Trimma sanguinellus Winterbottom & Southcott, 2007
Haemulidae:
Hapalogenys merguiensis Iwatsuki, Satapoomin & Amaoka, 2000
Plectorhinchus macrospilus Satapoomin & Randall, 2000
Pomadasys andamanensis McKay & Satapoomin, 1994
Nemipteridae:
Nemipterus aurora Russell, 1993
Nemipterus thosaporni Russell, 1991
Type materials
Paratypes (1)
Paratypes (2)
Holotype
Holotype, Paratypes (3)
Holotype, Paratypes (1)
Paratypes (2)
Paratypes (1)
Paratypes (1)
Paratypes (2)
Paratypes (1)
Holotype
Paratypes (1)
Paratypes (1)
42
Phuket mar. biol. Cent. Res. Bull.
Table 5. (continued)
Family: Species
Type materials
Opistognathidae:
Opistognathus variabilis Smith-Vaniz, 2009
Proscylliidae:
Proscyllium magnificum Last & Vongpanich, 2004
Pomacentridae:
Pomacentrus azuremaculatus Allen, 1991
Pomacentrus polyspinus Allen, 1991
Pomacentrus similis Allen, 1991
Scaridae:
Scarus maculipinna Westneat, Satapoomin & Randall, 2007
Serranidae:
Cephalopholis polyspila Randall & Satapoomin, 2000
Meganthias filiferus Randall & Heemstra, 2007
ACKNOWLEDGEMENTS
This paper was initially prepared as a
contribution to the concluding conference of the
10-year JSPS Multilateral Core University
Programme on “Coastal Marine Science” in Tokyo
during 26–29 October 2010, but I finally could
not make the trip. It was then presented as a plenary
talk at the seminar on “Biodiversity of Southern
Thailand in Focus: Status and Research Direction”
at Walailak University, Nakorn Sri Thammarat,
Thailand, during 10–12 November 2010. I am
grateful for the generous support made available
by the JSPS for me to be able to join several fish
workshops conducted over the years, which
ultimately gave me a wonderful opportunity to learn
more about the fish faunas of SE Asia. Special
Paratypes (2)
Holotype, Paratypes (3)
Holotype
Holotype
Paratypes (1)
Holotype, Paratypes (1)
Paratypes (4)
Holotype
thanks are due to Dr. Keiichi Matsuura [KM] of
the National Museum of Nature and Science,
Tokyo, for his considerable effort in leading the
fish diversity project under the JSPS multilateral
programme and encouraging me to write up this
paper. I am very grateful to Dr. Richard
Winterbottom [RW] of the Royal Ontario Museum
for sharing the research outcomes from the 1993
expedition to Phuket and inspiring my interest in
small cryptobenthic reef fishes. Both KM and RW
kindly gave their time for reviewing and improving
the first draft of this manuscript. Several past and
ongoing projects at PMBC which contribute to the
information and knowledge on the Andaman Sea
fish fauna have been continually supported by the
Thai government budget. I am also grateful to the
many staff members of the PMBC for their
assistance in the field.
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601 p.
Pokapunt, W. and M. Sinoda. 1978. Demersal fish resources in the Andaman Sea off the west coast of
Thailand for 1965–1975. Exploratory Fishing Division, Technical Paper No. 1. 28 p. (in
Thai)
Pokapunt, W. and J. Tantivala. 1987. Observations on demersal fish resources and trash fish
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Pokapunt, W., J. Tantivala, A. Munprasit and P. Samrankitdamrong. 1983. Some deep-sea fishes in the
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pls.
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Soc. 31(1): 1–7.
Manuscript received: 10 January 2011
Accepted: 1 August 2011
47
The fishes of southwestern Thailand
Appendix A
Checklist of marine fishes of southwestern
Thailand, the Andaman Sea
Abbreviations:
1. Habitat types – Cr (coral reefs); Mg (mangroves); Sg
(seagrass beds); OShr (other coastal and offshore areas)
2. Distribution categories – IP (Indo-Pacific); IWP (IndoWest Pacific); WCP (west central Pacific); WP (west
Pacific); IA (Indo-Australian Archipelago); IM (IndoMalayan Archipelago); IO (Indian Ocean); NIO (northern
Indian Ocean); EIO (eastern Indian Ocean); EEIO (eastern
extremity of Indian Ocean); AS (Andaman Sea); CIR
(circumtropical); COS (cosmopolitan) [see methods
section for definitions]
3. Voucher specimens – PMBC (denotes availability of
voucher specimens at the Reference Collection of the
Phuket Marine Biological Center)
CLASS MYXINI
Order Myxiniformes
Myxinidae
Eptatretus indrambaryai Wongratana, 1983: OShr; [AS];
PMBC
CLASS CHONDRICHTHYES
Order Chimaeriformes
Chimaeridae
Hydrolagus sp.: OShr; PMBC
Order Orectolobiformes
Parascylliidae
Parascyllium sp.: OShr; PMBC
Hemiscylliidae
Chiloscyllium griseum Müller & Henle, 1838: OShr; [NIOIM]; PMBC
Chiloscyllium hasselti Bleeker, 1852: Sg, OShr; [IM];
PMBC
Chiloscyllium indicum (Gmelin, 1788): OShr; [IWP];
PMBC
Chiloscyllium plagiosum (Bennett, 1830): OShr; [IWP]
Chiloscyllium punctatum Müller & Henle, 1838: Cr, Sg,
OShr; [EIO-WP]
Stegostomatidae
Stegostoma fasciatum (Hermann, 1793): Cr, OShr; [IP];
PMBC
Ginglymostomatidae
Nebrius ferrugineus (Lesson, 1830): Cr, OShr; [IWP]
Rhincodontidae
Rhincodon typus Smith, 1828: Cr, OShr; [COS]
Order Lamniformes
Alopiidae
Alopias pelagicus Nakamura, 1935: OShr; [CIR]
Alopias superciliosus (Lowe, 1840): OShr; [CIR]
Alopias vulpinus (Bonnaterre, 1788): OShr; [COS]
Lamnidae
Isurus oxyrinchus Rafinesque, 1810: OShr; [COS]
Order Carcharhiniformes
Scyliorhinidae
Atelomycterus marmoratus (Bennett, 1830): OShr; [IWP]
Cephaloscyllium umbratile Jordan & Fowler, 1903: OShr;
[WP]; PMBC
Halaelurus buergeri (Müller & Henle, 1838): OShr; [WP]
Halaelurus hispidus (Alcock, 1891): OShr; [EIO-WP];
PMBC
Proscylliidae
Proscyllium magnificum Last & Vongpanich, 2004: OShr;
[AS]; PMBC
Triakidae
Iago omanensis (Norman, 1939): OShr; [IO]; PMBC
Mustelus manazo Bleeker, 1854: OShr; [IWP]; PMBC
Mustelus mosis Hemprich & Ehrenberg, 1899: OShr; [IO];
PMBC
Hemigaleidae
Chaenogaleus macrostoma (Bleeker, 1852): OShr; [IWP]
Hemigaleus microstoma Bleeker, 1852: OShr; [EIO-WP];
PMBC
Hemipristis elongatus (Klunzinger, 1871): OShr; [IWP];
PMBC
Carcharhinidae
Carcharhinus albimarginatus (Rüppell, 1837): Cr, OShr;
[IP]
Carcharhinus altimus (Springer, 1950): OShr; [COS]
Carcharhinus amblyrhynchoides (Whitley, 1934): OShr;
[IWP]
Carcharhinus amblyrhynchos (Bleeker, 1856): OShr; [IP]
Carcharhinus amboinensis (Müller & Henle, 1839): OShr;
[IWP]
Carcharhinus brachyurus (Günther, 1870): OShr; [COS]
Carcharhinus brevipinna (Müller & Henle, 1839): OShr;
[CIR]
Carcharhinus dussumieri (Valenciennes, 1839): OShr;
[NIO-WP]
Carcharhinus falciformis (Müller & Henle, 1839): OShr;
[CIR]
Carcharhinus leucas (Valenciennes, 1839): OShr; [CIR]
Carcharhinus limbatus (Valenciennes, 1839): OShr; [COS]
Carcharhinus longimanus (Poey, 1861): OShr; [COS]
Carcharhinus macloti (Müller & Henle, 1839): OShr; [IWP];
PMBC
Carcharhinus melanopterus (Quoy & Gaimard, 1824): Cr,
OShr; [IP]; PMBC
Carcharhinus obscurus (LeSueur, 1818): OShr; [COS]
Carcharhinus plumbeus (Nardo, 1827): OShr; [CIR];
PMBC
Carcharhinus sealei (Pietschmann, 1916): OShr; [IWP]
Carcharhinus sorrah (Valenciennes, 1839): OShr; [IWP]
Galeocerdo cuvier (Péron & LeSueur, 1822): OShr; [CIR]
Loxodon macrorhinus Müller & Henle, 1839: OShr;
[IWP]
Negaprion acutidens (Rüppell, 1837): OShr; [IP]
Rhizoprionodon acutus (Rüppell, 1837): OShr; [CIR];
PMBC
Rhizoprionodon oligolinx Springer, 1964: OShr; [IWP]
48
Phuket mar. biol. Cent. Res. Bull.
Scoliodon laticuadus Müller & Henle, 1839: OShr; [IWP]
Triaenodon obesus (Rüppell, 1835): Cr, OShr; [IP]
Sphyrnidae
Eusphyra blochii (Cuvier, 1817): OShr; [NIO-WP]; PMBC
Sphyrna lewini (Griffith & Smith, 1834): OShr; [COS];
PMBC
Sphyrna mokarran (Rüppell, 1837): OShr; [CIR]
Sphyrna zygaena (Linnaeus, 1758): OShr; [CIR]; PMBC
Order Hexanchiformes
Hexanchidae
Heptranchias perlo (Bonnaterre, 1788): OShr; [COS]
Order Squaliformes
Squalidae
Etmopterus brachyurus Smith & Radcliffe, 1912: OShr;
[WP]
Squalus acanthias Linnaeus, 1758: OShr; [COS]; PMBC
Squalus mitzukurii Jordan & Snyder, 1903: OShr; [CIR];
PMBC
Order Squantiformes
Squatinidae
Squatina sp.1: OShr
Order Torpediniformes
Narcinidae
Narcine brunnea Annandale, 1909: OShr; [NIO-IM];
PMBC
Narcine indica Henle, 1834: OShr; [WP]
Narcine maculata (Shaw, 1804): OShr; [WP]; PMBC
Narcine prodosalis Bessednov, 1966: OShr; [EIO-WP]
Narcine timlei (Bloch & Schneider, 1801): OShr; [EIOWP]
Narke dipterygia (Bloch & Schneider, 1801): OShr; [NIOWP]
Temera hardwickii Gray, 1831: OShr; [IM]; PMBC
Order Pristiformes
Pristidae
Pristis microdon Latham, 1794: OShr; [CIR]; PMBC
Order Rajiformes
Rhinidae
Rhina ancylostoma (Bloch & Schneider, 1801): Cr, OShr;
[IP]; PMBC
Rhynchobatidae
Rhynchobatus palpebratus Compagno & Last, 2008: Cr,
OShr; [IA (presently known from west coast of Thailand
and northern Australia)]
Rhinobatidae
Rhinobatos granulatus Cuvier, 1829: OShr; [NIO-IM];
PMBC
Rhinobatos thouin (Anonymous in Lacepède, 1798): OShr;
[IWP]
Rhinobatos typus Bennett, 1830: OShr; [EIO-WP]
Rajidae
Anacanthobatis sp.1 : OShr, PMBC
Raja sp.: OShr
Order Myliobatiformes
Plesiobatidae
Plesiobatis daviesi Wallace, 1967: OShr; [IP]; PMBC
Urolophidae
Urolophus javanicus (Martens, 1864): OShr; [EEIO]
Dasyatidae
Dasyatis kuhlii (Müller & Henle, 1841): Cr, OShr; [IWP];
PMBC
Dasyatis zugei (Müller & Henle, 1841): OShr; [EIO-WP]
Himantura fai Jordan & Seale, 1906: OShr; [IWP]
Himantura gerrardi (Gray, 1851): Sg, OShr; [IP]; PMBC
Himantura imbricata (Bloch & Schneider, 1801): Mg, Sg,
OShr; [IWP]; PMBC
Himantura jenkinsii (Annadale, 1909): Cr, OShr; [EIO]
Himantura undulata (Bleeker, 1852): OShr; [EIO-WP]
Himantura walga (Müller & Henle, 1841): Sg, OShr; [EIOIM]; PMBC
Pastinachus sephen (Forsskål, 1775): Cr, OShr; [IWP]
Taeniura lymma (Forsskål, 1775): Cr; [IWP]; PMBC
Taeniura meyeni Müller & Henle, 1841: Cr; [IP]
Urogymnosus asperrimus (Bloch & Schneider, 1801): Cr;
[IP]
Gymnuridae
Gymnura poecilura (Shaw, 1804): Mg, OShr; [IP]; PMBC
Myliobatidae
Aetobatus narinari (Euphrasen, 1790): Cr, OShr; [COS]
Aetomylaeus maculatus (Gray, 1832): OShr; [EIO-WP]
Aetomylaeus milvus (Valenciennes, 1841): OShr; [IWP]
Aetomylaeus nichofii (Bloch & Schneider, 1801): OShr;
[IWP]; PMBC
Aetomylaeus vespertilio (Bleeker, 1852): OShr; [IWP]
Manta birostris (Donndorff, 1798): Cr, OShr; [CIR]
Mobula kuhlii (Valenciennes, in Müller & Henle, 1841):
OShr; [IWP]
Rhinoptera javanica Müller & Henle, 1841: OShr; [IWP];
PMBC
Rhinoptera sewelli Misra, 1946: OShr; [EIO]
CLASS ACTINOPTERYGII
Order Elopiformes
Elopidae
Elops hawaiiensis Regan, 1909: Mg, Sg, OShr; [WCP];
PMBC
Megalopidae
Megalops cyprinoides (Broussonet, 1782): Mg, OShr; [IP];
PMBC
Order Albuliformes
Albulidae
Albula glossodonta (Forsskål, 1775): OShr; [IP]
Albula oligolepis Iwatsuki & Hidaka, 2008: OShr; [IO];
PMBC
Notacanthidae
Notacanthus sp.: OShr; PMBC
Order Anguilliformes
Anguillidae
Anguilla bicolor bicolor McClelland, 1844: Mg; [IWP];
PMBC
Anguilla marmorata Quoy & Gaimard, 1824: OShr; [IP];
PMBC
Chlopsidae
49
The fishes of southwestern Thailand
Kaupichthys sp.: Cr
Muraenidae
Echidna delicatula (Kaup, 1856): OShr; [EIO-WP]; PMBC
Echidna nebulosa (Ahl, 1789): Cr; [IP]; PMBC
Echidna polyzona (Richardson, 1844): Cr; [IP]; PMBC
Enchelycore bayeri (Schultz, 1953): Cr; [IP]
Gymnomuraena zebra (Shaw, 1797): Cr; [IP]; PMBC
Gymnothorax annulatus (Smith & Bohlke, 1997): OShr;
[IM]
Gymnothorax buroensis (Bleeker, 1857): Cr; [IP]
Gymnothorax chilospilus Bleeker, 1865: Cr; [IP]; PMBC
Gymnothorax favagineus (Bloch & Schneider, 1801): Cr;
[IWP]; PMBC
Gymnothorax fimbriatus (Bennett, 1832): Cr; [IP]; PMBC
Gymnothorax flavimarginatus (Rüppell, 1828): Cr; [IP]
Gymnothorax javanicus (Bleeker, 1859): Cr; [IP]; PMBC
Gymnothorax margaritophorus Bleeker, 1864: Cr; [IP]
Gymnothorax melatremus Schultz, 1953: Cr; [IP]
Gymnothorax meleagris (Shaw & Nodder, 1795): Cr; [IP]
Gymnothorax moluccensis (Bleeker, 1864): Cr; [IWP]
Gymnothorax monochrous (Bleeker, 1856): Cr; [IP]
Gymnothorax permistus (Smith, 1962): Cr; [IO]; PMBC
Gymnothorax pindae Smith, 1962: Cr; [IP]; PMBC
Gymnothorax pseudoherrei Böhlke, 2000: Cr; [IWP]
Gymnothorax pseudothyrsoideus (Bleeker, 1852): Cr;
[NIO-WP]; PMBC
Gymnothorax reticularis Bloch, 1795: OShr; [IO]; PMBC
Gymnothorax richardsoni (Bleeker, 1852): Cr; [IWP];
PMBC
Gymnothorax rueppelliae (Mcclelland, 1845): Cr; [IP]
Gymnothorax thyrsoideus (Richardson, 1844): Cr, OShr;
[EIO-WCP]; PMBC
Gymnothorax undulatus (Lacepède, 1803): Cr; [IP]; PMBC
Gymnothorax zonipectis Seale, 1906: Cr; [IP]; PMBC
Rhinomuraena quaesita Garman, 1888: Cr; [IWP]
Scuticaria tigrina (Lesson, 1828): Cr; [IP]
Uropterygius concolor Rüppell, 1838: Cr; [IP]; PMBC
Uropterygius inornatus Gosline, 1958: Cr; [WCP]
Uropterygius xanthopterus Bleeker, 1859: Cr; [IP]
Ophichthidae
Allips concolor McCosker, 1972: Cr, Mg; [AS]
Apterichthus sp.: OShr; PMBC
Brachysomophis crocodilinus (Bennet, 1833): Cr, OShr;
[IP]
Echelus uropterus (Temminck & Schlegel, 1846): OShr;
[IWP]; PMBC
Muraenichthys gymnopterus (Bleeker, 1853): OShr; [EIOWP]
Myrichthys colubrinus (Boddaert, 1781): Cr; [IP]; PMBC
Neenchelys sp.: OShr; PMBC
Ophichthus evermanni Jordan & Richardson, 1909: OShr;
[WP]
Ophichthus sp.: OShr; PMBC
Pisonodophis cancrivorus (Richardson, 1844): Mg, Sg,
OShr; [IWP]; PMBC
Scolecenchelys gymnota (Bleeker, 1854): Sg; [IWP]
Scolecenchelys macroptera (Bleeker, 1857): Sg, OShr;
[IWP]; PMBC
Xyrias revulsus Jordan & Snyder, 1901: OShr; [IWP]
Colocongridae
Coloconger scholesi Chan, 1967: OShr; [IWP]; PMBC
Coloconger sp.: OShr; PMBC
Muraenesocidae
Congresox talabon (Cuvier, 1829): OShr; [EIO-IM];
PMBC
Congresox talabonoides (Bleeker, 1853): OShr; [IWP];
PMBC
Muraenosox bagio (Hamilton, 1822): OShr; [IWP]
Muraenosox cinereus (Forsskål, 1775): OShr; [IWP];
PMBC
Congridae
Ariosoma anago (Temminck & Schlegel, 1846): OShr;
[IWP]; PMBC
Bathycongrus guttulatus (Günther, 1887): OShr; [IP]
Bathyuroconger sp.: OShr; PMBC
Conger cinereus Rüppell, 1828: OShr; [IP]; PMBC
Gnathophis sp.: OShr; PMBC
Gorgasia preclara Böhlke & Randall, 1981: Cr; [EIO-IA]
Heteroconger hassi (Klausewitz & Eibl-Eibesfeldt, 1959):
Cr; [IP]
Rhynchoconger squaliceps (Alcock, 1894): OShr; [EIO]
Uroconger lepturus (Richardson, 1845): OShr; [IWP];
PMBC
Nettastomatidae
Nettastoma sp.: OShr
Order Clupeiformes
Pristigasteridae
Ilisha elongata (Bennett, 1830): OShr; [EIO-IM]; PMBC
Ilisha kampeni (Weber & Beaufort, 1913): Sg, OShr; [EIOIM]
Ilisha megaloptera (Swainson, 1839): Mg, OShr; [EIOIM]; PMBC
Ilisha melastoma (Schneider, 1801): Mg, OShr; [EIO-WP];
PMBC
Ilisha sirishai Seshagiri Rao, 1975: OShr; [NIO-IM]
Opisthopterus tardoore (Cuvier, 1829): Mg, OShr; [NIOIM]; PMBC
Pellona ditchela Valenciennes, 1847: Mg, OShr; [IWP];
PMBC
Raconda russeliana Gray, 1831: OShr; [EIO-IM]
Engraulidae
Coilia dussumieri Valenciennes, 1848: OShr; [EIO-IM];
PMBC
Coilia macrognathos Bleeker, 1852: OShr; [IM]
Coilia neglecta Whitehead, 1968: OShr; [EIO-IM]
Enchasicholina punctifer Fowler, 1938: OShr; [IP]
Encrasicholina devisi (Whitley, 1940): OShr; [IWP]; PMBC
Encrasicholina heteroloba (Rüppell, 1837): OShr; [IWP];
PMBC
Setipinna breviceps (Cantor, 1850): OShr; [EIO-IM]
Setipinna taty (Valenciennes, 1848): Mg, OShr; [EIO-IM]
Setipinna tenuifilis (Valenciennes, 1848): OShr; [EIO-WP]
50
Phuket mar. biol. Cent. Res. Bull.
Stolephorus andhraensis Babu Rao, 1966: OShr; [EIOIA]; PMBC
Stolephorus baganensis Hardenberg, 1933: Mg, OShr;
[EIO-IM]
Stolephorus chinensis (Günther, 1880): Mg, OShr; [IM]
Stolephorus commersonii Lacepède, 1803: Mg, Sg, OShr;
[IWP]; PMBC
Stolephorus dubiosus Wongratana, 1983: OShr; [EIO-IM]
Stolephorus indicus (van Hasselt, 1823): Mg, Sg, OShr;
[IWP]; PMBC
Stolephorus insularis Hardenberg, 1933: Mg, OShr; [NIOWP]
Stolephorus waitei Jordan & Seale, 1926: OShr; [EIO-IA];
PMBC
Thryssa baelama (Forster, 1775): OShr; [IWP]
Thryssa dussumieiri (Valenciennes, 1848): OShr; [NIOIM]
Thryssa hamiltonii (Gray, 1835): Mg, Sg, OShr; [NIOIA]; PMBC
Thryssa kammalensis (Bleeker, 1849): Mg, OShr; [IM]
Thryssa mystax (Schneider, 1801): OShr; [NIO-IM];
PMBC
Thryssa setirostris (Broussonet, 1782): Mg, Sg, OShr;
[IWP]; PMBC
Thryssa spinidens (Jordan & Seale, 1925): OShr; [EIO]
Chirocentridae
Chirocentrus dorab (Forsskål, 1775): Mg, Sg, OShr; [IWP];
PMBC
Chirocentrus nudus Swainson, 1839: OShr; [IWP]
Clupeidae
Amblygaster clupeoides Bleeker, 1849: OShr; [EIO-WP]
Amblygaster leiogaster (Valenciennes, 1847): OShr; [IWP]
Amblygaster sirm (Walbaum, 1792): OShr; [IWP]; PMBC
Anodontostoma chacunda (Hamilton, 1822): Mg, Sg, OShr;
[NIO-WP]; PMBC
Anodontostoma thailandiae Wongratana, 1983: OShr; [EIOIM]
Dussumieria acuta Valenciennes, 1847: OShr; [NIO-IM];
PMBC
Dussumieria elopsoides Bleeker, 1849: Mg, OShr; [IWP]
Escualosa thoracata (Valenciennes, 1847): Mg, Sg, OShr;
[NIO-IA]; PMBC
Herklotsichthys quadrimaculatus (Rüppell, 1837): Cr,
OShr; [IP]; PMBC
Hilsa kelee (Cuvier, 1829): Mg, Sg, OShr; [IWP]
Nematalosa galatheae Nelson & Rothman, 1973: OShr;
[EIO-IM]
Nematalosa nasus (Bloch, 1795): OShr; [EIO-WP]
Sardinella albella (Valenciennes, 1847): Mg, Sg, OShr;
[IWP]; PMBC
Sardinella fimbriata (Valenciennes, 1847): OShr; [EIO-IA];
PMBC
Sardinella gibbosa (Bleeker, 1849): OShr; [IWP]; PMBC
Sardinella lemura Bleeker, 1853: OShr; [WP]
Spratelloides delicatulus (Bennett, 1831): Cr, OShr; [IP];
PMBC
Spratelloides gracilis (Temminck & Schlegel, 1846): Cr,
Mg, Sg, OShr; [IP]
Tenualosa reevesii (Richardson, 1846): OShr; [WP]
Tenualosa toli (Valenciennes, 1847): OShr; [NIO-IM];
PMBC
Order Gonorynchiformes
Chanidae
Chanos chanos (Forsskål, 1775): Cr, Mg, OShr; [IP]
Order Siluriformes
Plotosidae
Plotosus canius Hamilton, 1822: Mg, Sg, OShr; [EIO-IA];
PMBC
Plotosus lineatus (Thunberg, 1787): Cr, Mg, Sg, OShr;
[IWP]; PMBC
Ariidae
Arius arius (Hamilton, 1822): OShr; [EIO-IM]; PMBC
Arius caelatus Valenciennes, 1840: Mg; [EIO-IM]
Arius gagora (Hamilton, 1822): OShr; [EIO]
Arius oetik Bleeker, 1846: OShr; [WP]
Arius venosus Valenciennes, 1840: OShr; [EIO-IM]
Nemapteryx nenga (Hamilton, 1822): OShr; [EIO-IM]
Nemuta thalassinus (Rüppell, 1837): OShr; [IWP]; PMBC
Osteogeneiosus militaris (Linnaeus, 1758): OShr; [EIOIM]
Plicofollis nella Bleeker, 1846: OShr; [WP]
Bagridae
Mystus gulio (Hamilton, 1822): Mg; [EIO-IM]; PMBC
Order Argentiniformes
Alepocephalidae
Alepocephalus bicolor Alcock, 1891: OShr; [IWP]; PMBC
Order Stomiiformes
Sternoptychidae
Argyropelecus affinis Garman, 1899: OShr; [CIR]; PMBC
Argyropelecus sladeni Regan, 1908: OShr; [CIR]; PMBC
Polyipnus asper Harold, 1994: OShr; [AS]; PMBC
Sternoptyx obscura Garman, 1899: OShr; [IWP]; PMBC
Stomiidae
Astronesthes lucifer Gilbert, 1905: OShr; [IP]
Chauliodus sloanei Bloch & Schneider, 1801: OShr; [COS]
Eustomias sp.: OShr; PMBC
Order Ateleopodiformes
Ateleopodidae
Ateleopus indicus Alcock, 1891: OShr; [EIO]; PMBC
Order Aulopiformes
Paraulopidae
Paraulopus brevirostris (Fourmanoir, 1981): OShr; [WP]
Synodontidae
Harpadon nehereus (Hamilton, 1822): OShr; [IWP]
Saurida elongata (Temminck & Schlegel, 1846): OShr;
[WP]; PMBC
Saurida gracilis (Quoy & Gaimard, 1824): Cr, OShr; [IP];
PMBC
Saurida longimanus Norman, 1939: OShr; [NIO]; PMBC
Saurida micropectoralis Shindo & Yamada, 1972: Mg,
OShr; [WP]; PMBC
Saurida nebulosa Valenciennes, 1849: Cr, Mg, Sg, OShr;
[IP]; PMBC
51
The fishes of southwestern Thailand
Saurida tumbil (Bloch, 1795): OShr; [IWP]; PMBC
Saurida undosquamis (Richardson, 1848): Cr, OShr;
[IWP]; PMBC
Synodus dermatogenys Fowler, 1912: Cr, OShr; [IP];
PMBC
Synodus hoshinonis Tanaka, 1917: OShr; [IWP]; PMBC
Synodus indicus (Day, 1873): OShr; [IWP]; PMBC
Synodus jaculum Russell & Cressey, 1979: Cr; [IP]; PMBC
Synodus rubromarmoratus Russell & Cressey, 1979: Cr;
[WP]
Synodus variegatus (Lacepède, 1801): Cr, OShr; [IP];
PMBC
Trachinocephalus myops (Bloch & Schneider, 1801): Mg,
Sg, OShr; [CIR]; PMBC
Chlorophthalmidae
Chlorophthalmus corniger Alcock, 1894: OShr; [EIO-WP]
Alepisauridae
Alepisaurus ferox Lowe, 1833: OShr; [COS]
Paralepididae
Lestolepis luetkeni (Ege, 1933): OShr; [IP]
Maculisudis longipinnis Kotthaus, 1967: OShr
Paralepis philippinus Fowler, 1934: OShr
Order Myctophiformes
Neoscopelidae
Neoscopelus macrolepidotus Jonhson, 1863: OShr; [COS]
Neoscopelus microchir Matsubara, 1943: OShr; [CIR];
PMBC
Myctophidae
Benthosema pterotum (Alcock, 1890): OShr; [IWP]
Diaphus coeruleus (Kluzinger, 1871): OShr; [IWP]
Diaphus splendidus (Brauer, 1904): OShr; [CIR]
Diaphus thiollierei Fowler, 1934: OShr; [IWP]
Order Lampriformes
Lophotidae
Lophotus lecepede Giorna, 1809: OShr; [CIR]; PMBC
Trachipteridae
Trachipterus jacksonensis (Ramsay, 1881): OShr; [COS]
Zu cristatus (Bonelli, 1820): OShr; [CIR]; PMBC
Order Polymixiiformes
Polymixiidae
Polymixia berndti Gilbert, 1905: OShr; [IP]
Order Gadiformes
Bregmacerotidae
Bregmaceros mcclellandi Thompson, 1840: OShr; [EIO];
PMBC
Macrouridae
Coelorinchus argentatus Smith & Radcliffe, 1912: OShr;
[IWP]
Coelorinchus macrorhynchus Smith & Radcliffe, 1912:
OShr; [WP]
Coelorinchus radcliffei Gilbert & Hubbs, 1920: OShr; [WP]
Coryphaenoides asprellus (Smith & Radcliff, 1912): OShr;
[IO]; PMBC
Malacocephalus laevis (Lowe, 1843): OShr; [COS]
Moridae
Physiculus roseus Alcock, 1891: OShr; [EIO-WP]
Carapidae
Encheliophis homei (Richardson, 1846): Sg; [IWP]; PMBC
Onuxodon fowleri (Smith, 1955): Cr; [IP]
Onuxodon parvibranchium (Fowler, 1927): Cr; [IP];
PMBC
Pyramodon ventralis Smith & Radcliffe, 1913: OShr; [IP]
Order Ophidiiformes
Ophidiidae
Glyptophidium argenteum Alcock, 1889: OShr; [IM];
PMBC
Glyptophidium macropus Alcock, 1894: OShr; [NIO]
Hypopleuron caninum Smith & Radcliffe, 1913: OShr;
[IWP]; PMBC
Neobythites fasciatus Smith & Radcliffe, 1913: OShr; [WP]
Bythitidae
Alionematichthys phuketensis Møller & Schwarzhans,
2008: Cr; [AS]
Alionematichthys piger (Alcock, 1890): Cr; [WCP]
Alionematichthys riukiuensis (Aoyaki, 1954): Cr; [WP]
Brosmophyciops pautzkei Schultz, 1960: Cr; [IWP]; PMBC
Dinematichthys iluocoeteoides Bleeker, 1855: Cr; [IWP];
PMBC
Order Batrachoidiformes
Batrachoididae
Allenbatrachus grunniens (Linnaeus, 1758): Mg, OShr;
[NIO-IM]; PMBC
Order Lophiiformes
Lophiidae
Lophiodes lugubris (Alcock, 1894): OShr; [EIO]
Lophiodes mutilus (Alcock, 1893): OShr; [IWP]; PMBC
Antennariidae
Antennarius coccineus (Cuvier, 1831): Cr; [IP]
Antennarius commerson (Latreille, 1804): Cr, OShr; [IP];
PMBC
Antennarius hispidus (Bloch & Schneider, 1801): Cr, OShr;
[IWP]
Antennarius maculatus (Desjardins, 1840): Cr; [IP]
Antennarius nummifer (Cuvier, 1817): Cr; [IP]; PMBC
Antennarius pictus (Shaw & Nodder, 1794): Cr; [IP]; PMBC
Chaunacidae
Chaunax pictus Lowe, 1846: OShr; [COS]
Ogcocephalidae
Halieutaea coccinea Alcock, 1889: OShr; [IO]
Halieutaea indica Annandale & Jenkins, 1910: OShr; [IWP]
Halieutaea stellata (Vahl, 1797): OShr; [WP]; PMBC
Halieutopsis micropa (Alcock, 1891): OShr; [IWP]
Order Mugiliformes
Mugilidae
Chelon macrolepis (Smith, 1846): Mg; [IP]; PMBC
Chelon parsia (Hamilton, 1822): Mg; [EIO]
Chelon subviridis (Valenciennes, 1836): Mg, Sg; [IWP];
PMBC
Crenimugil crenilabris (Forsskål, 1775): Cr; [IP]
Ellochelon vaigiensis (Quoy & Gaimard, 1825): Cr, Mg,
Sg; [IP]; PMBC
Moolgarda cunnesius (Valenciennes, 1836): Mg, Sg; [IWP];
PMBC
Moolgarda engeli (Bleeker, 1859): Mg; [IP]
52
Phuket mar. biol. Cent. Res. Bull.
Moolgarda pedaraki (Valenciennes, 1836): Mg, Sg; [IWP];
PMBC
Moolgarda perusii (Valenciennes, 1836): Mg, Sg; [IWP];
PMBC
Moolgarda seheli (Forsskål, 1775): Cr, Mg; [IP]; PMBC
Oedalechilus labiosus (Valenciennes, 1836): Cr; [IP];
PMBC
Paramugil parmatus (Cantor, 1849): Mg; [IM]; PMBC
Order Atheriniformes
Phallostethidae
Neostethus bicornis Regan, 1916: Mg; [IM]
Neostethus lankesteri Regan, 1916: Mg; [IM]; PMBC
Phenacostethus posthon Roberts, 1971: Mg; [AS]
Atherinidae
Atherinomorus duodecimalis (Valenciennes, 1835): Cr, Mg,
Sg, OShr; [IP]; PMBC
Atherinomorus lacunosus (Forster & Schneider, 1801):
OShr; [EIO-WP]; PMBC
Atherinomorus pinguis (Lacepède, 1803): Sg, OShr; [IWP]
Atherion elymus Jordan & Starks, 1901: Cr, OShr; [WP];
PMBC
Hypoatherina temminckii (Bleeker, 1853): Cr, OShr [IP];
PMBC
Hypoatherina valenciennei (Bleeker, 1853): Mg, Sg, OShr;
[WP]; PMBC
Order Beloniformes
Andrianichthyidae
Oryzias javanicus (Bleeker, 1854): Mg; [IM]; PMBC
Exocoetidae
Cheilopogon arcticeps (Günther, 1866): OShr; [WP]
Cheilopogon spilopterus (Valenciennes, 1846): OShr; [WP];
PMBC
Exocoetus volitans (Linnaeus, 1758): OShr; [COS]; PMBC
Hirundichthys speculiger (Valenciennes, 1847): OShr; [CIR]
Parexocoetus brachypterus (Richardson, 1846): OShr; [IP];
PMBC
Hemiramphidae
Dermogenys siamensis Fowler, 1934: Mg; [IM]
Hemiramphus far (Forsskål, 1775): Cr, Sg, OShr; [IWP];
PMBC
Hyporhamphus balinensis (Bleeker, 1858): OShr; [IWP]
Hyporhamphus dussumieri (Valenciennes, 1846): Sg, OShr;
[IP]; PMBC
Hyporhamphus limbatus (Valenciennes, 1846): Mg, Sg,
OShr; [NIO-WP]
Hyporhamphus melanopterus Collette & Parin, 1978: Mg,
OShr; [IM]; PMBC
Hyporhamphus quoyi Valenciennes, 1847: Mg, Sg, OShr;
[IWP]; PMBC
Hyporhamphus unicuspis Collette & Parin, 1978: OShr;
[NIO]
Zenarchopterus buffornis (Valenciennes, 1847): Mg, Sg,
OShr; [EIO-IA]; PMBC
Zenarchopterus dispar (Valenciennes, 1847): Mg, Sg, OShr;
[IWP]; PMBC
Zenarchopterus dunckeri Mohr, 1926: Mg, Sg, OShr; [WP];
PMBC
Belonidae
Ablennes hians (Valenciennes, 1846): Cr, OShr; [COS];
PMBC
Strongylura incisa (Valenciennes, 1846): OShr; [EIO-WP];
PMBC
Strongylura leiura (Bleeker, 1850): OShr; [IWP]; PMBC
Strongylura strongylura (van Hasselt, 1823): Mg, Sg, OShr;
[NIO-IA]; PMBC
Tylosurus acus melanotus (Bleeker, 1850): Cr, OShr; [IP]
Tylosurus crocodilus crocodilus (Peron & Lesueur, 1821):
Cr, Mg, Sg, OShr; [CIR]
Order Cyprinodontiformes
Aplocheilidae
Aplocheilus panchax Day, 1875: Mg; [EIO-IM]; PMBC
Order Beryciformes
Diretmidae
Diretmoides veriginae Kotlyar, 1987: OShr; [EIO-WP];
PMBC
Monocentridae
Monocentris japonica (Houttuyn, 1782): OShr; [IWP];
PMBC
Trachichthyidae
Gephyroberyx darwinii (Johnson, 1866): OShr; [COS];
PMBC
Berycidae
Beryx splendens Lowe, 1834: OShr; [COS]
Centroberyx druzhinini (Busakhin, 1981): OShr; [IWP];
PMBC
Holocentridae
Myripristis adustus (Bleeker, 1853): Cr, OShr; [IP]
Myripristis berndti (Jordan & Evermann, 1903): Cr, OShr;
[IP]; PMBC
Myripristis botche Cuvier, 1829: Cr, OShr; [IWP]; PMBC
Myripristis chryseres Jordan & Evermann, 1903: Cr, OShr;
[IP]
Myripristis hexagona (Lacepède, 1802): Cr, OShr; [IWP];
PMBC
Myripristis kuntee Cuvier, 1831: Cr, [IP]
Myripristis murdjan (Forsskål, 1775): Cr; [IP]
Myripristis violacea Bleeker, 1851: Cr; [IP]
Myripristis vittatus Valenciennes, 1831: Cr; [IP]
Neoniphon sammara (Forsskål, 1775): Cr, OShr; [IP];
PMBC
Ostichthys japonicus (Cuvier, 1829): OShr; [WP]; PMBC
Sargocentron caudimaculatum (Rüppell, 1835): Cr; [IP];
PMBC
Sargocentron cornutum (Bleeker, 1853): Cr; [WP]; PMBC
Sargocentron diadema (Lacepède, 1802): Cr; [IP]
Sargocentron melanospilos (Bleeker, 1858): Cr; [IWP]
Sargocentron rubrum (Forsskål, 1775): Cr, OShr; [IWP];
PMBC
Sargocentron spiniferum (Forsskål, 1775): Cr; [IP]
Order Zeiformes
Grammicolepididae
Xenolepidichthys dalgleishi Gilchrist, 1922: OShr; [CIR]
Zeidae
Cyttopsis rosea (Lowe, 1843): OShr; [COS]; PMBC
53
The fishes of southwestern Thailand
Zenopsis nebulosa (Temminck & Schlegel, 1845): OShr;
[IP]
Order Gasterosteiformes
Pegasidae
Eurypegasus draconis (Linnaeus, 1766): Cr, OShr; [IP];
PMBC
Pegasus volitans Linnaeus, 1758: Cr, Sg, OShr; [IWP];
PMBC
Solenostomidae
Solenostomus cyanopterus Bleeker, 1852: Cr; [IP]
Solenostomus paradoxus (Pallas, 1770): Cr, Sg; [IWP];
PMBC
Syngnathidae
Bhanotia fasciolata (Dumeril, 1870): Cr, OShr; [IA];
PMBC
Choeroichthys brachysoma (Bleeker, 1855): Cr; [IP];
PMBC
Choeroichthys sculptus (Günther, 1870): Cr, OShr; [IP]
Corythoichthys amplexus Dowson & Randall, 1906: Cr,
OShr; [IWP]
Corythoichthys flavofasciatus (Rüppell, 1838): Cr; [IP]
Corythoichthys haematopterus (Bleeker, 1851): Cr, OShr;
[IWP]; PMBC
Corythoichthys schultzi Herald, 1953: Cr; [IP]; PMBC
Doryrhamphus janssi (Herald & Randall, 1972): Cr, OShr;
[WP]; PMBC
Dunckerocampus multiannulatus (Regan, 1903): Cr; [IO]
Halicampus grayi Kaup, 1856: OShr; [NIO-WP]; PMBC
Halicampus mataafae (Jordan & Seale, 1906): Cr, OShr;
[IP]
Halicampus spinirostris (Dawson & Allen, 1981): Cr;
[EIO-WP]
Hippichthys cyanospilus (Bleeker, 1854): Sg, OShr; [IWP];
PMBC
Hippichthys heptagonus (Bleeker, 1853): Sg, OShr; [IWP];
PMBC
Hippichthys penicillus (Cantor, 1849): Mg, Sg, OShr; [NIOWP]; PMBC
Hippichthys spicifer (Rüppell, 1840): Mg, OShr; [IP];
PMBC
Hippocampus comes Cantor, 1850: Cr, Sg; [IM]; PMBC
Hippocampus kelloggi Jordan & Snyder, 1901: OShr;
[IWP]; PMBC
Hippocampus kuda Bleeker, 1852: Mg, Sg, OShr;[NIOWCP]; PMBC
Hippocampus spinosissimus Weber, 1913: OShr; [EIO-IA];
PMBC
Hippocampus trimaculatus Leach, 1814: OShr; [EIO-WP];
PMBC
Phoxocampus belcheri (Kaup, 1856): Cr, OShr; [IWP]
Syngnathoides biaculeatus (Bloch, 1785): Sg, OShr; [IWP];
PMBC
Trachyrhamphus bicoarctatus (Bleeker, 1857): Cr, Sg,
OShr; [IWP]; PMBC
Trachyrhamphus longirostris Kaup, 1856: OShr; [IWP];
PMBC
Trachyrhamphus serratus (Temminck & Schlegel, 1850):
OShr; [EIO-WP]; PMBC
Aulostomidae
Aulostomus chinensis (Linnaeus, 1766): Cr; [IP]; PMBC
Fistulariidae
Fistularia commersonii Rüppell, 1835: Cr, Sg; [IP]; PMBC
Fistularia petimba Lacepède, 1803: OShr; [CIR]; PMBC
Centriscidae
Aeoliscus strigatus (Günther, 1860): Cr, Sg, OShr; [IWP]
Centriscus cristatus De Vis, 1885: OShr; [WP]
Centriscus scutatus Linnaeus, 1758: Cr, Sg, OShr; [IWP];
PMBC
Order Sybranchiformes
Sybranchidae
Macrotrema caligans (Cantor, 1849): Mg; [IM]
Order Scorpaeniformes
Dactylopteridae
Dactyloptena macracantha (Bleeker, 1854): OShr; [EIOWP]; PMBC
Dactyloptena orientalis (Cuvier, 1829): Cr, OShr; [IP];
PMBC
Dactyloptena perterseni (Nystrom, 1887): OShr; [IWP];
PMBC
Scorpaenidae
Ablabys taenianotus (Cuvier, 1829): Cr; [EIO-WP]
Apistus carinatus (Bloch & Schneider, 1801): OShr; [IWP]
Choridactylus multibarbus Richardson, 1848: OShr; [IWP]
Dendrochirus biocellatus (Fowler, 1938): Cr; [IP]
Dendrochirus brachypterus (Cuvier, 1829): Cr, OShr; [IP];
PMBC
Dendrochirus zebra (Cuvier, 1829): Cr, Sg, OShr; [IWP];
PMBC
Inimicus cuvieri (Gray, 1832): Cr, OShr; [IM]
Lioscorpius longiceps Günther, 1880: OShr; [WP]
Minous coccineus Alcock, 1890: OShr; [IWP]
Minous inermis Alcock, 1889: OShr; [IO]
Minous monodactylus (Bloch & Schneider, 1801): OShr;
[IWP]; PMBC
Minous trachycephalus (Bleeker, 1854): OShr; [EIO-WP]
Neomerinthe amplisquamiceps (Fowler, 1938): OShr; [WP]
Paracentropogon longispinis (Bleeker, 1829): Cr, OShr;
[EIO-WP]
Parapterois heterurus (Bleeker, 1856): OShr; [IWP]
Parascorpaena aurita (Rüppell, 1838): Cr, OShr; [IWP];
PMBC
Parascorpaena mossambica (Peters, 1855): Cr; [IP]
Pontinus macrocephalus (Sauvage, 1882): OShr; [WP];
PMBC
Pterois antennata (Bloch, 1787): Cr, OShr; [IP]; PMBC
Pterois miles (Bennett, 1828): Cr, Sg, OShr; [IO]; PMBC
Pterois mombasae (Smith, 1957): Cr, OShr; [IO]
Pterois radiata Cuvier, 1829: Cr, OShr; [IP]; PMBC
Pterois russellii Bennett, 1831: Sg, OShr; [IWP]; PMBC
Rhinopias eschmeyeri Conde, 1977: Cr; [IWP]
Rhinopias frondosa (Günther, 1891): Cr, OShr; [IWP]
Richardsonichthys leucogaster (Richardson, 1848): OShr;
[IWP]
54
Phuket mar. biol. Cent. Res. Bull.
Scorpaenodes albaiensis (Evermann & Seale, 1907): Cr;
[IWP]; PMBC
Scorpaenodes guamensis (Quoy & Gaimard, 1824): Cr,
OShr; [IP]; PMBC
Scorpaenodes hirsutus (Smith, 1957): Cr; [IP]
Scorpaenodes minor (Smith, 1958): Cr; [IP]
Scorpaenodes parvipinnis (Garrett, 1863): Cr, OShr; [IP];
PMBC
Scorpaenodes scaber (Ramsay & Ogilby, 1886): Cr; [EIOWCP]
Scorpaenodes varipinnis Smith, 1957: Cr; [IWP]
Scorpaenopsis diabolus Cuvier, 1829: Cr, OShr; [IP];
PMBC
Scorpaenopsis longispina Randall & Eschmeyer, 2001: Cr;
[IO]
Scorpaenopsis neglecta Heckel, 1837: Sg, OShr; [EIO-WP];
PMBC
Scorpaenopsis oxycephala (Bleeker, 1849): Cr, OShr;
[IWP]; PMBC
Scorpaenopsis possi Randall & Eschmeyer, 2001: Cr; [IP]
Scorpaenopsis ramaraoi Randall & Eschmeyer, 2001: Cr,
OShr; [NIO-WP]
Scorpaenopsis vesosa (Cuvier, 1829): OShr; [IWP]
Sebastapistes fowleri (Pietschmann, 1934): Cr; [IP]
Sebastapistes mauritiana (Cuvier, 1829): Cr, OShr; [IP]
Sebastapistes strongia (Cuvier, 1829): Cr; [IP]
Setarches guentheri Johnson, 1862: OShr; [CIR]
Synanceia horrida (Linnaeus, 1766): Cr, OShr; [EIO-WP];
PMBC
Synanceia verrucosa (Smith, 1956): Cr, OShr; [IP]
Taenianotus triacanthus Lacepède, 1802: Cr; [IP]
Trachicephalus uranoscopus (Bloch & Schneider, 1801):
Mg, Sg, OShr; [EIO-WP], PMBC
Vespicula trachinoides (Cuvier & Valenciennes, 1829): Mg,
Sg, OShr; [WP]; PMBC
Caracanthidae
Caracanthus unipinna (Gray, 1831): Cr; [IWP]; PMBC
Aploactinidae
Acanthosphex leurynnis (Jordan & Seale, 1905): Mg, Sg,
OShr; [EIO-IM]; PMBC
Cocotropus echinatus (Cantor, 1850): Mg, Sg; [AS]
Cocotropus larvatus Poss & Allen, 1987: OShr; [WP]
Paraploactis obbesi (Weber, 1913): Cr, OShr; [WP]
Sthenopus mollis Richardson, 1848: OShr; [WP]
Xenaploactis cautes Poss & Eschmeyer, 1980: OShr; [IM
(presently known from AS and Gulf of Thailand)]
Triglidae
Lepidotrigla punctipectoralis Fowler, 1938: OShr; [IWP]
Lepidotrigla spiloptera Günther, 1880: OShr; [IWP];
PMBC
Peristediidae
Peristedion liorhynchus Günther, 1871: OShr; [IWP];
PMBC
Peristedion moluccense Bleeker, 1850: OShr; [WP]; PMBC
Satyrichthys adeni (Lloyd, 1907): OShr; [IWP]; PMBC
Satyrichthys sp.: OShr; PMBC
Bembridae
Brachybembras sp.: OShr
Plathycephalidae
Cociella punctata (Cuvier, 1829): Cr, Mg, Sg, OShr; [IWP];
PMBC
Elates ransonneti (Steindachner, 1877): OShr; [IA]; PMBC
Grammoplites scaber (Linnaeus, 1758): Mg, Sg, OShr;
[NIO-IM]; PMBC
Inegocia japonica (Tilesius, 1812): Mg, Sg, OShr; [EIOWP]; PMBC
Kumococius rodericensis Cuvier in Cuvier & Valenciennes,
1829: OShr; [NIO-IA]; PMBC
Platycephalus indicus (Linnaeus, 1758): Mg, Sg, OShr;
[IWP]; PMBC]
Rogadias pristiger (Cuvier, 1829): OShr; [IWP]; PMBC
Rogadias serratus (Cuvier, 1829): OShr; [IWP]; PMBC
Sorsogona melanoptera (Knapp & Wongratana, 1987);
OShr; [NIO]
Sorsogona tuberculata (Cuvier, 1829): OShr; [NIO-IA];
PMBC
Suggrundus macracanthus (Bleeker, 1869): OShr; [EIOIA]; PMBC
Sunagocia carbunculus (Valenciennes, 1833): Mg, Sg, OShr;
[EIO-IM]; PMBC
Sunagocia otaitensis (Cuvier, 1829): Cr, OShr; [IP]; PMBC
Thrysanophrys celebicus (Bleeker, 1854): Cr; [IWP]
Thrysanophrys chitonae Schultz, 1966: Cr, OShr; [IP]
Hoplichthyidae
Hoplichthys citrinus Gilbert, 1905: OShr; [WP]
Order Perciformes
Ambassidae
Ambassis buruensis Bleeker, 1857: Mg; [WP]; PMBC
Ambassis dussumieri Cuveir, 1828: Sg; [IWP]
Ambassis gymnocephalus (Lacepède, 1802): Mg; [IWP];
PMBC
Ambassis interruptus Bleeker, 1852: Mg, Sg; [IA]; PMBC
Ambassis macracanthus Bleeker, 1849: Mg, Sg; [EIO-IM];
PMBC
Ambassis nalua (Hamilton, 1822): Mg, Sg; [EIO-IA];
PMBC
Ambassis urotaenia Bleeker, 1852: Mg, Sg; [IWP]; PMBC
Ambassis vachellii Richardson, 1846: Mg, Sg; [IWP];
PMBC
Latidae
Lates calcalifer (Bloch, 1790): Mg; [NIO-WP]; PMBC
Acropomatidae
Acropoma argentistigma Okamoto & Ida, 2002: OShr; [AS]
Synagrops japonicus (Döderlein, 1883): OShr; [IP]; PMBC
Synagrops malayanus Weber, 1913: OShr; [WCP]
Synagrops philippinensis (Günther, 1880): OShr; [EIOWP]; PMBC
Serranidae
Aethaloperca rogaa (Forsskål, 1775): Cr, OShr; [IP];
PMBC
Anyperodon leucogrammicus (Valnciennes, 1828): Cr, OShr;
[IP]; PMBC
55
The fishes of southwestern Thailand
Cephalopholis argus Bloch & Schneider, 1801: Cr, OShr;
[IP]; PMBC
Cephalopholis aurantia (Valenciennes, 1828): OShr; [IWP]
Cephalopholis boenak (Bloch, 1790): Cr, Sg, OShr; [IWP];
PMBC
Cephalopholis formosa (Shaw & Nodder, 1812): Cr, OShr;
[EIO-WP]; PMBC
Cephalopholis leopardus (Lacepède, 1801): Cr; [IP]
Cephalopholis miniata (Forsskål, 1775): Cr, OShr; [IP];
PMBC
Cephalopholis polyspila Randall & Satapoomin, 2000: Cr,
OShr; [AS]; PMBC
Cephalopholis sexmaculata (Rüppell, 1830): Cr; [IP]
Cephalopholis sonnerati (Valenciennes, 1828): Cr, OShr;
[IP]; PMBC
Cephalopholis urodeta (Bloch & Schneider, 1801): Cr,
OShr; [IP]; PMBC
Chelidoperca margaritifera Weber, 1913: OShr; [WP];
PMBC
Cromileptes altivelis (Valenciennes, 1828): Cr, OShr; [WP];
PMBC
Diploprion bifasciatum Cuvier, 1828: Cr, OShr; [EIO-WP];
PMBC
Epinephelus amblycephalus (Bleeker, 1857): OShr; [WP];
PMBC
Epinephelus areolatus (Forsskål, 1775): Cr, OShr; [IWP];
PMBC
Epinephelus bleekeri (Vaillant, 1877): Cr, Mg, Sg, OShr;
[NIO-IA]; PMBC
Epinephelus caeruleopunctatus (Bloch, 1790): Cr, OShr;
[IWP]; PMBC
Epinephelus coioides (Hamilton, 1822): Cr, Mg, Sg, OShr:
[IWP]; PMBC
Epinephelus epistictus (Temminck & Schlegel, 1842): OShr;
[IWP]; PMBC
Epinephelus erythrurus (Valenciennes, 1828): Cr, Sg, OShr;
[NIO-IM]; PMBC
Epinephelus fasciatus (Forsskål, 1775): Cr, OShr; [IP];
PMBC
Epinephelus flavocaeruleus (Lacepède, 1801): Cr, OShr;
[IO]
Epinephelus fuscoguttatus (Forsskål, 1775): Cr, OShr; [IP];
PMBC
Epinephelus heniochus Fowler, 1904: OShr; [WP]
Epinephelus hexagonatus (Bloch & Schneider, 1801): Cr;
[IP]
Epinephelus lanceolatus (Bloch, 1790): Cr, OShr; [IP];
PMBC
Epinephelus latifasciatus (Temminck & Schlegel, 1842):
OShr; [IWP]
Epinephelus longispinis (Kner, 1864): Cr, OShr; [IO]
Epinephelus macrospilos (Bleeker, 1855): Cr; [IP]; PMBC
Epinephelus malabaricus (Bloch & Schneider, 1801): Cr,
Mg, OShr; [IWP]; PMBC
Epinephelus merra Bloch, 1793: Cr, OShr; [IP]; PMBC
Epinephelus morrhua (Valenciennes, 1833): OShr; [IP];
PMBC
Epinephelus octofasciatus Griffin, 1926: OShr; [IP]; PMBC
Epinephelus ongus (Bloch, 1790): Cr, OShr; [IWP]; PMBC
Epinephelus polyphekadion (Bleeker, 1849): Cr, OShr; [IP];
PMBC
Epinephelus quoyanus (Valenciennes, 1830): Cr, Sg, OShr;
[WP]; PMBC
Epinephelus radiatus (Day, 1867): OShr; [IWP]; PMBC
Epinephelus sexfasciatus (Valenciennes, 1828): Mg, Sg,
OShr; [IA]; PMBC
Epinephelus spilotoceps Schultz, 1953: Cr; [IP]
Epinephelus tauvina (Forsskål, 1775): Cr, OShr; [IP];
PMBC
Epinephelus tukula Morgan, 1959: Cr, OShr; [IWP]
Epinephelus undulosus (Quoy & Gaimard, 1824): Cr, OShr;
[IWP]; PMBC
Grammistes sexlineatus (Thunberg, 1792): Cr; [IP]
Grammistops ocellatus Schultz, 1953: Cr; [IP]
Liopropoma africanum (Smith, 1954): Cr; [IO]; PMBC
Liopropoma susumi (Jordan & Seale, 1906): Cr; [IP];
PMBC
Meganthias filiferus Randall & Heemstra, 2007: OShr; [AS];
PMBC
Plectropomus areolatus (Rüppell, 1830): Cr, OShr; [IP]
Plectropomus laevis (Lacepède, 1801): Cr, OShr; [IP]
Plectropomus pessuliferus Fowler, 1901: Cr, OShr; [IWP]
Pogonoperca ocellata Günther, 1859: Cr, OShr; [IO];
PMBC
Pseudanthias evansi (Smith, 1954): Cr; [IO]; PMBC
Pseudanthias ignitus (Randall & Lubbock, 1981): Cr; [EIO]
Pseudanthias rubrizonatus (Randall, 1983): Cr; [WP]
Pseudanthias squamipinnis (Peters, 1855): Cr; [IWP];
PMBC
Pseudogramma polyacanthum (Bleeker, 1856): Cr; [IP];
PMBC
Rabaulichthys stigmaticus Randall & Pyle, 1989: Cr, OShr;
[IO]
Variola albimarginata Baissac, 1953: Cr, OShr; [IWP]
Variola louti (Forsskål, 1775): Cr, OShr; [IP]; PMBC
Ostracoberycidae
Ostracoberyx dorygenys Fowler, 1934: OShr; [IWP]
Pseudochromidae
Amsichthys knighti (Allen, 1987): Cr; [WP]
Blennodesmus scapularis Günther, 1871: Cr; [IA]; PMBC
Pseudochromis andamanensis Lubbock, 1980: Cr; [EEIO];
PMBC
Pseudochromis caudalis Boulenger, 1898: Cr; [IO]
Pseudochromis coccinicauda Tickell in Day, 1888: Cr;
[EIO]
Pseudochromis fuscus Müller & Troschel, 1849: Cr; [EIOIA]; PMBC
Pseudochromis tapeinosoma Bleeker, 1853: Cr; [IA];
PMBC
Pseudoplesiops rosae Schultz, 1953: Cr; [EIO-WCP];
PMBC
Plesiopidae
Calloplesiops altivelis (Steindachner, 1903); Cr; [IP]
Plesiops auritus Mooi, 1995: Cr; [EIO]
56
Phuket mar. biol. Cent. Res. Bull.
Plesiops caeruleolineatus Rüppell, 1835: Cr; [IP]; PMBC
Plesiops corallicola Bleeker, 1853: Cr; [WCP]; PMBC
Opistognathidae
Opistognathus cyanospilotus Smith-Vaniz, 2009: Cr;
[EEIO]
Opistognathus nigromarginatus Rüppell, 1830: Cr, OShr;
[IWP]; PMBC
Opistognathus rosenbergii (Bleeker, 1856): Cr, OShr;
[EIO-WP]; PMBC
Opistognathus variabilis Smith-Vaniz, 2009: Cr; [EIO-WP];
PMBC
Priacanthidae
Heteropriacanthus cruentatus (Lacepède, 1801): Cr, OShr;
[COS]; PMBC
Priacanthus blochii Bleeker, 1853: OShr; [IWP]; PMBC
Priacanthus hamrur (Forsskål, 1775): Cr, OShr; [IP];
PMBC
Priacanthus macracanthus Cuvier, 1829: Cr, OShr; [WP];
PMBC
Priacanthus sagittarius Starnes, 1988: OShr; [IWP]
Priacanthus tayenus Richardson, 1846: OShr; [NIO-WP];
PMBC
Pristigenys niphonia (Cuvier, 1829): OShr; [IWP]
Apogonidae
Apogon abrogramma Fraser & Lachner, 1985: Cr; [IWP]
Apogon angustatus (Smith & Radcliffe, 1911): Cr; [IP];
PMBC
Apogon apogonides (Bleeker, 1856): Cr; [IWP]
Apogon aureus (Lacepède, 1802): Cr, OShr; [IWP]
Apogon bryx Fraser, 1998: OShr; [IWP]
Apogon campbelli Smith, 1949: Cr; [IO]
Apogon compressus (Smith & Radcliffe, 1911): Cr; [WP];
PMBC
Apogon cookii Macleay, 1881: Cr; [IWP]; PMBC
Apogon cyanosoma Bleeker, 1853: Cr; [IWP]; PMBC
Apogon endekataenia Bleeker, 1852: Cr; [IA]; PMBC
Apogon fasciatus (White, 1790): OShr; [IWP]; PMBC
Apogon fleurieu (Lacepède, 1802): Cr, OShr; [IWP]
Apogon frenatus Valenciennes, 1832: Cr; [IP]; PMBC
Apogon hyalosoma Bleeker, 1852: Mg, Sg; [WP]; PMBC
Apogon kallopterus Bleeker, 1856: Cr; [IP]; PMBC
Apogon kalosoma (Bleeker, 1852): Cr; [EIO-WP]
Apogon lateralis Valenciennes, 1832: Cr, Mg, Sg; [IWP];
PMBC
Apogon moluccensis Valenciennes, 1832: Cr, Sg; [IA];
PMBC
Apogon nigrofasciatus Lachner, 1953: Cr; [IP]; PMBC
Apogon novaeguinae Valenciennes, 1832: Mg, Sg; [IWP];
PMBC
Apogon ocellicaudus Allen, Kuiter & Randall, 1994: Cr;
[IA]
Apogon pleuron Fraser, 2005: Sg, OShr; [EIO-WP]; PMBC
Apogon poecilopterus Cuvier, 1828: OShr; [EIO-WP]
Apogon semiornatus Peters, 1876: Cr; [IWP]
Apogon smithi (Kotthaus, 1970): OShr; [IWP]; PMBC
Apogon sp.1: Cr; [AS]
Apogon striatodes Gon, 1996: OShr; [IM]; PMBC
Apogon striatus (Smith & Radcliffe, 1912): OShr; [NIOWP]; PMBC
Apogon taeniophorus Regan, 1908: Cr; [IP]; PMBC
Apogon talboti Smith, 1961: Cr; [IWP]
Apogon thermalis Cuvier, 1829: Cr, Sg; [IWP]; PMBC
Apogon trimaculatus Cuvier, 1828: Cr, OShr; [WP]; PMBC
Apogon truncatus Bleeker, 1854: OShr; [IWP]; PMBC
Apogonichthyoides nigripinnis (Cuvier, 1828): OShr; [EIOIA]; PMBC
Apogonichthyoides sialis (Jordan & Thompson, 1914): Cr;
[EIO-WP]
Apogonichthys sp.: OShr
Archamia ataenia Randall & Satapoomin, 1999: Cr; [AS];
PMBC
Archamia bleekeri Günther, 1859: Cr; [IWP]
Archamia fucata (Cantor, 1850): Cr, Sg, OShr; [IP]; PMBC
Archamia macroptera (Cuvier, 1828): Cr; [EIO-WP]
Cheilodipterus artus Smith, 1961: Cr; [IP]; PMBC
Cheilodipterus macrodon (Lacepède, 1802): Cr; [IP];
PMBC
Cheilodipterus quinquelineatus Cuvier, 1828: Cr; [IP];
PMBC
Foa fo (Jordan & Seale, 1906): Sg, OShr; [IWP]; PMBC
Fowleria aurita (Valenciennes, 1831): Cr; [IWP]; PMBC
Fowleria isostigma (Jordan & Seale, 1960): Cr; [IWP];
PMBC
Fowleria vaiulae (Jordan & Seale, 1906): Cr; [IWP];
PMBC
Fowleria variegata (Valenciennes, 1832): Cr, Mg, Sg;
[IWP]; PMBC
Gymnapogon urospilotus Lachner, 1953: Cr; [WCP]
Neamia octospina Smith & Radcliffe, 1912: Cr; [IWP]
Nectamia fusca (Quoy & Gaimard, 1824): Cr; [IWP];
PMBC
Nectamia luxuria Fraser, 2008: Cr; [EIO-WP]; PMBC
Nectamia savayensis (Günther, 1872): Cr; [IP]
Pseudamia hayashii Randall, Lachner, & Fraser, 1985: Cr;
[IP]
Pseudamia sp.1: Cr
Rhabdamia cypselurus Weber, 1909: Cr; [IWP]; PMBC
Rhabdamia gracilis (Bleeker, 1856): Cr; [IWP]; PMBC
Siphamia tubifer Weber, 1909: Cr; [WP]
Zoramia leptacantha (Bleeker, 1856): Cr; [IWP]; PMBC
Zoramia perlita (Fraser & Lachner, 1985): Cr; [IM]
Sillaginidae
Sillaginopsis panijus (Hamilton, 1822): OShr; [EIO];
PMBC
Sillago aeolus Jordan & Evermann, 1902: Cr, Mg, Sg, OShr;
[IWP]; PMBC
Sillago chondropus Bleeker, 1849: OShr; [IWP]; PMBC
Sillago intermedius Wongratana, 1977: OShr; [EIO-IM]
Sillago lutea McKay, 1985: OShr; [EIO-IA]
Sillago sihama (Forsskal, 1775): Mg, Sg, OShr; [IWP];
PMBC
Malacanthidae
57
The fishes of southwestern Thailand
Branchiostegus sawakinensis Amirthalingam, 1969: OShr;
[IWP]; PMBC
Hoplolatilus cuniculus Randall & Dooley, 1974: Cr; [IP]
Hoplolatilus luteus Allen & Kuiter, 1989: Cr, OShr; [EEIO];
PMBC
Malacanthus brevirostris Guichenot, 1848: Cr, OShr; [IP];
PMBC
Malacanthus latovittatus (Lacepède, 1801): Cr; [IP]
Lactariidae
Lactarius lactarius (Bloch & Schneider, 1801): OShr;
[NIO-WP]; PMBC
Coryphaenidae
Coryphaena equiselis Linnaeus, 1758: OShr; [COS]; PMBC
Coryphaena hippurus Linnaeus, 1758: OShr; [COS];
PMBC
Rachycentridae
Rachycentron canadum (Linnaeus, 1766): Cr, Sg, OShr;
[CIR]; PMBC
Echeneidae
Echeneis naucrates Linnaeus, 1758: Cr, OShr; [COS];
PMBC
Phtheirichthys lineatus (Menzies, 1791): OShr; [COS]
Remora remora (Linnaeus, 1758): OShr; [COS]
Carangidae
Alectis ciliaris (Bloch, 1788): Cr, OShr; [CIR]; PMBC
Alectis indica (Rüppell, 1830): Cr, Sg, OShr; [IWP]; PMBC
Alepes djedaba (Forsskål, 1775): Mg, OShr; [IWP]; PMBC
Alepes kleinii (Bloch, 1793): Mg, Sg, OShr; [NIO-WP];
PMBC
Alepes melanoptera (Swainson, 1839): OShr; [NIO-IM];
PMBC
Alepes vari (Cuvier, 1833): Cr, Mg, OShr; [IWP]
Atropus atropus (Schneider, 1801): OShr; [NIO-WP]
Atule mate (Cuvier, 1833): Cr, Mg, Sg, OShr; [IP]; PMBC
Carangoides armatus (Rüppell, 1830): Mg, Sg, OShr;
[IWP]; PMBC
Carangoides caeruleopinnatus (Rüppell, 1830): Cr, Sg,
OShr; [IWP]; PMBC
Carangoides chrysophrys (Cuvier, 1833): OShr; [IWP]
Carangoides equula (Temminck & Schlegel, 1844): Cr,
OShr; [IP]
Carangoides ferdua (Forsskål, 1775): Cr, OShr; [IP];
PMBC
Carangoides fulvoguttatus (Forsskål, 1775): OShr; [IWP];
PMBC
Carangoides gymnostethus (Cuvier, 1833): OShr; [IWP];
PMBC
Carangoides hedlandensis (Whitley, 1933): Cr, OShr;
[IWP]; PMBC
Carangoides malabaricus (Bloch & Schneider, 1801):
OShr; [IWP]; PMBC
Carangoides oblongus (Cuvier, 1833): OShr; [IWP];
PMBC
Carangoides orthogrammus (Jordan & Gilbert, 1882): Cr,
OShr; [IP]
Carangoides plagiotaenia Bleeker, 1857: Cr, OShr; [IP];
PMBC
Carangoides praeustus (Bennett, 1830): Mg, Sg, OShr;
[NIO-IM]; PMBC
Carangoides talamparoides Bleeker, 1852: OShr; [NIOIA]
Caranx heberi (Bennett, 1830): Cr, OShr; [NIO-WP]
Caranx ignobilis (Forsskål, 1775): Cr, Mg, Sg, OShr; [IP];
PMBC
Caranx melampygus Cuvier, 1833: Cr, OShr; [IP]; PMBC
Caranx papuensis Alleyne & Macleay, 1877: Cr, OShr;
[IP]
Caranx sexfasciatus Quoy & Gaimard, 1824: Cr, Mg, Sg,
OShr; [IP]; PMBC
Decapterus kurroides Bleeker, 1855: OShr; [IWP]; PMBC
Decapterus macarellus (Cuvier, 1833): OShr; [CIR];
PMBC
Decapterus macrosoma Bleeker, 1851: Cr, OShr; [IP];
PMBC
Decapterus maruadsi (Temminck & Schlegel, 1843): OShr;
[WP]; PMBC
Decapterus russelli (Rüppell, 1830): Cr, OShr; [IWP];
PMBC
Elagatis bipinnulata (Quoy & Gaimard, 1825): Cr, OShr;
[CIR]; PMBC
Gnathanodon speciosus (Forsskål, 1775): Cr, OShr; [IP];
PMBC
Megalaspis cordyla (Linnaeus, 1758): Cr, OShr; [IP];
PMBC
Naucrates ductor (Linnaeus, 1758): OShr; [CIR]; PMBC
Parastromateus niger (Bloch, 1795): OShr; [IWP]; PMBC
Scomberoides commersonianus Lacepède, 1801: Cr, Mg,
Sg, OShr; [IWP]; PMBC
Scomberoides lysan (Forsskål, 1775): Cr, Mg, Sg, OShr;
[IP]; PMBC
Scomberoides tala (Cuvier, 1832): OShr; [EIO-WP];
PMBC
Scomberoides tol (Cuvier, 1832): Cr, Mg, Sg, OShr; [IWP];
PMBC
Selar boops (Cuvier, 1833): Cr, OShr; [IWP]; PMBC
Selar crumenophthalmus (Bloch, 1793): Cr, OShr; [CIR];
PMBC
Selaroides leptolepis (Cuvier, 1833): Cr, Sg, OShr; [NIOWP]; PMBC
Seriola rivoliana Valenciennes, 1833: OShr; [CIR]; PMBC
Seriolina nigrofasciata (Rüppell, 1829): Cr, OShr; [IWP];
PMBC
Trachinotus bailloni (Lacepède, 1801): Cr, OShr; [IP];
PMBC
Trachinotus blochii (Lacepède, 1801): Cr, OShr; [IP]
Trachinotus botla (Shaw, 1803): OShr; [IO]; PMBC
Ulua mentalis (Cuvier, 1833): Cr, OShr; [IWP]; PMBC
Uraspis helvola (Forster, 1801): OShr; [IP]; PMBC
Uraspis uraspis (Günther, 1860): OShr; [IWP]; PMBC
Menidae
Mene maculata (Bloch & Schneider, 1801): OShr; [IWP];
PMBC
Leiognathidae
Equulites berbis (Valenciennes, 1835): OShr; [IWP]; PMBC
58
Phuket mar. biol. Cent. Res. Bull.
Equulites elongatus (Günther, 1860): OShr; [IWP]; PMBC
Equulites leuciscus (Günther, 1860): Mg, Sg, OShr; [IWP];
PMBC
Equulites oblongus (Valenciennes, 1835): Mg, OShr; [IWP];
PMBC
Equulites stercorarius (Everman & Seale, 1907): Mg, Sg,
OShr; [WP]; PMBC
Eubleekeria jonesi (James, 1971): Mg, Sg, OShr; [EIOWP]; PMBC
Gazza dentex (Valenciennes, 1835): OShr; [IO]; PMBC
Gazza minuta (Bloch, 1797): Mg, Sg, OShr; [IWP]; PMBC
Gazza rhombea Kimura, Yamashita & Iwatsuki, 2000:
OShr; [EIO-WP]; PMBC
Karalla daura (Cuvier, 1829): Mg, OShr; [IWP]; PMBC
Leiognathus equulus (Forsskål, 1775): Mg, Sg, OShr;
[IWP]; PMBC
Leiognathus fasciatus (Lacepède, 1803): Mg, OShr; [IWP];
PMBC
Leiognathus longispinis (Valenciennes, 1835): Mg, Sg,
OShr; [EIO-IA]; PMBC
Nuchequula blochii (Valenciennes, 1835): Mg, OShr; [NIOWP]; PMBC
Nuchequula gerroides (Bleeker, 1851): Mg, Sg, OShr; [EIOWP]; PMBC
Photopectoralis bindus (Valenciennes, 1835): Mg, OShr;
[IWP]; PMBC
Secutor hanedai Moshizuki & Hayashi, 1989: Mg, Sg,
OShr; [IM]; PMBC
Secutor indicius Monkolprasit, 1973: OShr; [IM]; PMBC
Secutor interruptus (Valenciennes, 1835): Mg, OShr; [EIOWP]
Secutor insidiator (Bloch, 1787): Mg, Sg, OShr; [IWP];
PMBC
Secutor megalolepis Moshizuki & Hayashi, 1989: Mg, Sg,
OShr; [IA]; PMBC
Bramidae
Brama dussumieri Cuvier, 1831: OShr; [CIR]; PMBC
Lutjanidae
Aphareus furca (Lacepède, 1801): OShr; [IP]; PMBC
Aphareus rutilans Cuvier, 1830: Cr, OShr; [IP]; PMBC
Aprion virescens Valenciennes, 1830: Cr, OShr; [IP];
PMBC
Lipocheilus carnolabrum (Chan, 1970): OShr; [NIO-WP];
PMBC
Lutjanus argentimaculatus (Forsskål, 1775): Cr, Mg, Sg,
OShr; [IP]; PMBC
Lutjanus bengalensis (Bloch, 1790): Cr, OShr; [IO]
Lutjanus biguttatus (Valenciennes 1830): Cr, Mg, Sg, OShr;
[EIO-IA]; PMBC
Lutjanus bohar (Forsskål, 1775): Cr, OShr; [IP]; PMBC
Lutjanus decussatus (Cuvier, 1828): Cr, OShr; [EIO-IA];
PMBC
Lutjanus ehrengergii (Peters, 1869): OShr; [IWP]
Lutjanus erythropterus Bloch, 1790: Cr, OShr; [NIO-IA]
Lutjanus fulviflamma (Forsskål, 1775): Cr, Mg, Sg, OShr;
[IWP]; PMBC
Lutjanus fulvus (Schneider, 1801): Cr, OShr; [IP]; PMBC
Lutjanus gibbus (Forsskål, 1775): Cr, OShr; [IP]; PMBC
Lutjanus johnii (Bloch, 1792): Cr, Mg, Sg, OShr; [IWP];
PMBC
Lutjanus kasmira (Forsskål, 1775): Cr, OShr; [IP]; PMBC
Lutjanus lemniscatus (Valenciennes, 1828): Cr, Mg, OShr;
[EIO-IA]; PMBC
Lutjanus lunulatus (Park, 1797): Cr, Mg, Sg; [NIO-IA];
PMBC
Lutjanus lutjanus Bloch, 1790: Cr, OShr; [IWP]; PMBC
Lutjanus madras (Valenciennes, 1831): Cr, Mg, OShr;
[IWP]; PMBC
Lutjanus malabaricus Schneider, 1801: Cr, OShr; [NIOWP]; PMBC
Lutjanus monostigma (Cuvier, 1828): Cr, OShr; [IP];
PMBC
Lutjanus quinquelineatus Bloch, 1790: Cr, OShr; [NIOWP]; PMBC
Lutjanus rivulatus (Cuvier, 1828): Cr, OShr; [IP]; PMBC
Lutjanus rufolineatus (Valenciennes, 1830): OShr; [WP]
Lutjanus russelli (Bleeker, 1849): Cr, Mg, Sg, OShr; [IWP];
PMBC
Lutjanus sebae (Cuvier, 1828): Cr, OShr; [IWP]; PMBC
Lutjanus timorensis (Quoy & Gaimard, 1824): OShr; [WP]
Lutjanus vitta (Quoy & Gaimard, 1824): Cr, Sg, OShr;
[IWP]; PMBC
Macolor macularis Fowler, 1931: Cr, OShr; [EIO-WP]
Macolor niger (Forsskål, 1775): Cr, OShr; [IP]; PMBC
Paracaesio xanthurus (Bleeker, 1869): OShr; [INP]
Pinjalo lewisi Randall, Allen & Anderson, 1987: Cr; [NIOWP]
Pinjalo pinjalo (Bleeker, 1850): OShr; [IWP]; PMBC
Pristipomoides flavipinnis Shinohara, 1963: OShr; [WCP]
Pristipomoides multidens (Day, 1871): OShr; [IWP]; PMBC
Pristipomoides typus Bleeker, 1852: OShr; [IA]; PMBC
Symphorichthys spilurus (Günther, 1874): Cr, OShr; [WP]
Symphorus nematophorus (Bleeker, 1860): Cr, Sg, OShr;
[WP]; PMBC
Caesionidae
Caesio caerulaurea Lacepède, 1801: Cr, Mg, Sg, OShr;
[IP]; PMBC
Caesio cuning (Bloch, 1791): Cr, Sg, OShr; [EIO-WP];
PMBC
Caesio lunaris Cuvier, 1830: Cr, OShr; [IWP]; PMBC
Caesio teres Seale, 1906: Cr, OShr; [IP]
Caesio varilineata Carpenter, 1987: Cr, OShr; [IWP]
Caesio xanthonota Bleeker, 1853: Cr, OShr; [IO]; PMBC
Dipterygonotus balteatus (Valenciennes, 1830): Cr, OShr;
[IWP]; PMBC
Gymnocaesio gymnoptera (Bleeker, 1856): Cr; [IWP];
PMBC
Pterocaesio chrysozona (Cuvier, 1830): Cr, OShr; [IWP];
PMBC
Pterocaesio pisang (Bloch, 1853): Cr, OShr; [IWP]; PMBC
Pterocaesio randalli Carpenter, 1985: Cr; [EIO-IM]
Pterocaesio tessellata Carpenter, 1987: Cr, OShr; [EIOIA]; PMBC
59
The fishes of southwestern Thailand
Pterocaesio tile (Cuvier, 1830): Cr, OShr; [IP]; PMBC
Lobotidae
Lobotes surinamensis (Bloch, 1790): Mg, Sg, OShr; [COS];
PMBC
Gerreidae
Gerres erythrourus (Bleeker, 1791): Mg, Sg, OShr; [EIOWP]; PMBC
Gerres filamentosus (Cuvier, 1829): Cr, Mg, Sg, OShr;
[IP]; PMBC
Gerres limbatus Cuvier, 1830: Mg, OShr; [EIO-WP];
PMBC
Gerres longirostris (Lacepède, 1801): Cr, Mg, OShr; [IWP]
Gerres lucidus (Cuvier, 1830): Cr; [EIO-WP]
Gerres macracanthus Bleeker, 1854: Mg, Sg, OShr; [IWP];
PMBC
Gerres oblongus (Cuvier, 1830): Cr, Mg, Sg, OShr; [IP];
PMBC
Gerres oyena (Forsskål, 1775): Cr, Mg, Sg, OShr; [IWP];
PMBC
Gerres phaiya Iwatsuki & Heemstra, 2001: Mg, Sg, OShr;
[EIO]; PMBC
Gerres setifer (Hamilton, 1822): OShr; [EIO]
Gerres shima Iwatsuki, Kimura & Yoshino, 2007: OShr;
[WP]
Pentaprion longimanus (Cantor, 1850): OShr; [EIO-WP];
PMBC
Haemulidae
Diagramma pictum pictum (Thunberg, 1792): Cr, Sg, OShr;
[WP]; PMBC
Hapalogenys merguiensis Iwatsuki, Satapoomin &
Amaoka, 2000: OShr; [AS]; PMBC
Plectorhinchus chaetodonoides Lacepède, 1800: Cr, OShr;
[IWP]; PMBC
Plectorhinchus gibbosus (Lacepède, 1802): Cr, Mg, Sg,
OShr; [IP]; PMBC
Plectorhinchus macrospilus Satapoomin & Randall, 2000:
Cr; [AS]; PMBC
Plectorhinchus vittatus (Linnaeus, 1758): Cr, OShr; [IWP]
Pomadasys andamanensis Mckay & Satapoomin, 1994:
Cr; [AS]; PMBC
Pomadasys argenteus (Forsskål, 1775): OShr; [IWP];
PMBC
Pomadasys argyreus (Valenciennes, 1833): Mg, OShr;
[NIO-IM]; PMBC
Pomadasys auritus (Cuvier, 1830): OShr; [IA]
Pomadasys furcatus (Bloch & Schneider, 1801): OShr;
[IM]; PMBC
Pomadasys kaakan (Cuvier, 1830): Mg, Sg, OShr; [IWP];
PMBC
Pomadasys maculatus (Bloch, 1793): OShr; [IWP]; PMBC
Pomadasys olivaceus (Day, 1875): OShr; [IO]
Pomadasys unimaculatus Tian, 1982: Mg, Sg, OShr; [WP];
PMBC
Nemipteridae
Nemipterus balinensoides (Popta, 1918): OShr; [WP]
Nemipterus bathybius Snyder, 1911: OShr; [IA]; PMBC
Nemipterus bipunctatus (Ehrenberg, 1830): Cr, OShr; [IO];
PMBC
Nemipterus furcosus (Valenciennes, 1830): Cr, OShr; [EIOWP]; PMBC
Nemipterus hexodon (Quoy & Gaimard, 1824): OShr; [WP];
PMBC
Nemipterus japonicus (Bloch, 1791): OShr; [IWP]; PMBC
Nemipterus marginatus (Valenciennes, 1830): OShr; [WP]
Nemipterus mesoprion (Bleeker, 1853): OShr; [IM];
PMBC
Nemipterus nematophorus (Bleeker, 1853): OShr; [EIOIM]; PMBC
Nemipterus peronii (Valenciennes, 1830): Cr, Mg, OShr;
[IWP]; PMBC
Nemipterus tambuloides (Bleeker, 1853): OShr; [IM];
PMBC
Nemipterus thosaporni Russell, 1991: OShr; [IM]; PMBC
Nemipterus zysron (Bleeker, 1856-57): Cr, OShr; [IWP];
PMBC
Parascolopsis aspinosa (Rao & Rao, 1981): OShr; [NIO]
Parascolopsis eriomma (Jordan & Richardson, 1909):
OShr; [IWP]; PMBC
Parascolopsis inermis (Schlegel, 1843): OShr; [EIO-WP];
PMBC
Scolopsis affinis Peters, 1877: Cr, Sg, OShr; [WP]; PMBC
Scolopsis bilineatus (Bloch, 1793): Cr, Sg, OShr; [EIOWP]; PMBC
Scolopsis ciliatus (Lacepède, 1802): Cr, Mg, Sg, OShr;
[WP]; PMBC
Scolopsis lineatus Quoy & Gaimard, 1824: Cr, OShr;
[WCP]; PMBC
Scolopsis margaritifer (Cuvier, 1830): Cr, OShr; [WP]
Scolopsis monogramma (Cuvier, 1830): Cr, Sg, OShr;
[WP]; PMBC
Scolopsis taeniopterus (Valenciennes, 1830): Mg, Sg, OShr;
[IA]; PMBC
Scolopsis vosmeri (Bloch, 1792): Cr, OShr; [IWP]; PMBC
Scolopsis xenochrous Günther, 1872: Cr, OShr; [EIO-WP]
Lethrinidae
Gnathodentex aurolineatus (Lacepède, 1802): Cr; [IP]
Gymnocranius elongatus Senta, 1973: OShr; [IWP]
Gymnocranius grandoculis (Valenciennes, 1830): Cr, OShr;
[IP]; PMBC
Gymnocranius griseus (Schlegel, 1844): Cr, OShr; [EIOWP]; PMBC
Gymnocranius microdon (Bleeker, 1851): Cr, OShr; [WCP]
Lethrinus conchyliatus (Smith, 1959): OShr; [IO]
Lethrinus crocineus Smith, 1959: Cr, OShr; [IO]; PMBC
Lethrinus erythracanthus Cuvier, 1830: Cr, OShr; [IP]
Lethrinus erythropterus Valenciennes, 1830: Cr, OShr;
[IWP]
Lethrinus harak (Forsskål, 1775): Cr, OShr; [IP]; PMBC
Lethrinus lentjan (Lacepède, 1802): Cr, Sg, OShr; [IWP];
PMBC
Lethrinus mahsena (Forsskål, 1775): Cr; [IO]; PMBC
Lethrinus microdon Valenciennes, 1830: Cr, OShr; [IWP]
60
Phuket mar. biol. Cent. Res. Bull.
Lethrinus nebulosus (Forsskål, 1775): Cr, OShr; [IP];
PMBC
Lethrinus obsoletus (Forsskål, 1775): Cr, OShr; [IP]
Lethrinus olivaceus Valenciennes, 1830: Cr, OShr; [IP]
Lethrinus ornatus Valenciennes, 1830: Cr, Sg, OShr; [EIOWP]; PMBC
Lethrinus rubrioperculatus Sato, 1978: Cr; [IP]; PMBC
Lethrinus variegatus Valenciennes, 1830: OShr; [IWP];
PMBC
Lethrinus xanthochilus Klunzinger, 1870: Cr, OShr; [IP]
Monotaxis grandoculis (Forsskål, 1775): Cr, OShr; [IP];
PMBC
Wattsia mossambrica (Smith, 1957): OShr; [IWP]; PMBC
Sparidae
Acanthopagrus berda (Forsskål, 1775): Mg, OShr; [IWP]
Argyrops spinifer (Forsskål, 1775): Cr, OShr; [IWP]; PMBC
Polynemidae
Eleutheronema tetradactylum (Shaw, 1804): Mg, OShr;
[NIO-IA]; PMBC
Eleutheronema tridactylum (Bleeker, 1849): OShr; [IM]
Filimanus perplexa Feltes, 1991: OShr; [EEIO]; PMBC
Filimanus similis Feltes, 1991: OShr; [NIO]
Filimanus xanthonema (Valenciennes, 1831): OShr; [EEIO]
Leptomelanosoma indicum (Shaw, 1804): Mg, OShr; [NIOIM]; PMBC
Polydactylus microstomus (Bleeker, 1851): Mg, OShr; [EIOWP]; PMBC
Polydactylus plebeius (Broussonet, 1782): Mg, OShr; [IWP];
PMBC
Polydactylus sexfilis (Valenciennes, 1831): OShr; [IP]
Polydactylus sextarius (Bloch, 1801): OShr; [EIO-WP]
Polydactylus siamensis Motomura, Iwatsuki & Yoshino,
2001: OShr; [IM (presently known from west coast of
Thailand and Gulf of Thailand)]; PMBC
Sciaenidae
Dendrophysa russelli (Cuvier, 1830): Mg, Sg, OShr; [EIOIM]; PMBC
Johnius amblycephalus (Bleeker, 1855): OShr; [NIO-IA];
PMBC
Johnius belangerii (Cuvier, 1830): Mg, OShr; [NIO-IM]
Johnius borneensis (Bleeker, 1851): OShr; [NIO-IA]
Johnius carutta Bloch, 1793: OShr; [NIO]; PMBC
Johnius coitor (Hamilton, 1822): OShr: [EIO-IM]
Johnius dussumieri (Cuvier, 1830): OShr; [NIO]
Johnius latifrons Sasaki, 1992: OShr; [IM]
Macrospinosa cuja (Hamilton, 1822): OShr; [EIO]
Nibea soldado (Lacepède, 1802): Mg, OShr; [EIO-IA]
Otolithes cuvieri Trewavas, 1974: Mg, OShr; [IO]
Otolithes ruber (Bloch & Schneider, 1801): OShr; [IWP]
Otolithoides biauritus (Cantor, 1850): OShr; [EIO-IM]
Panna microdon (Bleeker, 1849): OShr; [IM]
Pennahia anea (Bloch, 1793): Mg, Sg, OShr; [NIO-IM];
PMBC
Protonibea diacanthus (Lacepède, 1802): OShr; [NIO-IA];
PMBC
Mullidae
Mulloidichthys flavolineatus (Lacepède, 1801): Cr, Sg, OShr;
[IP]
Mulloidichthys vanicolensis (Valenciennes, 1831): Cr, OShr;
[IP]; PMBC
Parupeneus barberinus (Lacepède, 1801): Cr, OShr; [IP];
PMBC
Parupeneus cyclostomus (Lacepède, 1801): Cr, OShr; [IP];
PMBC
Parupeneus heptacanthus (Lacepède, 1801): Cr, Sg, OShr;
[IWP]; PMBC
Parupeneus indicus (Shaw, 1803): Cr, Sg, OShr; [IWP];
PMBC
Parupeneus macronema (Lacepède, 1801): Cr, OShr; [IWP];
PMBC
Parupeneus pleurostigma (Bennett, 1830): Cr, OShr; [IP];
PMBC
Parupeneus trifasciatus Lacepède, 1801: Cr, OShr; [IO]
Upeneus asymmetricus Lachner, 1954: OShr; [IWP];
PMBC
Upeneus guttatus (Day, 1868): OShr; [IWP]
Upeneus moluccensis (Bleeker, 1855): OShr; [IWP]
Upeneus sulphureus Cuvier, 1829: Mg, Sg, OShr; [IWP];
PMBC
Upeneus sundaicus (Bleeker, 1855): Mg, OShr; [NIO-IA];
PMBC
Upeneus tragula Richardson, 1846: Cr, Mg, Sg, OShr;
[IWP]; PMBC
Upeneus vittatus (Forsskål, 1775): OShr; [IP]; PMBC
Pempheridae
Parapriacanthus ransonneti Steindachner, 1870: Cr; [WP];
PMBC
Pempheris mangula Cuvier, 1829: Cr; [IWP]; PMBC
Pempheris otaitensis Cuvier, 1831: Cr; [IWP]
Pempheris vanicolensis Cuvier, 1831: Cr; [IM]; PMBC
Monodactylidae
Monodactylus argenteus (Linnaeus, 1758): Cr, Mg, OShr;
[IWP]; PMBC
Toxotidae
Toxotes chatareus (Hamiton, 1822): Mg; [EIO-IA]; PMBC
Toxotes jaculatrix (Pallas, 1766): Mg; [EIO-WP]; PMBC
Kyphosidae
Kyphosus cinerascens (Forsskål, 1775): Cr; [IP]; PMBC
Kyphosus vaigiensis (Quoy & Gaimard, 1825): Cr; [IP];
PMBC
Drepaneidae
Drepane longimana (Bloch & Schneider, 1801): Mg, Sg,
OShr; [IWP]; PMBC
Drepane punctata (Linnaeus, 1758): Mg, Sg, OShr; [IWP];
PMBC
Chaetodontidae
Chaetodon andamanensis Kuiter & Debelius, 1999: Cr;
[EIO]; PMBC
Chaetodon auriga Forsskål, 1775: Cr; [IP]; PMBC
Chaetodon bennetti Cuvier, 1831: Cr; [IP]
Chaetodon citrinellus Cuvier, 1831: Cr; [IP]; PMBC
Chaetodon collare Bloch, 1787: Cr; [NIO]; PMBC
61
The fishes of southwestern Thailand
Chaetodon decussatus Cuvier, 1831: Cr; [EIO]; PMBC
Chaetodon ephippium Cuvier, 1831: Cr; [IP]; PMBC
Chaetodon falcula Bloch, 1793: Cr; [IO]; PMBC
Chaetodon gardineri Norman, 1939: Cr; [IO]; PMBC
Chaetodon guttatissimus Bennett, 1832: Cr; [IO]; PMBC
Chaetodon interruptus Ahl, 1922: Cr; [IO]
Chaetodon kleinii Bloch, 1790: Cr; [IP]; PMBC
Chaetodon lineolatus Cuvier, 1831: Cr; [IP]; PMBC
Chaetodon lunula (Lacepède, 1803): Cr; [IP]; PMBC
Chaetodon madagaskariensis Ahl, 1923: Cr; [IO]; PMBC
Chaetodon melannotus Bloch & Schneider, 1801: Cr; [IP];
PMBC
Chaetodon meyeri Bloch & Schneider, 1801: Cr; [IP];
PMBC
Chaetodon octofasciatus Bloch, 1787: Cr; [EIO-WP];
PMBC
Chaetodon oxycephalus Bleeker, 1853: Cr; [EIO-WP]
Chaetodon rafflesi Bennett, 1830: Cr; [EIO-WCP]; PMBC
Chaetodon semeion Bleeker, 1855: Cr; [EIO-WCP]
Chaetodon triangulum Cuvier, 1831: Cr; [IO]; PMBC
Chaetodon trifascialis Quoy & Gaimard, 1825: Cr; [IP];
PMBC
Chaetodon trifasciatus Park, 1797: Cr; [IO]; PMBC
Chaetodon vagabundus Linnaeus, 1758: Cr; [IP]; PMBC
Chaetodon xanthocephalus Bennett, 1832: Cr; [IO]; PMBC
Chelmon rostratus (Linnaeus, 1758): Cr; [WP]; PMBC
Coradion altivelis Mcculloch, 1916: Cr; [WP]
Coradion chrysozonus (Cuvier, 1831): Cr, OShr; [WP];
PMBC
Forcipiger flavissimus Jordan & Mcgregor, 1898: Cr; [IP];
PMBC
Forcipiger longirostris (Broussonet, 1782): Cr; [IP]
Hemitaurichthys zoster (Bennett, 1831): Cr; [IO]; PMBC
Heniochus acuminatus (Linnaues, 1758): Cr, Mg, Sg, OShr;
[IP]; PMBC
Heniochus diphreutes Jordan, 1903: Cr, OShr; [IP]
Heniochus pleurotaenia Ahl, 1923: Cr; [EIO]; PMBC
Heniochus singularius Smith & Radcliffe, 1911: Cr; [EIOWCP]; PMBC
Parachaetodon ocellatus (Cuvier, 1831): Cr, Sg: [EIO-WP];
PMBC
Pomacanthidae
Apolemichthys trimaculatus (Lacapède, 1831): Cr; [IWP];
PMBC
Apolemichthys xanthurus (Bennett, 1832): Cr; [IO]; PMBC
Centropyge eibli Klausewitz, 1963: Cr; [EIO]; PMBC
Centropyge flavipectoralis Randall & Klausewitz, 1977:
Cr; [EIO]
Centropyge multispinis (Playfair, 1866): Cr; [IO]; PMBC
Chaetodontoplus mesoleucus (Bloch, 1787): Cr; [WP]
Genicanthus caudovittatus (Günther, 1860): Cr, OShr; [IO];
PMBC
Pomacanthus annularis (Bloch, 1787): Cr, OShr; [IWP];
PMBC
Pomacanthus imperator (Bloch, 1787): Cr; [IP]; PMBC
Pomacanthus navarchus (Cuvier, 1831): Cr; [IA]
Pomacanthus semicirculatus (Cuvier, 1831): Cr; [IP]
Pomacanthus sexstriatus (Cuvier, 1831): Cr; [WP]
Pomacanthus xanthometopon (Bleeker, 1853): Cr; [EIOWCP]
Pygoplites diacanthus (Boddaert, 1772): Cr; [IP]; PMBC
Terapontidae
Pelates quadrilineatus (Bloch, 1790): Mg, Sg; [IWP];
PMBC
Pseudoterapon theraps (Cuvier, 1829): Mg, Sg, OShr;
[IWP]; PMBC
Terapon jarbua (Forsskål, 1775): Cr, Mg, Sg, OShr; [IWP];
PMBC
Terapon puta (Cuvier, 1829): Mg, Sg, OShr; [IWP]; PMBC
Kuhliidae
Kuhlia mugil (Bloch & Schneider, 1801): Cr; [IP]; PMBC
Cirrhitidae
Cirrhitichthys falco Randall, 1963: Cr; [EIO-WP]
Cirrhitichthys oxycephalus (Bleeker, 1855): Cr; [IP];
PMBC
Oxycirrhites typus Bleeker, 1857: Cr; [IP]
Paracirrhites arcatus (Cuvier, 1829): Cr; [IP]
Paracirrhites fosteri (Schneider, 1801): Cr; [IP]; PMBC
Cepolidae
Acanthocepola abbreviata (Valenciennes, 1835): OShr;
[NIO-IM]; PMBC
Acanthocepola indica (Day, 1888): OShr; [IWP]
Pomacentridae
Abudefduf bengalensis (Bloch, 1787): Cr; [EIO-WP];
PMBC
Abudefduf notatus (Day, 1869): Cr; [IWP]; PMBC
Abudefduf septemfasciatus (Cuvier, 1830): Cr; [IP]; PMBC
Abudefduf sordidus (Forsskål, 1775): Cr; [IP]; PMBC
Abudefduf vaigiensis (Quoy & Gaimard, 1825): Cr; [IP];
PMBC
Amblyglyphidodon aureus (Cuvier, 1830): Cr; [WCP];
PMBC
Amblyglyphidodon indicus Allen & Randall, 2002: Cr; [IO]
Amblyglyphidodon leucogaster (Bleeker, 1847): Cr; [WP];
PMBC
Amphiprion akallopisos Bleeekr, 1853: Cr; [IO]; PMBC
Amphiprion clarkii (Bennett, 1830): Cr, Sg; [IWP]; PMBC
Amphiprion ephippium (Bloch, 1790): Cr; [EIO]; PMBC
Amphiprion ocellaris Cuvier, 1830: Cr; [WP]; PMBC
Amphiprion sebae Bleeekr, 1853: Cr, Sg; [EIO]; PMBC
Cheiloprion labiatus (Day, 1877): Cr; [IA]; PMBC
Chromis atripectoralis Welander & Schultz, 1951: Cr;
[IWP]; PMBC
Chromis cinerascens (Cuvier, 1830): Cr; [EIO-IA]; PMBC
Chromis delta Randall, 1987: Cr; [EIO-WP]
Chromis dimidiata (Kluzinger, 1871): Cr; [IO]; PMBC
Chromis elerae Fowler & Bean, 1928: Cr; [EIO-WP]
Chromis flavipectoralis Randall, 1990: Cr; [EIO]; PMBC
Chromis lepidolepis Bleeker, 1877: Cr; [IP]
Chromis opercularis (Günther, 1866): Cr; [IO]; PMBC
Chromis ternatensis (Bleeker, 1856): Cr; [IP]; PMBC
Chromis viridis (Cuvier, 1830): Cr; [IP]; PMBC
62
Phuket mar. biol. Cent. Res. Bull.
Chromis weberi Fowler & Bean, 1928: Cr; [IP]; PMBC
Chromis xouthos Allen & Erdmann, 2005: Cr; [AS]
Chrysiptera biocellata (Quoy & Gaimard, 1825): Cr; [IP];
PMBC
Chrysiptera brownriggii (Bennett, 1828): Cr; [IP]; PMBC
Chrysiptera glauca (Cuvier, 1830): Cr; [IP]
Chrysiptera rollandi (Whitley, 1961): Cr; [IA]; PMBC
Chrysiptera unimaculata (Cuvier, 1830): Cr; [IWP]; PMBC
Dascyllus aruanus (Linnaeus, 1758): Cr; [IP]; PMBC
Dascyllus carneus Fischer, 1885: Cr; [IO]; PMBC
Dascyllus trimaculatus (Rüppell, 1828): Cr; [IP]; PMBC
Dischistodus perspicillatus (Cuvier, 1830): Cr; [WP];
PMBC
Hemiglyphidodon plagiometopon (Bleeker, 1852): Cr; [IA];
PMBC
Lepidozygus tapeinosoma (Bleeker, 1856): Cr; [IP]; PMBC
Neoglyphidodon melas (Cuvier, 1830): Cr; [IWP]; PMBC
Neoglyphidodon nigroris (Cuvier, 1830): Cr; [WP]; PMBC
Neopomacentrus anabatoides (Bleeker, 1847): Cr; [IA];
PMBC
Neopomacentrus bankieri (Richardson, 1846): Cr; [WP]
Neopomacentrus cyanomos (Bleeker, 1856): Cr; [IWP];
PMBC
Neopomacentrus filamentosus (Macleay, 1883): Cr; [IA];
PMBC
Neopomacentrus sororius Randall & Allen, 2005: Cr; [IO];
PMBC
Neopomacentrus taeniurus (Bleeker, 1856): Cr; [IWP];
PMBC
Neopomacentrus violascens (Bleeker, 1848): Cr; [IA]
Plectroglyphidodon dickii (Leinard, 1839): Cr; [IP]; PMBC
Plectroglyphidodon johnstonianus Fowler & Ball, 1924:
Cr; [IP]; PMBC
Plectroglyphidodon lacrymatus (Quoy & Gaimard, 1825):
Cr; [IP]; PMBC
Plectroglyphidodon leucozona (Bleeker, 1859): Cr; [IP];
PMBC
Pomacentrus adelus Allen, 1991: Cr; [IA]; PMBC
Pomacentrus alleni Burgess, 1981: Cr; [EEIO]; PMBC
Pomacentrus amboinensis Bleeker, 1868: Cr; [WCP];
PMBC
Pomacentrus azuremaculatus Allen, 1991: Cr; [EEIO];
PMBC
Pomacentrus chrysurus Cuvier, 1830: Cr; [EIO-WP];
PMBC
Pomacentrus lepidogenys Fowler & Bean, 1928: Cr; [WP];
PMBC
Pomacentrus moluccensis Bleeker, 1853: Cr; [WP]; PMBC
Pomacentrus nagasakiensis Tanaka, 1917: Cr; [EIO-WP];
PMBC
Pomacentrus pavo (Bloch, 1787): Cr; [IP]; PMBC
Pomacentrus philippinus Evermann & Seale, 1907: Cr;
[EIO-WP]; PMBC
Pomacentrus polyspinus Allen, 1991: Cr; [AS]; PMBC
Pomacentrus similis Allen, 1991: Cr; [EIO]; PMBC
Pomacentrus tripunctatus Cuvier, 1830: Cr; [EIO-WP];
PMBC
Pomacentrus xanthosternus Allen, 1991: Cr; [EEIO];
PMBC
Premnas biaculeatus (Bloch, 1790): Cr; [IA]
Pristotis obtusirostris (Günther, 1862): Cr, OShr; [IWP];
PMBC
Stegastes fasciolatus (de Vis, 1884): Cr; [IP]; PMBC
Stegastes nigricans (Lacepède, 1803): Cr; [IP]; PMBC
Stegastes obreptus (Whitley, 1948): Cr; [EIO-WP]; PMBC
Stegastes punctatus (Quoy & Gaimard, 1825): Cr; [IWP];
PMBC
Teixeirichthys jordani (Rutter, 1897): OShr; [IWP]
Labridae
Anampses caeruleopunctatus Rüppell, 1829: Cr; [IP];
PMBC
Anampses lineatus Randall, 1972: Cr; [IO]; PMBC
Anampses meleagrides Valenciennes, 1840: Cr; [IWP];
PMBC
Anampses twistii Bleeker, 1856: Cr; [IP]; PMBC
Bodianus axillaris (Bennett, 1832): Cr; [IP]
Bodianus diana (Lacepède, 1801): Cr; [IO]; PMBC
Bodianus mesothorax (Bloch & Schneider, 1801): Cr; [WP];
PMBC
Bodianus neilli (Day, 1867): Cr; [EIO]; PMBC
Cheilinus chlorourus (Bloch, 1791): Cr, OShr; [IP]; PMBC
Cheilinus fasciatus (Bloch, 1791): Cr; [IP]; PMBC
Cheilinus oxycephalus Bleeker, 1853: Cr; [IP]
Cheilinus trilobatus Lacepède, 1801: Cr; [IP]; PMBC
Cheilinus undulatus Rüppell, 1835: Cr; [IP]
Choerodon robustus (Günther, 1862): OShr; [IWP];
PMBC
Cirrhilabrus cyanopleura (Bleeker, 1851): Cr; [EEIO];
PMBC
Cirrhilabrus exquisitus Smith, 1957: Cr; [IP]; PMBC
Cirrhilabrus joanallenae Allen, 2000: Cr; [AS]
Coris batuensis (Bleeker, 1856): Cr; [EIO-WCP]; PMBC
Coris cuvieri (Bennett, 1831): Cr; [IO]; PMBC
Diproctacanthus xanthurus (Bleeker, 1856): Cr; [IA]
Epibulus insidiator (Pallas, 1770): Cr; [IP]; PMBC
Gomphosus caeruleus Lacepède, 1801: Cr; [IO]; PMBC
Halichoeres argus (Bloch, 1791): Cr; [WP]; PMBC
Halichoeres bicolor (Bloch & Schneider, 1801): Cr, Mg,
Sg; [EIO-IM]; PMBC
Halichoeres chloropterus (Bloch, 1791): Cr; [IA]; PMBC
Halichoeres cosmetus Randall & Smith, 1982: Cr; [IO]
Halichoeres hortulanus (Lacepède, 1801): Cr; [IP]; PMBC
Halichoeres kallochroma (Bleeker, 1835): Cr; [AS];
PMBC
Halichoeres leucoxanthus Randall & Smith, 1982: Cr;
[EIO]; PMBC
Halichoeres margaritaceus (Valenciennes, 1839): Cr;
[WCP]
Halichoeres marginatus Rüppell, 1835: Cr; [IP]; PMBC
Halichoeres nebulosus (Valenciennes, 1839): Cr; [IWP];
PMBC
Halichoeres nigrescens (Bloch & Schneider, 1801): Cr, Sg;
[IWP]; PMBC
Halichoeres scapularis (Bennett, 1832): Cr; [IWP]; PMBC
63
The fishes of southwestern Thailand
Halichoeres timorensis (Bleeker, 1852): Cr; [EIO-IM];
PMBC
Halichoeres vrolikii (Bleeker, 1855): Cr; [EIO]; PMBC
Halichoeres zeylonicus (Bennett, 1832): Cr, OShr; [IO];
PMBC
Hemigymnus fasciatus (Bloch, 1792): Cr; [IP]; PMBC
Hemigymnus melapterus (Bloch, 1791): Cr; [IP]; PMBC
Hologymnosus annulatus (Lacepède, 1801): Cr; [IP];
PMBC
Hologymnosus doliatus (Lacepède, 1801): Cr; [IP]
Iniistius baldwini (Jordan & Evermann, 1903): OShr; [IP];
PMBC
Iniistius bimaculatus (Rüppell, 1829): OShr; [IO]; PMBC
Iniistius pavo (Valenciennes, 1840): Cr, OShr; [IP]; PMBC
Labrichthys unilineatus (Guichenot, 1847): Cr; [IP]; PMBC
Labroides bicolor Fowler & Bean, 1928: Cr; [IP]
Labroides dimidiatus (Valenciennes, 1839): Cr; [IP];
PMBC
Labropsis xanthonota Randall, 1981: Cr; [IWP]
Leptojulis chrysotaenia Randall & Ferraris, 1981: Cr; [EIO]
Leptojulis cyanopleura (Bleeker, 1853): Cr, OShr; [NIOWP]; PMBC
Leptojulis urostigma Randall, 1996: OShr; [IA]
Macropharyngodon ornatus Randall, 1978: Cr; [IWP];
PMBC
Novaculichthys taeniourus (Lacepède, 1801): Cr; [IP]
Oxycheilinus areanatus (Valenciennes, 1840): Cr; [IP]
Oxycheilinus bimaculatus (Valenciennes, 1840): Cr; [IP]
Oxycheilinus celebicus (Bleeker, 1853): Cr; [WP]; PMBC
Oxycheilinus digramma (Lacepède, 1801): Cr, OShr; [IP];
PMBC
Oxycheilinus orientalis (Günther, 1862): Cr; [IWP]
Paracheilinus mccoskeri Randall & Harmelin-Vivien, 1977:
Cr; [IO]; PMBC
Pseudocheilinus evanidus Jordan & Evermann, 1903: Cr;
[IP]
Pseudocheilinus hexataenia (Bleeker, 1857): Cr; [IP];
PMBC
Pseudodax moluccanus (Valenciennes, 1840): Cr; [IP];
PMBC
Pseudojuloides kaleidos Kuiter & Randall, 1995: Cr; [EIO];
PMBC
Pteragogus cryptus Randall, 1981: Cr; [IWP]; PMBC
Stethojulis albovittata (Bonnaterre, 1788): Cr; [IO]; PMBC
Stethojulis bandanensis (Bleeker, 1851): Cr; [WCP]
Stethojulis interrupta (Bleeker, 1851): Cr; [IWP]; PMBC
Stethojulis strigiventer (Bennett, 1832): Cr, Sg; [IP]
Stethojulis trilineata (Bloch & Schneider, 1801): Cr; [EIOWCP]; PMBC
Suezichthys caudovittatus (Steindachner, 1898): OShr; [IO];
PMBC
Thalassoma amblycephalum (Bleeker, 1856): Cr; [IP];
PMBC
Thalassoma hardwicke (Bennett, 1830): Cr; [IP]; PMBC
Thalassoma jansenii (Bleeker, 1856): Cr; [EIO-WP];
PMBC
Thalassoma lunare (Linnaeus, 1758): Cr; [IP]; PMBC
Thalassoma purpureum (Forsskål, 1775): Cr; [IP]
Thalassoma trilobatum (Lacepède, 1801): Cr; [IP]; PMBC
Wetmorella nigropinnata (Seale, 1901): Cr; [IP]; PMBC
Xiphocheilus quadrimaculatus (Günther, 1880): OShr;
[EIO-WP]
Scaridae
Bolbometopon muricatum (Valenciennes, 1840): Cr; [IP];
PMBC
Calotomus carolinus (Valenciennes, 1840): Cr; [IP]; PMBC
Cetoscarus bicolor (Rüppell, 1829): Cr; [IP]
Chlorurus capistratoides (Bleeker, 1847): Cr; [IO]; PMBC
Chlorurus rhakoura Randall & Anderson, 1997: Cr; [EIO];
PMBC
Chlorurus sordidus (Forsskål, 1775): Cr; [IP]; PMBC
Chlorurus strongylocephalus (Bleeker, 1854): Cr; [IO];
Chlorurus troschelii (Bleeker, 1853): Cr; [EEIO]
Hipposcarus harid (Forsskål, 1775): Cr; [IO]
Scarus frenatus Lacepède, 1802: Cr; [IP]; PMBC
Scarus ghobban Forsskål, 1775: Cr, Sg; [IP]; PMBC
Scarus maculipinna Westneat, Satapoomin & Randall,
2007: Cr; [EEIO]; PMBC
Scarus niger Forsskål, 1775: Cr; [IP]; PMBC
Scarus prasiognathos Valenciennes, 1840: Cr; [EIO-WP];
PMBC
Scarus quoyi Valenciennes, 1840: Cr; [EIO-WP]; PMBC
Scarus rubroviolaceus Bleeker, 1847: Cr; [IP]; PMBC
Scarus russelii Valenciennes, 1840: Cr; [IO]; PMBC
Scarus scaber Valenciennes, 1840: Cr; [IO]; PMBC
Scarus tricolor Bleeker, 1847: Cr; [IWP]; PMBC
Scarus viridifucatus (Smith, 1956): Cr; [IO]; PMBC
Champsodontidae
Champsodon nudivittis (Ogilby, 1895): OShr; [IWP]
Champsodon vorax Günther, 1867: OShr; [EIO-WP]
Pinguipedidae
Parapercis alboguttata (Günther, 1872): OShr; [NIO-IA];
PMBC
Parapercis clathrata Ogilby, 1910: Cr, OShr; [WCP];
PMBC
Parapercis cylindrica (Bloch, 1792): Cr: [EIO-WP], PMBC
Parapercis hexophthalma (Cuvier, 1829): Cr, OShr; [IWP];
PMBC
Parapercis maculata (Bloch & Schneider, 1801): OShr;
[IO]
Parapercis millepunctata (Günther, 1860): Cr, OShr; [EIOWCP]; PMBC
Parapercis punctata (Cuvier, 1829): Mg, OShr; [IO];
PMBC
Parapercis quadrispinosa (Weber, 1913): OShr; [EIO];
PMBC
Parapercis schauinslandi (Steindachner, 1900): Cr; [IP]
Parapercis snyderi Jordan & Starks, 1905: Cr, OShr; [WP]
Parapercis tetracantha (Lacepède, 1800): Cr, OShr; [IWP];
PMBC
Parapercis xanthozona (Bleeker, 1849): Cr, OShr; [IWP];
PMBC
Trichonotidae
Trichonotus setiger Bloch & Schneider, 1801: OShr; [WP]
64
Phuket mar. biol. Cent. Res. Bull.
Trichonotus sp.1: Cr, OShr
Percophidae
Bembrops caudimaculata Steindachner, 1876: OShr; [IWP]
Bembrops platyrhynchus (Alcock, 1894): OShr; [IWP];
PMBC
Ammodytidae
Bleekeria kallolepis Günther, 1862: OShr; [EIO]
Uranoscopidae
Ichthyoscopus lebeck (Schneider, 1801): OShr; [EIO];
PMBC
Uranoscopus affinis Cuvier, 1829: OShr; [IWP]; PMBC
Uranoscopus bicinctus Temminck & Schlegel, 1843: OShr;
[EIO-WP]
Uranoscopus cognatus Cantor, 1849: OShr; [EIO-WP]
Uranoscopus oligolepis Bleeker, 1878: OShr; [IO]
Tripterygiidae
Ennaepterygius fasciatus (Weber, 1909): Cr; [IP]
Ennaepterygius nanus (Schultz, 1960): Cr; [WP]
Ennaepterygius philippinus (Peters, 1868): Cr; [IWP]
Enneapterygius sp.1:
Enneapterygius tutuilae Jordan & Seale, 1906: Cr; [IP];
PMBC
Helcogramma chica Rosenblatt, 1960: Cr; [EIO-WP]
Helcogramma lacuna Williams & Howe, 2003: Cr; [AS]
Helcogramma obtusirostre Klunzinger, 1871: Cr; [IWP]
Helcogramma rosea Holleman, 2006: Cr; [EIO]
Helcogramma sp.1: Cr
Helcogramma springeri Hansen, 1986: Cr; [WP]; PMBC
Helcogramma striata Hansen, 1986: Cr; [EIO-WP]; PMBC
Norfolkia brachylepis (Schultz, 1960): Cr; [IP]
Ucla xenogrammus Holleman, 1993: Cr; [IP]; PMBC
Blenniidae
Alticus saliens (Lacepède, 1800): Cr; [IP]; PMBC
Andamia reyi Sauvage, 1880: Cr; [IM]; PMBC
Aspidontus dussumieri (Valenciennes, 1836): Cr; [IP]
Aspidontus tractus Fowler, 1903: Cr; [IO]; PMBC
Atrosalarias fuscus (Rüppell, 1835): Cr; [IO]; PMBC
Blenniella chrysospilos (Bleeker, 1857): Cr; [IP]
Blenniella cyanostigma (Bleeker, 1849): Cr; [IO]
Blenniella leopardus (Fowler, 1904): Cr; [AS]; PMBC
Blenniella periophthalmus (Valenciennes, 1836): Cr; [IWP]
Cirripectes auritus Carlson, 1980: Cr; [IP]
Cirripectes castaneus (Valenciennes, 1836): Cr; [IWP]
Cirripectes filamentosus (Alleyne & Macleay, 1877): Cr;
[IWP]; PMBC
Cirripectes stigmaticus Strasburg & Schultz, 1953: Cr;
[IWP]
Crossosalarias macrospilus Smith-Vaniz & Springer,
1971: Cr; [WP]
Ecsenius bicolor (Day, 1888): Cr; [EIO-WP]; PMBC
Ecsenius lubbocki Springer, 1988: Cr; [EEIO]; PMBC
Ecsenius midas Starck, 1969: Cr; [IP]; PMBC
Ecsenius nalolo Smith, 1959: Cr; [IO]; PMBC
Ecsenius paroculus Springer, 1988: Cr; [EEIO]; PMBC
Enchelyurus kraussi (Klunzinger, 1871): Cr; [IWP]; PMBC
Entomacrodus striatus (Quoy & Gaimard, 1836): Cr; [IP];
PMBC
Entomacrodus vermiculatus (Valenciennes, 1836): Cr; [IO];
PMBC
Exallias brevis (Kner, 1968): Cr; [IP]
Glyptoparus delicatulus Smith, 1959: Cr; [IWP]
Istiblennius bellus (Günther, 1861): Cr; [IP]; PMBC
Istiblennius dussumieri (Valenciennes, 1836): Cr; [IWP];
PMBC
Istiblennius edentulus (Schneider, 1801): Cr; [IP]; PMBC
Istiblennius lineatus (Valenciennes, 1836): Cr; [EIO-WCP]
Laiphognathus multimaculatus Smith, 1955: Cr; [IWP];
PMBC
Meiacanthus smithi Klausewitz, 1916: Cr; [EIO]; PMBC
Meiacanthus urostigma Smith-Vaniz, Satapoomin & Allen,
2001: Cr; [AS]; PMBC
Omobranchus elongatus (Peters, 1855): Cr; [IWP]; PMBC
Omobranchus fasciolatus (Valenciennes, 1836): Mg; [IO];
PMBC
Omobranchus ferox (Herre, 1927): Mg; [IP]; PMBC
Omobranchus obliquus (Garman, 1903): Cr; [WCP];
PMBC
Omobranchus punctatus (Valenciennes, 1836): Mg; [IWP];
PMBC
Omobranchus smithi (Rao, 1973): Mg; [EIO-IM]; PMBC
Omobranchus zebra (Bleeker, 1868): Mg; [EIO-IM]
Petroscirtes breviceps (Valenciennes, 1836): Cr, Mg, Sg;
[IWP]; PMBC
Petroscirtes mitratus Rüppell, 1830: Cr, Sg; [IP]; PMBC
Petroscirtes thepassi Bleeker, 1853: Sg; [WP]
Petroscirtes variabilis Cantor, 1850: Sg; [EIO-IA]; PMBC
Plagiotremus phenax Smith-Vaniz, 1976: Cr; [EIO]; PMBC
Plagiotremus rhinorhynchos (Bleeker, 1852): Cr; [IP];
PMBC
Plagiotremus tapeinosoma (Bleeker, 1857): Cr; [IP];
PMBC
Plagiotremus townsendi (Regan, 1905): Cr; [IO]
Praealticus oortii (Bleeker, 1851): Cr; [IM]
Praealticus triangulus (Chapman, 1951): Cr; [EEIO]
Salarias fasciatus (Bloch, 1786): Cr; [IWP]; PMBC
Salarias guttatus Valenciennes, 1836: Cr; [IA]; PMBC
Salarias sinuosus Snyder, 1908: Cr; [WP]; PMBC
Xiphasia setifer Swainson, 1839: OShr; [IWP]
Gobiesocidae
Diademichthys lineatus (Sauvage, 1883): Cr; [IWP]
Discotrema crinophilum Briggs, 1976: Cr; [WP]
Lepadichthys lineatus Briggs, 1966: Cr; [IWP]
Callionymidae
Bathycallionymus kaianus (Günther, 1880): OShr; [IWP]
Calliurichthys japonicus (Houttuyn, 1782): OShr; [WP];
PMBC
Dactylopus dactylopus (Valenciennes, 1837): Mg, Sg, OShr;
[WP]; PMBC
Repomucenus belcheri recurvispinnis (Li, 1966): OShr;
[EIO-IM]
Repomucenus doryssus (Jordan & Fowler, 1903): OShr;
[WP]
Repomucenus filamentosus (Valenciennes, 1837): OShr; [IP]
65
The fishes of southwestern Thailand
Repomucenus hindsi (Richardson, 1844): Sg, OShr; [NIOIM]
Repomucenus octostigmatus (Fricke, 1981): OShr; [IM]
Repomucenus sagitta (Pallas, 1770): OShr; [EIO-IM]
Repomucenus schaapii (Bleeker, 1852): Mg, Sg, OShr; [EIOIM]; PMBC
Paradiplogrammus enneactis (Bleeker, 1879): Mg, Sg;
[WP]; PMBC
Synchiropus lineolatus (Valenciennes, 1837): Cr; [IWP]
Synchiropus stellatus Smith, 1963: Cr; [IO]
Eleotridae
Bostrychus sinensis (Lacepède, 1801): Mg; [EIO-WP]
Butis butis (Hamilton, 1822): Mg, Sg; [IWP]; PMBC
Butis humeralis (Valenciennes, 1837): Mg; [EIO-WP];
PMBC
Butis koilomatodon (Bleeker, 1849): Mg, Sg; [IWP]; PMBC
Calumia godeffroyi (Günther, 1877): Cr; [IP]
Eleotris melanosoma Bleeker, 1852): Mg; [IWP]; PMBC
Odonteleotris macrodon (Bleeker, 1853): Mg; [EIO-IM];
PMBC
Ophiocara aporos (Bleeker, 1854): Mg; [IWP]; PMBC
Ophiocara porocephala (Valenciennes, 1837): Mg; [IWP];
PMBC
Xenisthmidae
Xenisthmus africanus Smith, 1958: Cr; [IO]
Xenisthmus polyzonatus (Klunzinger, 1817): Cr; [IP]
Gobiidae
Acentrogobius audax Smith, 1959: Sg; [IWP]; PMBC
Acentrogobius caninus (Valenciennes, 1837): Mg, Sg; [IWP];
PMBC
Acentrogobius chlorostigmatoides (Bleeker, 1849): OShr;
[WP]; PMBC
Acentrogobius cyanomos (Bleeker, 1849): Mg, Sg; [EIOIM]; PMBC
Acentrogobius janthinopterus (Bleeker, 1852): Mg; [WP];
PMBC
Acentrogobius madraspatensis (Day, 1868): Mg, Sg; [EIOWP]; PMBC
Acentrogobius multifasciatus (Herre, 1927): Cr, Sg; [WP];
PMBC
Acentrogobius sp.1: Sg
Acentrogobius viridipunctatus (Valenciennes, 1837): Mg;
[IWP]; PMBC
Amblyeleotris aurora (Polunin & Lubbock, 1977): Cr; [IO]
Amblyeleotris diagonalis Polunin & Lubbock, 1979: Cr;
[IP]; PMBC
Amblyeleotris downingi Randall, 1994: Cr; OShr; [NIO];
PMBC
Amblyeleotris fontanesii (Bleeker, 1852): Cr, OShr; [WP];
PMBC
Amblyeleotris gymnocephala (Bleeker, 1853): Cr; [IA];
PMBC
Amblyeleotris latifasciata Polunin & Lubbock, 1979: Cr;
[IA]; PMBC
Amblyeleotris periophthalma (Bleeker, 1853): Cr; [IWP];
PMBC
Amblyeleotris steinitzi (Klausewits, 1974): Cr; [IWP];
PMBC
Amblyeleotris triguttatus Randall, 1994: Cr; [IO]
Amblyeleotris wheeleri (Polunin & Lubbock, 1977): Cr;
[IWP]
Amblygobius hectori (Smith, 1957): Cr; [IWP]; PMBC
Amblygobius nocturnus (Herre, 1945): Cr; [NIO-WCP];
PMBC
Amblygobius semicinctus (Bennett, 1833): Cr; [IO]; PMBC
Amblygobius stethophthalmus (Bleeker, 1851): Cr; [IM]
Amoya gracilis (Bleeker, 1875): Mg; [IA]; PMBC
Amoya moloanus (Herre, 1927): Mg, Sg; [WP]; PMBC
Apocryptodon madurensis (Bleeker, 1849): Mg, Sg; [EIOWP]; PMBC
Arcygobius baliurus (Valenciennes, 1837): Sg; [IWP];
PMBC
Asterropteryx dfh sp.3: Cr
Asterropteryx ensiferus (Bleeker, 1874): Cr; [IP]; PMBC
Asterropteryx semipunctatus Rüppell, 1828: Cr; [IP];
PMBC
Barbuligobius boehlkei Lachner & McKinney, 1974: Cr;
[IWP]
Bathygobius cocosensis (Bleeker, 1854): Cr; [IP]; PMBC
Bathygobius crassiceps (Jordan & Seale, 1906): Cr; [IP];
PMBC
Bathygobius cyclopterus (Valenciennes, 1837): Cr; [IWP];
PMBC
Bathygobius fuscus (Rüppell, 1830): Cr, Sg; [IWP]; PMBC
Bathygobius karachiensis Hoda & Goren, 1990: Cr; [EIO]
Bathygobius laddi (Fowler, 1931): Cr; [IP]; PMBC
Bathygobius meggitti (Hora & Mukuji, 1936): Cr; [WP]
Boleophthalmus boddarti (Pallas, 1770): Mg; [NIO-WP];
PMBC
Brachygobius kabiliensis Inger, 1958: Mg; [IM]
Bryaninops amplus Larson, 1985: Cr; [IP]
Bryaninops erythops (Jordan & Seale, 1906): Cr; [EIOWP]
Bryaninops loki Larson, 1985: Cr; [IWP]
Bryaninops natans Larson, 1985: Cr; [IWP]; PMBC
Bryaninops ridens Smith, 1959: Cr; [IWP]; PMBC
Bryaninops youngei (Davis & Cohen, 1969): Cr; [IP]
Cabillus tongarevae (Fowler, 1927): Cr; [EIO-WP]; PMBC
Callogobius andamanensis Menon & Chatterjee, 1974: Mg;
[AS]
Callogobius dfh sp.3: Cr
Callogobius dfh sp.6: Cr
Callogobius dfh sp.11: Cr
Callogobius dfh sp.13: Cr
Callogobius dfh sp.15: Cr
Callogobius dfh sp.16: Cr
Callogobius flavobrunneus (Smith, 1958): Cr; [IO]; PMBC
Callogobius maculipinnis (Fowler, 1918): Cr; [IWP]
Callogobius plumatus (Smith, 1959): Cr; [IO]
Callogobius rw-th sp.2: Mg
Callogobius sclateri (Steindachner, 1880): Cr; [IP]; PMBC
Cristatogobius nonatoae (Ablan, 1940): Mg; [WP]; PMBC
66
Phuket mar. biol. Cent. Res. Bull.
Cryptocentrus caeruleomaculatus (Herre, 1933): Cr, Sg;
[IP]; PMBC
Cryptocentrus cyanotaenia (Bleeker, 1853): Cr, Mg, OShr;
[IA]; PMBC
Cryptocentrus fasciatus (Playfair & Gunther, 1867): Cr;
[IWP]; PMBC
Cryptocentrus leucostictus (Günther, 1871): Cr; [WCP]
Cryptocentrus maudae Fowler, 1937: Cr; [IA]; PMBC
Cryptocentrus melanopus (Bleeker, 1860): Cr, Sg, OShr;
[WP]; PMBC
Cryptocentrus pavoninoides (Bleeker, 1849): Cr, OShr;
[IA]; PMBC
Cryptocentrus sp.1 (dfh sp.27): Cr, Sg; [WP]; PMBC
Cryptocentrus strigilliceps (Jordan & Seale, 1906): Cr, Sg;
[IP]; PMBC
Ctenogobiops aurocingulus (Herre, 1935): Cr; [WCP]
Ctenogobiops crocineus Smith, 1959: Cr; [IWP]; PMBC
Ctenogobiops feroculus Lubbock & Polunin, 1977: Cr;
[IWP]
Ctenogobiops pomastictus Lubbock & Polunin, 1977: Cr;
[WP]; PMBC
Drombus globiceps (Hora, 1923): Mg; [EIO-WP]; PMBC
Drombus key (Smith, 1947): Cr, Mg, Sg; [IO]; PMBC
Drombus ocyurus (Jordan & Seale, 1907): Mg, Sg; [WP];
PMBC
Drombus triangularis (Weber, 1911): Mg, Sg; [IWP];
PMBC
Eugnathogobius illotus (Larson, 1999): Mg; [IM]; PMBC
Eugnathogobius microps Smith, 1931: Mg; [IM (presently
known from west coast of Thailand, Gulf of Thailand
and southern Vietnam)]
Eugnathogobius mindora (Herre, 1945): Mg; WP
Eugnathogobius polylepis (Wu & Ni, 1985): Mg; [IA]
Eugnathogobius variegatus (Peters, 1869): Mg; [IM];
PMBC
Eviota afelei Jordan & Seale, 1906: Cr; [WCP]
Eviota albolineata Jewett & Lachner, 1983: Cr; [IP]; PMBC
Eviota herrei Jordan & Seale, 1906: Cr; [WCP]
Eviota indica Lachner & Karnella, 1980: Cr; [IO]; PMBC
Eviota infulata (Smith, 1956): Cr; [IP]
Eviota mikiae Allen, 2001: Cr; [EIO]
Eviota nigriventris Giltay, 1933: Cr; [IA]; PMBC
Eviota pellucida Larson, 1976: Cr; [WCP]
Eviota prasina (Klunzinger, 1871): Cr; [IWP]; PMBC
Eviota prasites Jordan & Seale, 1906: Cr; [IP]
Eviota punctulata Jewett & Lachner, 1983: Cr; [WCP];
PMBC
Eviota queenslandica Whitley, 1932: Cr; [WP]; PMBC
Eviota rw-th sp.1: Cr
Eviota rw-th sp.2: Cr
Eviota rw-th sp.4: Cr
Eviota sebreei Jordan & Seale, 1906: Cr; [IP]; PMBC
Eviota sigillata Jewett & Lachner, 1983: Cr; [IWP]; PMBC
Eviota spilota Lachner & Karnella, 1980: Cr; [WP]; PMBC
Eviota zebrina Lachner & Karnella, 1978: Cr: [IWP];
PMBC
Eviota zonura Jordan & Seale, 1906: Cr; [WCP]
Exyrias bellissimus (Smith, 1959): Cr; [IWP]; PMBC
Exyrias puntang (Bleeker, 1851): Mg; [EIO-WP]
Favonigobius melanobranchus (Fowler, 1934): Cr, Mg,
Sg; [IWP]; PMBC
Favonigobius reichei (Bleeker, 1853): Cr, Mg, Sg; [IWP];
PMBC
Flabelligobius latruncularius (Klausewitz, 1974): Cr; [IO]
Flabelligobius russus (Cantor, 1849): OShr; [IM]; PMBC
Fusigobius dfh sp.4: Cr
Fusigobius duospilus Hoese & Reader, 1985: Cr; [IP];
PMBC
Fusigobius humeralis (Randall, 2001): Cr; [IP]
Fusigobius inflamaculatus Randall, 1994: Cr; [IWP]
Fusigobius maximus (Randall, 2001): Cr; [IWP]
Fusigobius melacron (Randall, 2001): Cr; [WP]
Fusigobius neophytus (Günther, 1877): Cr; [IP]; PMBC
Fusigobius pallidus (Randall, 2001): Cr; [IWP]
Fusigobius signipinnis Hoese & Obika, 1988: Cr; [WP]
Gladiogobius rex Shibukawa & Allen, 2007: Cr; [IO];
PMBC
Glossogobius bicirrhosus (Weber, 1894): Mg; [WP];
PMBC
Glossogobius circumspectus (Macleay, 1883): Mg; [WP];
PMBC
Glossogobius guiris (Hamilton, 1822): Mg; [IWP]
Glossogobius sparsipapillus Akihito & Meguro, 1976: Mg;
[WP]; PMBC
Gnatholepis anjerensis (Bleeker, 1850): Cr; [IP]; PMBC
Gnatholepis cauerensis (Bleeker, 1853): Cr; [IP]
Gobiodon citrinus (Rüppell, 1853): Cr; [IWP]
Gobiodon dfh sp.11: Cr
Gobiodon erythrospilus Bleeker, 1857: Cr; [WP]
Gobiodon histrio (Valenciennes, 1837): Cr; [IWP]; PMBC
Gobiodon prolixus Winterbottom & Harold, 2005: Cr;
[EIO-WP]
Gobiodon quinquestrigatus (Cuvier & Valenciennes, 1837):
Cr; [WCP]
Gobiodon rivulatus (Rüppell, 1828): Cr; [IWP]; PMBC
Gobiodon rw-th sp.1: Cr
Gobiodon rw-th sp.2: Cr
Gobiodon rw-th sp.3: Cr
Gobiopsis aporia Lachner & McKinney, 1978: Cr; [EIOWP]; PMBC
Gobiopsis macrostoma Steindachner, 1861: Mg; [NIOIM]; PMBC
Gobiopsis quinquecincta (Smith, 1931): Cr; [EIO-WP];
PMBC
Gobiopterus brachypterus (Bleeker, 1855): Mg; [IA];
PMBC
Gobiopterus panayensis (Herre, 1944): Mg; [WP]; PMBC
Gobiopterus rw-th sp.1: Mg
Hemigobius hoevenii (Bleeker, 1851): Mg; [IA]
Hemigobius mingi Herre, 1936: Mg; [IM]
Istigobius decoratus (Herre, 1927): Cr; [IWP]; PMBC
Istigobius diadema (Steindachner, 1877): Cr, Mg; [EIOIA]; PMBC
67
The fishes of southwestern Thailand
Istigobius goldmanni (Bleeker, 1852): Cr; [WP]; PMBC
Istigobius ornatus (Rüppell, 1830): Cr; [IP]; PMBC
Istigobius rigilius (Herre, 1953): Cr; [WCP]
Lophogobius bleekeri Popta, 1921: Mg; [IM]; PMBC
Mahidolia mystacina (Valenciennes, 1837): Cr, Sg, OShr;
[IWP]; PMBC
Mangarinus seshaiyai (Jacop & Rangarajan, 1960): Mg;
[EIO]
Mangarinus waterousi Herre, 1943: Mg; [WP]; PMBC
Mugilogobius chulae (Smith, 1932): Mg; [IM]; PMBC
Mugilogobius fasciatus Larson, 2001: Mg; [IM (presently
known from west coast of Thailand and Singapore)];
PMBC
Mugilogobius mertoni (Weber, 1911): Mg; [IWP]
Mugilogobius tigrinus Larson, 2001: Mg; [IM (presently
known from west coast of Thailand, Singapore and Gulf
of Thailand)]; PMBC
Myersina adonis Shibukawa & Satapoomin, 2006; Mg, Sg;
[IM (presently known from west coast of Thailand and
Singapore]; PMBC
Myersina crocatus (Wongratana, 1975): Cr, OShr; [IM];
PMBC
Myersina filifer (Cuvier & Valenciennes, 1837): OShr; [IWP];
PMBC
Myersina yangii (Chen, 1960): Cr, OShr; [WP]; PMBC
Oligolepis acutipennis (Valenciennes, 1837): Mg; [IWP]
Oplopomus caninoides (Bleeker, 1852): Cr, OShr; [EIOWP]; PMBC
Oplopomus oplopomus (Valenciennes, 1837): Cr, OShr;
[IP]; PMBC
Oxyurichthys microlepis (Bleeker, 1849): Mg, Sg; [IWP];
PMBC
Oxyurichthys ophthalmonema (Bleeker, 1856): Mg, Sg;
[EIO-WP]
Oxyurichthys papuensis (Valenciennes, 1837): Cr, OShr;
[IWP]; PMBC
Oxyurichthys tentacularis (Valenciennes, 1837): Mg, Sg;
[WP]; PMBC
Palutrus scapulopunctatus (De Beaufort, 1912): Cr; [IWP];
PMBC
Pandaka lidwilli (McCulloch, 1917): Mg; [WP]; PMBC
Pandaka pygmaea Herre, 1927: Mg; [IM]
Parachaeturichthys polynema (Bleeker, 1853): Mg; [IWP];
PMBC
Paragobiodon modestus (Regan, 1908): Cr; [IP]; PMBC
Paratrypauchen microcephalus (Bleeker, 1860): Mg, Sg;
[IWP]
Periophthalmus argentilineatus Valenciennes, 1837: Mg;
[IWP]; PMBC
Periophthalmus chrysospilos Bleeker, 1852: Mg; [EIOIM]; PMBC
Periophthalmus kalolo Lesson, 1830: Mg; [IWP]; PMBC
Periophthalmus minutus Eggert, 1935: Mg; [IA]; PMBC
Periophthalmus novemradiatus (Hamilton, 1822): Mg;
[EIO-IA]; PMBC
Periophthalmus septemradiatus (Hamilton, 1822): Mg;
[EIO-IM]
Periophthalmus walailakae Darumas & Tantochodok,
2002: Mg; [IM (presently known from west coast of
Thailand and Singapore)]; PMBC
Pleurosicya bilobata (Koumans, 1941): Sg; [IWP]
Pleurosicya coerulea Larson, 1990: Cr; [IP]
Pleurosicya fringilla Larson, 1990: Cr; [IWP]; PMBC
Pleurosicya labiata (Weber, 1913): Cr; [EIO-IA]; PMBC
Pleurosicya micheli Fourmanoir, 1971: Cr; [IP]
Pleurosicya mossambica Smith, 1959: Cr; [IP]; PMBC
Pleurosicya plicata Larson, 1990: Cr; [IWP]
Pleurosicya prognatha Goren, 1984: Cr; [IWP]
Priolepis cinctus (Regan, 1908): Cr; [IWP]; PMBC
Priolepis inhaca (Smith, 1949): Cr; [IWP]
Priolepis nuchifasciatus (Günther, 1873): Cr; [IA]
Priolepis semidoliatus (Valenciennes, 1837): Cr; [IP];
PMBC
Psammogobius biocellatus (Valenciennes, 1837): Mg, Sg;
[IWP]; PMBC
Pseudapocryptes lanceolatus (Bloch & Schneider, 1801):
Mg; [EIO-IM]
Pseudogobius javanicus (Bleeker, 1856): Mg; [IM]; PMBC
Redigobius bikolanus Herre, 1927: Mg; [WP]; PMBC
Redigobius chrysosoma (Bleeker, 1857): Mg; [IA]; PMBC
Scartelaos cantoris (Day, 1871): Mg, Sg; [AS]; PMBC
Scartelaos histophorus (Valenciennes, 1837): Mg, Sg; [NIOWP]; PMBC
Silhouettea sp.1: Cr, Sg; PMBC
Stigmatogobius pleurostigma (Bleeker, 1849): Mg; [IM]
Stigmatogobius sandanundio (Hamilton, 1822): Mg; [EIO];
PMBC
Stonogobiops nematodes Hoese & Randall, 1982: Cr; [IA]
Sueviota lachneri Winterbottom & Hoese, 1988: Cr; [IWP];
PMBC
Taenioides cirratus (Blyth, 1860): Mg, Sg, OShr; [IWP];
PMBC
Taenioides gracilis (Valenciennes, 1837): OShr; [IWP];
PMBC
Trimma emeryi Winterbottom, 1985: Cr; [IO]
Trimma flammeum (Smith, 1959): Cr; [IO]; PMBC
Trimma fucatum Winterbottom & Southcott, 2007: Cr;
[AS]; PMBC
Trimma naudei Smith, 1956: Cr; [IWP]; PMBC
Trimma okinawae (Aoyaki, 1949): Cr; [WCP]; PMBC
Trimma sanguinellus Winterbottom & Southcott, 2007:
Cr; [AS]; PMBC
Trimma striata (Herre, 1945): Cr; [EIO-WP]; PMBC
Trimma talori Lobel, 1979: Cr; [IP]
Trimma winterbottomi Randall & Downing, 1994: Cr; [IO];
PMBC
Trypauchen vagina (Bloch & Schneider, 1801): OShr; [NIOIM]; PMBC
Valenciennea immaculata (Ni, 1981): Sg; [WP]
Valenciennea limicola Hoese & Larson, 1994: Cr; [WP];
PMBC
Valenciennea mularis (Valenciennes, 1837): Cr, Sg, OShr;
[IA]
Valenciennea parva Hoese & Larson, 1994: Cr; [IWP]
68
Phuket mar. biol. Cent. Res. Bull.
Valenciennea puellaris (Tomiyama, 1956): Cr, OShr; [IP];
PMBC
Valenciennea randalli Hoese & Larson, 1994: Cr, OShr;
[WP]; PMBC
Valenciennea sexguttata (Valenciennes, 1837): Cr, OShr;
[IP]; PMBC
Valenciennea strigata (Broussonet, 1782): Cr; [IP]; PMBC
Valenciennea wardii (Playfair, 1867): Cr, OShr; [IWP];
PMBC
Vanderhorstia ambanoro (Fourmanoir, 1957): Cr, OShr;
[IWP]; PMBC
Vanderhorstia ornatissima Smith, 1959: Cr; [IP]
Yongeichthys nebulosus (Forsskål, 1775): Cr, Mg, Sg, OShr;
[IWP]; PMBC
Microdesmidae
Gunnellichthys curiosus Dawson, 1968: Cr; [IP]
Gunnellichthys viridescens Dawson, 1968: Cr; [IP]
Ptereleotridae
Nemateleotris decora Randall & Allen, 1973: Cr; [IWP]
Nemateleotris magnifica Fowler, 1928: Cr; [IP]
Oxymetopon compressus Chan, 1966: Cr, OShr; [WP];
PMBC
Parioglossus formosus (Smith, 1931): Cr; [WP]; PMBC
Parioglossus palustris (Herre, 1945): Mg; [WP]
Parioglossus philippinus (Herre, 1940): Cr; [IWP]; PMBC
Ptereleotris arabica Randall & Hoese, 1985: Cr, OShr;
[IO]; PMBC
Ptereleotris evides (Jordan & Hubbs, 1925): Cr; [IP]
Ptereleotris heteroptera (Bleeker, 1855): Cr; [IP]; PMBC
Ptereleotris microlepis (Bleeker, 1856): Cr; [IP]; PMBC
Ptereleotris monoptera (Bleeker, 1855): Cr; [IP]
Ptereleotris zebra (Fowler, 1938): Cr; [IP]
Kurtidae
Kurtus indicus Bloch, 1786: OShr; [EIO-IM]; PMBC
Ephippidae
Ephippus orbis (Bloch, 1787): Mg, Sg, OShr; [IWP];
PMBC
Platax boersi Bleeker, 1852: Cr; [IP]
Platax orbicularis (Forsskål, 1775): Cr, Mg, Sg, OShr; [IP];
PMBC
Platax pinnatus (Linnaeus, 1758): Cr, OShr; [WP]; PMBC
Platax teira (Forsskål, 1775): Cr, OShr; [IWP]; PMBC
Scatophagidae
Scatophagus argus (Linnaeus, 1758): Mg, Sg; [NIO-WP];
PMBC
Siganidae
Siganus argenteus (Quoy & Gaimard, 1825): Cr; [IP];
PMBC
Siganus canaliculatus (Park, 1797): Cr, Mg, Sg, OShr; [NIOIA]; PMBC
Siganus corallinus (Valenciennes, 1835): Cr; [IWP]; PMBC
Siganus fuscescens (Houttuyn, 1782): Cr; [WP]; PMBC
Siganus guttatus (Bloch, 1787): Cr, Mg, Sg, OShr; [IM]
Siganus javus (Linnaeus, 1766): Cr, Mg, Sg, OShr; [NIOWP]
Siganus lineatus (Valenciennes, 1835): Cr; [EIO-WP]
Siganus magnificus (Burgess, 1977): Cr; [EEIO]; PMBC
Siganus puelloides Woodland & Randall, 1979: Cr; [EIO];
PMBC
Siganus stellatus laqueus Von Bonde, 1934: Cr, OShr; [IO];
PMBC
Siganus vermiculatus (Valenciennes, 1835): Cr, Mg, Sg,
OShr; [EIO-WP]; PMBC
Siganus virgatus (Valenciennes, 1835): Cr, Sg, OShr; [EIOWP]; PMBC
Zanclidae
Zanclus cornutus (Linnaeus, 1758): Cr; [IP]; PMBC
Acanthuridae
Acanthurus bariene Lesson, 1830: Cr; [WP-EIO]; PMBC
Acanthurus blochii Valenciennes, 1835: Cr; [IP]
Acanthurus guttatus Forster, 1801: Cr; [IP]
Acanthurus leucocheilus Herre, 1927: Cr; [IWP]
Acanthurus leucosternon Bennett, 1832: Cr; [IO]; PMBC
Acanthurus lineatus (Linnaeus, 1758): Cr; [IP]; PMBC
Acanthurus maculiceps (Ahl, 1923): Cr; [EIO-WCP]
Acanthurus mata (Cuvier, 1829): Cr; [IP]; PMBC
Acanthurus nigricauda Duncker & Mohr, 1929: Cr; [IP];
PMBC
Acanthurus nigrofuscus (Forsskål, 1775): Cr; [IP]
Acanthurus olivaceus Forster, 1801: Cr; [WCP]
Acanthurus tennenti Günther, 1861: Cr; [IO]
Acanthurus thompsoni (Fowler, 1923): Cr; [IP]; PMBC
Acanthurus triostegus (Linnaeus, 1758): Cr; [IP]; PMBC
Acanthurus tristis Tickell In Day, 1888: Cr; [IO]; PMBC
Acanthurus xanthopterus Valenciennes, 1835: Cr; [IP];
PMBC
Ctenochaetus binotatus Randall, 1955: Cr; [IP]; PMBC
Ctenochaetus striatus (Quoy & Gaimard, 1825): Cr; [IP];
PMBC
Ctenochaetus truncatus Randall & Clements, 2001: Cr;
[IO]; PMBC
Naso brachycentron (Valenciennes, 1835): Cr; [IP]
Naso brevirostris (Valenciennes, 1835): Cr; [IP]; PMBC
Naso caeruleocauda Randall, 1994: Cr; [IA]
Naso elegans (Rüppell, 1829): Cr; [IO]; PMBC
Naso hexacanthus (Bleeker, 1855): Cr; [IP]; PMBC
Naso lopezi Herre, 1927: Cr; [WP]
Naso mcdadei Johnson, 2002: Cr; [IWP]
Naso thynnoides (Valenciennes, 1835): Cr; [IP]
Naso unicornis (Forsskål, 1775): Cr; [IP]; PMBC
Naso vlamingii (Valenciennes, 1835): Cr; [IP]; PMBC
Paracanthurus hepatus (Linnaeus, 1766): Cr; [IP]
Zebrasoma desjardinii (Bennett, 1835): Cr; [IO]; PMBC
Zebrasoma scopas (Cuvier, 1829): Cr; [IP]; PMBC
Sphyraenidae
Sphyraena baracuda (Walbaum, 1792): Cr, Sg, OShr;
[COS]; PMBC
Sphyraena forsteri Cuvier, 1829: Cr, OShr; [IP]; PMBC
Sphyraena jello Cuvier, 1829: Cr, Mg, Sg, OShr; [IP];
PMBC
Sphyraena obtusata Cuvier, 1829: Cr, Sg, OShr; [IWP];
PMBC
69
The fishes of southwestern Thailand
Sphyraena pinguis Günther, 1874: Cr; [IWP]
Sphyraena putnamae Jordan & Seale, 1905: Mg, Sg, OShr;
[IWP]
Sphyraena qenie Klunzinger, 1870: Cr; [IP]
Gempylidae
Gempylus serpens Cuvier, 1829: OShr; [COS]
Neoepinnula orientalis (Gilchrist & von Bonde, 1924): OShr;
[IWP]; PMBC
Nesiarchus nasutus Johnson, 1862: OShr; [COS]; PMBC
Promethichthys prometheus (Cuvier, 1832): OShr; [COS]
Rexea bengalensis (Alcock, 1884): OShr; [IWP]
Thyrsitoides marleyi Fowler, 1929: OShr; [IWP]
Trichiuridae
Eupleurogrammus muticus (Gray, 1831): OShr; [NIO-IM]
Tentoriceps cristatus (Klunzinger, 1884): OShr; [IWP];
PMBC
Trichiurus gangeticus Gupta, 1966: OShr; [EIO]; PMBC
Trichiurus lepturus Linnaeus, 1758: Mg, Sg, OShr; [COS];
PMBC
Scombridae
Acanthocybium solandi (Cuvier, 1831): Cr, OShr; [COS]
Auxis rochei rochei (Risso, 1810): OShr; [COS]; PMBC
Auxis thazard thazard (Lacepède, 1800): OShr; [COS];
PMBC
Euthynnus affinis (Cantor, 1849): Cr, OShr; [IP]; PMBC
Grammatorcynos bilineatus (Rüppell, 1836): Cr, OShr;
[IP]; PMBC
Gymnosarda unicolor (Rüppell, 1838): Cr, OShr; [IP];
PMBC
Katsuwonus pelamis (Linnaeus, 1758): OShr; [COS];
PMBC
Rastrelliger brachysoma (Bleeker, 1851): OShr; [WP];
PMBC
Rastrelliger faughni Matsui, 1967: OShr; [EIO-WP];
PMBC
Rastrelliger kanagurta (Cuvier, 1817): Cr, Mg, Sg, OShr;
[IP]; PMBC
Scomberomorus commerson (Lacepède, 1800): Cr, Mg,
OShr; [IP]; PMBC
Scomberomorus guttatus (Bloch & Schneider, 1801): OShr;
[NIO-WP]; PMBC
Scomberomorus lineolatus (Cuvier, 1831): OShr; [NIOIM]
Thunnus alalunga (Bonnaterre, 1788): OShr; [COS]
Thunnus albacares (Bonnaterre, 1788): OShr; [COS];
PMBC
Thunnus obesus (Lowe, 1839): OShr; [COS]
Thunnus tonggol (Bleeker, 1851): OShr; [IWP]; PMBC
Xiphiidae
Xiphias gladius Linnaeus, 1758: OShr; [COS]
Istiophoridae
Istiophorus platypterus (Shaw & Nodder, 1792): OShr;
[IP]; PMBC
Makaira indica (Cuvier, 1832): OShr; [IP]
Tetrapturus audax (Philippi, 1887): OShr; [IP]
Centrolophidae
Psenopsis obscura Headrich, 1967: OShr; [IM]
Nomeidae
Cubiceps pauciradiatus Günther, 1872: OShr; [CIR];
PMBC
Cubiceps whiteleggii (Waite, 1894): OShr; [IWP]
Ariommatidae
Ariomma indicum (Day, 1870): OShr; [IWP]; PMBC
Stromateidae
Pampus argenteus (Euphrasen, 1788): OShr; [NIO-WP];
PMBC
Pampus chinensis (Euphrasen, 1788): OShr; [NIO-WP]
Caproidae
Antigonia rubescens (Günther, 1860): OShr; [WP]
Order Pleuronectiformes
Psettodidae
Psettodes erumei (Bloch & Schneider, 1801): Mg, OShr;
[IWP]; PMBC
Citharidae
Brachypleura novaezeelandiae Günther, 1862: OShr;
[NIO-IA]; PMBC
Paralichthyidae
Pseudorhombus argus Weber, 1913: OShr; [IA]
Pseudorhombus arsius (Hamilton, 1822): Mg, Sg, OShr;
[IWP]; PMBC
Pseudorhombus dupliciocellatus Regan, 1905: OShr; [IA];
PMBC
Pseudorhombus elevatus Ogilby, 1912: Mg, OShr; [IWP];
PMBC
Pseudorhombus malayanus Bleeker, 1866: Mg, OShr;
[NIO-IM]
Pseudorhombus quinquocellatus Weber & de Beaufort,
1929: OShr; [EIO-IA]; PMBC
Pseudorhombus triocellatus (Schneider, 1801): OShr; [EIOIA]; PMBC
Bothidae
Arnoglossus aspilos (Bleeker, 1851): Cr; [NIO-IA]; PMBC
Arnoglossus macrolophus Alcock, 1889: OShr; [IWP]
Arnoglossus tapeinosoma (Bleeker, 1865): OShr; [NIOWP]
Arnoglossus tenuis Günther, 1880: OShr; [WP]; PMBC
Asterorhombus intermedius (Bleeker, 1866): OShr; [IWP]
Bothus pantherinus (Rüppell, 1830): OShr; [IP]
Chascanopsetta lugubris Alcock, 1894: OShr; [IWP]
Engyprosopon grandisquama (Temminck & Schlegel,
1846): OShr; [IWP]; PMBC
Engyprosopon latifrons (Regan, 1908): OShr; [IWP];
PMBC
Engyprosopon maldivensis (Regan, 1908): OShr; [EIO-WP]
Grammatobothus polyophthalmus (Bleeker, 1866): OShr;
[EIO-WP]; PMBC
Laeops guentheri Alcock, 1890: OShr; [NIO-IM]
Laeops kitaharae (Smith & Pope, 1906): OShr; [IWP]
Poecilopsettidae
Poecilopsetta colorata Günther, 1880: OShr; [EIO-IM]
Samaridae
Samaris cristatus Gray, 1831: OShr; [IWP]; PMBC
70
Phuket mar. biol. Cent. Res. Bull.
Soleidae
Aesopia cornuta Kaup, 1858: OShr; [IWP]; PMBC
Aseraggodes dubius Weber, 1913: OShr; [WP]
Aseraggodes satapoomini Randall & Desoutter-Meniger,
2007: Cr; [AS]
Brachirus pan (Hamilton, 1822): OShr; [EIO-IM]
Heteromycteris oculus (Alcock, 1889): OShr; [IWP];
PMBC
Liachirus melanospilos (Bleeker, 1854): OShr; [EIO-WP]
Pardachirus pavoninus (Lacepède, 1802): Cr, Sg, OShr;
[EIO-WP]; PMBC
Solea ovata Richardson, 1846: Sg, OShr; [EIO-IM]; PMBC
Synaptura commersonii (Lacepède, 1802): Sg, OShr; [NIOIM]; PMBC
Zebrias altipinnis (Alcock, 1890): OShr; [EIO]
Zebrias quagga Kaup, 1858: Mg, Sg, OShr; [IWP]; PMBC
Zebrias synapturoides (Jenkins, 1910): OShr; [NIO-WP]
Cynoglossidae
Cynoglossus arel (Schneider, 1801): Mg, OShr; [NIO-IM]
Cynoglossus bilineatus (Lacepède, 1802): Mg, OShr; [NIOWP]; PMBC
Cynoglossus cynoglossus (Hamilton, 1822): OShr; [NIOIM]; PMBC
Cynoglossus kopsi (Bleeker, 1851): OShr; [NIO-WP]
Cynoglossus lida (Bleeker, 1851): Mg, Sg, OShr; [IWP];
PMBC
Cynoglossus lingua Hamilton, 1822: Mg, OShr; [IWP]
Cynoglossus macrolepidotus (Bleeker, 1851): OShr; [WP];
PMBC
Cynoglossus monopus (Bleeker, 1849): OShr; [EIO-IM]
Cynoglossus oligolepis (Bleeker, 1854): OShr; [WP];
PMBC
Cynoglossus puncticeps (Richardson, 1846): Sg, OShr;
[NIO-IA]; PMBC
Cynoglossus semifasciatus Day 1877: Mg, Sg, OShr; [EIO];
PMBC
Cynoglossus sibogae Weber, 1913: OShr; [IM]; PMBC
Paraplagusia bilineata (Bloch, 1787): OShr; [IWP]
Paraplagusia blochii (Bleeker, 1851): OShr; [NIO-WP]
Symphurus septemstriatus (Alcock, 1891): OShr; [EIOIM]
Order Tetraodontiformes
Triacanthodidae
Halimochirurgus alcocki Weber, 1913: OShr; [IWP]
Tydemania navigatoris Weber, 1913: OShr; [IWP]; PMBC
Triacanthidae
Pseudotriacanthus strigilifer (Cantor, 1849): OShr; [NIOIA]; PMBC
Triacanthus biaculeatus (Bloch, 1786): Mg, Sg, OShr;
[NIO-WP]; PMBC
Triacanthus nieuhofi Bleeker, 1852: Sg, OShr; [EIO-IA]
Trixiphichthys weberi (Chaudhuri, 1910): OShr; [EIO-IA];
PMBC
Balistidae
Abalistes stellatus (Anonymous, 1798): Cr, OShr; [IWP];
PMBC
Balistapus undulatus (Park, 1797): Cr, OShr; [IP]; PMBC
Balistoides conspicillum (Bloch & Schneider, 1801): Cr,
OShr; [IP]; PMBC
Balistoides viridescens (Bloch & Schneider, 1801): Cr,
OShr; [IP]; PMBC
Canthidermis maculatus (Bloch, 1786): OShr; [COS];
PMBC
Canthidermis rotundatus (Marion de Procé, 1822): OShr;
[IWP]; PMBC
Melichthys indicus Randall & Klausewitz, 1973: Cr, OShr;
[IO]; PMBC
Melichthys vidua (Solander, 1844): Cr; [IP]
Odonus niger (Rüppell, 1837): Cr, OShr; [IP]; PMBC
Pseudobalistes flavimarginatus (Rüppell, 1829): Cr, OShr;
[IP]
Pseudobalistes fuscus (Bloch & Schneider, 1801): Cr, OShr;
[IP]; PMBC
Rhinecanthus aculeatus (Linnaeus, 1758): Cr, OShr; [IP]
Rhinecanthus rectangulus (Bloch & Schneider, 1801): Cr,
OShr; [IP]
Rhinecanthus verrucosus (Linnaeus, 1758): Cr, OShr; [IWP]
Sufflamen bursa (Bloch & Schneider, 1801): Cr; [IP]
Sufflamen chrysopterum (Bloch & Schneider, 1801): Cr,
OShr; [IWP]; PMBC
Sufflamen fraenatum (Latreille, 1804): Cr, OShr; [IP];
PMBC
Xanthichthys auromarginatus (Bennett, 1831): Cr, OShr;
[IP]
Monacanthidae
Acreichthys tomentosus (Linnaeus, 1758): Sg; [IA]
Aluterus monoceros (Linnaeus, 1758): OShr; [CIR]; PMBC
Aluterus scriptus (Osbeck, 1765): Cr, OShr; [CIR]; PMBC
Amanses scopas (Cuvier, 1829): Cr; [IP]; PMBC
Anacanthus barbatus Gray, 1831: Sg, OShr; [NIO-IA]
Cantherhines dumerilli (Hollard, 1854): Cr, OShr; [IP]
Cantherhines multilineatus (Tanaka, 1918): OShr; [IWP;
PMBC
Cantherhines pardalis (Rüppell, 1837): Cr, OShr; [IP];
PMBC
Monacanthus chinensis Osbeck, 1765: Cr, Mg, Sg, OShr;
[WP]; PMBC
Oxymonacanthus longirostris (Bloch & Schneider, 1801):
Cr; [IP]; PMBC
Paraluteres sp.1: Cr; [AS]
Paramonacanthus japonicus (Tilesius, 1810): OShr; [EIOWP]
Paramonacanthus tricuspis (Hollard, 1854): Mg, Sg, OShr;
[EIO]; PMBC
Pseudomonacanthus macrurus (Bleeker, 1857): Mg, Sg,
OShr; [IWP]; PMBC
Pseudomonacanthus peroni (Hollard, 1854): OShr; [IA]
Ostraciidae
Lactoria cornuta (Linnaeus, 1758): Cr, Mg, Sg, OShr; [IP];
PMBC
Lactoria diaphana (Bloch & Schneider, 1801): Mg, OShr;
[IP]
71
The fishes of southwestern Thailand
Ostracion cubicus Linnaeus, 1758: Cr, Mg, Sg, OShr; [IP];
PMBC
Ostracion meleagris Shaw, 1796: Cr; [IP]; PMBC
Ostracion nasus Bloch, 1785: OShr; [EIO-WP]; PMBC
Ostracion rhinorhynchus Bleeker, 1852: Cr, OShr; [IWP]
Ostracion solorensis Bleeker, 1853: Cr; [IWP]
Tetrosomus gibbosus (Linnaeus, 1758): Cr, OShr; [IWP];
PMBC
Tetrosomus reipublicae (Whitley, 1930): OShr; [IWP]
Tetraodontidae
Amblyrhynchotes honckenii (Bloch, 1795): OShr; [IWP]
Arothron caeruleopunctatus Matsuura, 1994: Cr, Mg, Sg;
[EIO-WP]
Arothron hispidus (Linnaeus, 1758): Cr, Mg, Sg, OShr;
[IP]; PMBC
Arothron immaculatus (Bloch & Schneider, 1801): Cr, OShr;
[IWP]; PMBC
Arothron manilensis (Marion de Porcé, 1822): Cr, Sg, OShr;
[WCP]
Arothron mappa (Lesson, 1830): Cr, OShr; [IP]; PMBC
Arothron nigropunctatus (Bloch & Schneider, 1801): Cr,
OShr; [IP]; PMBC
Arothron reticularis (Bloch & Schneider, 1801): Mg, OShr;
[IWP]; PMBC
Arothron stellatus (Bloch & Schneider, 1801): Cr, OShr;
[IP]; PMBC
Canthigaster smithae Allen & Randall, 1977: Cr; [IO]
Canthigaster solandri (Richardson, 1844): Cr, OShr; [IP];
PMBC
Chelonodon patoca (Hamilton, 1822): Mg, Sg, OShr; [IWP];
PMBC
Lagocephalus gloveri Abe & Tabeta, 1983: OShr; [IWP]
Lagocephalus inermis (Temminck & Schlegel, 1844): OShr;
[IWP]; PMBC
Lagocephalus lunaris (Bloch & Schneider, 1801): Mg, Sg,
OShr; [IWP]; PMBC
Lagocephalus scleratus (Gmelin, 1789): OShr; [IWP];
PMBC
Lagocephalus spadiceus (Richardson, 1844): OShr; [IWP]
Takifugu oblongus (Bloch, 1786): Mg, Sg, OShr; [IWP];
PMBC
Tetraodon fluviatilis Hamilton, 1822: OShr; [EIO-IM];
PMBC
Tetraodon nigroviridis Procé, 1822: Mg, Sg, OShr; [EIOWP]; PMBC
Torquigener hypselogeneion (Bleeker, 1852): Cr; [IWP];
PMBC
Diodontidae
Cyclichthys orbicularis (Bloch, 1785): Mg, Sg, OShr;
[IWP]; PMBC
Diodon eydouxii Brisout de Barneville, 1846: OShr; [IWP]
Diodon holocanthus Linnaeus, 1758: OShr; [CIR]; PMBC
Diodon hystrix Linnaeus, 1758: Cr, OShr; [CIR]; PMBC
Diodon liturosus Shaw, 1804: Cr, Sg, OShr; [IP]; PMBC
Molidae
Masturus lanceolatus (Lienard, 1840): OShr; [COS]
Mola mola (Linnaeus, 1758): OShr; [COS]; PMBC
72
Phuket mar. biol. Cent. Res. Bull.
Appendix B
Selected taxonomic literature pertaining to the records of fishes from southwestern Thailand,
the Andaman Sea
Class Myxini
Wongratana, T. 1983. Eptatretus indrambaryai, a new species of hagfish (Myxinidae) from the Andaman Sea. Nat Hist.
Bull. Siam Soc. 31(2): 139–150.
Class Chondrichthyes
Compagno, L.J.V. and P.R. Last. 2008. A new species of wedgefish, Rhynchobatus palpebratus sp. nov. (Rhynchobatoidei:
Rhynchobatidae), from the Indo-West Pacific. In: P.R. Last, W.T. White and J.J. Pogonoski (eds.). Descriptions
of New Australian Chondrichthyans. CISRO Marine and Atmospheric Research Paper No. 022. pp. 227–240.
Krajangdara, T., S. Puteeka, R. Phoonsawat, S. Vibunpant, R. Sojittosakul and R. Chayakul. 2006. Sharks found in Thai
waters and adjacent areas. Marine Fisheries Research and Development Bureau, Department of Fisheries.
Technical Paper No. 17/2006. 99 p. (in Thai)
Monkolprasit, S. 1984. The cartilaginous fishes (Class Elasmobranchii) found in Thai waters and adjacent areas.
Department of Fishery Biology, Faculty of Fisheries, Kasetsart University, Bangkok, Thailand. 175 p.
Class Actinopterygii
Order Albuliformes
Hidaka, K., Y. Iwatsuki and J.E. Randall. 2008. A review of the Indo-Pacific bonefishes of the Albula argentea complex,
with a description of a new species. Ichthyol. Res. 55(1): 53–64.
Order Anguilliformes
McCosker, J.E. 1972. Two new genera and two new species of western Pacific snake-eels (Apodes: Ophichthidae).
Proc. Cal. Acad. Sci. 39(10): 111–119.
McCosker, J.E. and J.E. Randall. 2001. Revision of the snake-eel genus Brachysomophis (Anguilliformes: Ophichthidae),
with descriptions of two new species and comments on the species of Mystriophis. Indo-Pacific Fishes 33: 1–
32, 2 pls.
Böhlke, E.B. 2000. Notes on the identity of small, brown, unpatterned Indo-Pacific moray eels, with descriptions of
three new species (Anguilliformes: Muraenidae). Pac. Sci. 54(4): 395–416.
Böhlke, E.B. and J.E. Randall. 2000. A review of the moray eels (Anguilliformes; Muraenidae) of the Hawaiian Islands,
with descriptions of two new species. Proc. Acad. nat. Sci. Philad. 150: 203–278, 9 pls.
Order Clupeiformes
Whitehead, P.J.P. 1985. FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (suborder Clupeoidei). An annotated
and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies, and wolf-herrings. Part
1 – Chirocentridae, Clupeidae and Pristigasteridae. FAO Species Synopsis No. 125, Vol. 7, Pt. 1: 1–303.
Whitehead, P.J.P., G.J. Nelson and T. Wongratana. 1988. FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world
(suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats,
shads, anchovies, and wolf-herrings. Part 2 – Engraulididae. FAO Species Synopsis No. 125, Vol. 7, Pt. 2: 305–
579.
Order Ophidiiformes
Markle, D.F. and J.E. Olney. 1990. Systematics of the pearlfishes (Pisces: Carapidae). Bull. Mar. Sci. 47(2): 269–410.
Møller, P.R. and W. Schwarzhans. 2008. Review of the Dinematichthyini (Teleostei: Bythitidae) of the Indo-west
Pacific. Part IV. Dinematichthys and two new genera with descriptions of nine new species. The Beagle,
Records of the Museum of Art Galleries of the Northern Territory 24: 87–146.
Nielsen, J.G. and Y. Machida. 1988. Revision of the Indo-Pacific bathyal fish genus Glyptophidium (Ophidiiformes,
Ophidiidae). Jap. J. Ichthyol. 35(3): 289–319.
Order Lophiiformes
Bradbury, M.G. 1988. Rare fishes of the deep-sea genus Halieutopsis: a review with descriptions of four new species
(Lophiiformes: Ogcocephalidae). Fieldiana, Zool. 44: 1–22.
Order Atheriniformes
Kimura, S., D. Golani, Y. Iwatsuki, M. Tabuchi and T. Yoshino. 2007. Redescriptions of the Indo-Pacific atherinid fishes
Atherinomorus forskalii, Atherinomorus lacunosus, and Atherinomorus pinguis. Ichthyol. Res. 54(2): 145–
159.
Parenti, L.R. 1989. A phylogenetic revision of the phallostethid fishes (Atherinomorpha, Phallostethidae). Proc. Cal.
Acad. Sci. 46(11): 243–277.
Order Beloniformes
73
The fishes of southwestern Thailand
Parenti, L.R. 2008. A phylogenetic analysis and taxonomic revision of ricefishes, Oryzias and relatives (Beloniformes,
Andrianichthyidae). Zool. J. Linn. Soc. 154: 494–610.
Order Beryciformes
Randall, J.E. 1998. Revision of the Indo-Pacific squirrelfishes (Beryciformes: Holocentridae: Holocentrinae) of the
genus Sargocentron, with descriptions of four new species. Indo-Pacific Fishes 27: 1–105, 11 pls.
Order Gasterosteiformes
Dawson, C.E. 1985. Indo-Pacific Pipefishes (Red Sea to the Americas). The Gulf Coast Research Laboratory, Ocean
Springs, Mississippi. 230 p.
Order Scorpaeniformes
Imamura, H. 1996. Phylogeny of the family Platycephalidae and related taxa (Pisces: Scorpaeniformes). Species Diversity
1(2): 123–233.
Knapp, L.W. and T. Wongratana. 1987. Sorsogona melanoptera, a new flathead fish from the northern Indian Ocean
(Teleostei: Platycephalidae). Proc. Biol. Soc. Wash. 100(2): 381–385.
Poss, S.G. and W.N. Eschmeyer. 1980. Xenaploactis, a new genus for Prosopodasys asperrimus Günther (Pisces:
Aploactinidae), with descriptions of two new species. Proc. Cal. Acad. Sci. 42(8): 287–293.
Randall, J.E. and W.N. Eschmeyer. 2001. Revision of the Indo-Pacific scorpionfish genus Scorpaenopsis, with descriptions
of eight new species. Indo-Pacific Fishes 34: 1–79, 12 pls.
Wongratana, T. 1975. A record of a flathead fish, Thysanophrys papillolabium Schultz, in the Andaman Sea, with a key
to fourteen species of the Platycephalidae from Thai waters (Pisces: Platycephalidae). Phuket mar. biol. Cent.
Res. Bull. 7: 1–9.
Order Perciformes
Family Acropomatidae
Okamoto, M. and H. Ida. 2002. Acropoma argentistigma, a new species from the Andaman Sea, off southern Thailand
(Perciformes: Acropomatidae). Ichthyol. Res. 49(3): 281–285.
Family Serranidae
Randall, J.E. and C.C. Baldwin. 1997. Revision of the serranid fishes of the subtribe Pseudogrammina, with descriptions
of five new species. Indo-Pacific Fishes 26: 1–56, 1 pl.
Randall, J.E. and P.C. Heemstra. 1991. Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with
descriptions of five new species. Indo-Pacific Fishes 20: 1–332, 41 pls.
Randall, J.E. and P.C. Heemstra. 2007. Meganthias filiferus, a new species of anthiine fish (Perciformes: Serranidae),
from the Andaman Sea off southwestern Thailand. Phuket mar. biol. Cent. Res. Bull. 68: 5–9.
Randall, J.E. and R. Lubbock. 1981. A revision of the serranid fishes of the subgenus Mirolabrichthys (Anthiinae:
Anthias), with descriptions of five new species. Contr. Sci. 333: 1–27.
Randall, J.E. and U. Satapoomin. 2000. Cephalopholis polyspila, a new species of grouper (Perciformes: Serranidae:
Epinephelinae) from southwestern Thailand and Sumatra. Phuket mar. biol. Cent. Res. Bull. 63: 1–7.
Family Pseudochromidae
Gill, A.C. 2004. Revision of the Indo-Pacific dottyback fish subfamily Pseudochrominae (Perciformes: Pseudochromidae).
Smithiana Monograph 1. 213 p, 12 pls.
Winterbottom, R. and J.E. Randall. 1994. Two new species of congrogadins (Teleostei: Pseudochromidae), with range
extensions for four other species. Can. J. Zool. 72: 750–756.
Family Plesiopidae
Mooi, R.D. 1995. Revision, phylogeny, and discussion of biology and biogeography of the fish genus Plesiops
(Perciformes: Plesiopidae). Royal Ontario Museum Life Sciences Contributions No. 159. iv + 107 p.
Family Opistognathidae
Smith-Vaniz, W.F. 2009. Three new species of Indo-Pacific jawfishes (Opistognathus: Opistognathidae), with the
posterior end of the upper jaw produced as a thin flexible lamina. Aqua, Int. J. Ichthyol. 15(2): 69–108.
Family Apogonidae
Gon, O. 1996. Revision of the cardinalfish subgenus Jaydia (Perciformes, Apogonidae, Apogon). Trans. Roy. Soc. S. Afr.
51: 147–194.
Gon, O. 2000. The taxonomic status of the cardinalfish species Apogon niger, A. nigripinnis, A. pharaonis, A. sialis, and
related species (Perciformes: Apogonidae). J.L.B. Smith Inst. Ichthyol. Spec. Publ. 65: 1–20.
Gon, O. and J.E. Randall. 2003. Revision of the Indo-Pacific cardinalfish genus Archamia (Perciformes: Apogonidae),
with description of a new species. Indo-Pacific Fishes 35: 1–49, 3 pls.
Fraser, T.H. 2005. A review of the species in the Apogon fasciatus group with a description of a new species of
cardinalfish from the Indo-west Pacific (Perciformes: Apogonidae). Zootaxa 924: 1–30.
Fraser, T.H. 2008. Cardinafishes of the genus Nectamia (Apogonidae, Perciformes) from the Indo-Pacific region with
descriptions of four new species. Zootaxa 1691: 1–52.
74
Phuket mar. biol. Cent. Res. Bull.
Fraser, T.H. and E.A. Lachner. 1985. A revision of the cardinalfish subgenera Pristiapogon and Zoramia (Genus Apogon)
of the Indo-Pacific region (Teleostei: Apogonidae). Smithson. Contr. Zool. 42. 47 p.
Randall, J.E., T.H. Fraser and E.A. Lachner. 1990. On the validity of the Indo-Pacific cardinalfishes Apogon aureus
(Lacepède) and A. fleurieu (Lacepède), with description of a related new species from the Red Sea. Proc. Biol.
Soc. Wash. 103(1): 39–62.
Family Sillaginidae
McKay, R.J. 1985. A revision of the fishes of the family Sillaginidae. Mem. Qd. Mus. 22(1): 1–73.
Family Leiognathidae
Kimura, S., T. Ito, T. Peristiwady, Y. Iwatsuki and T. Yoshino. 2005. The Leiognathus splendens complex (Perciformes:
Leiognathidae) with the description of a new species, Leiognathus kupanensis Kimura and Peristiwady.
Ichthyol. Res. 52(3): 275–291.
Kimura, S., R. Kimura and K. Ikejima. 2008. Revision of the genus Nuchequula with descriptions of three new species
(Perciformes: Leiognathidae). Ichthyol. Res. 55(1): 22–42.
Kimura, S., T. Yamashita and Y. Iwatsuki. 2000. A new species, Gazza rhombea, from the Indo-west Pacific, with a
redescription of G. achlamys Jordan & Starks, 1917 (Perciformes: Leiognathidae). Ichthyol. Res. 47(1): 1–12.
Mochizuki, K. and M. Hayashi. 1989. Revision of the leiognathid fishes of the genus Secutor, with two new species. Sci.
Rept. Yokosuka City Mus. 37: 83–95.
Yamashita, T., S. Kimura and Y. Iwatsuki. 1998. Validity of the leiognathid fish, Gazza dentex (Valenciennes in Cuvier &
Valenciennes, 1835), with designation of a lectotype, and redescription of G. minuta (Bloch, 1795). Ichthyol.
Res. 45(3): 271–280.
Family Lutjanidae
Allen, G.R. and F.H. Talbot. 1985. Review of the snappers of the genus Lutjanus (Pisces: Lutjanidae) from the IndoPacific, with description of a new species. Indo-Pacific Fishes 11: 1–87, 10 pls.
Family Caesionidae
Carpenter, K.E. 1987. Revision of the Indo-Pacific fish family Caesionidae (Lutjanoidea), with descriptions of five new
species. Indo-Pacific Fishes 15: 1–56, 7 pls.
Family Gerreidae
Iwatsuki, Y. and P.C. Heemstra. 2001. Gerres phaiya: a new species of gerreid fish (Teleostei: Perciformes: Gerreidae)
from India, with comments on Gerres poieti and the Gerres erythrourus complex. Copeia 2001(4): 1043–
1049.
Iwatsuki, Y., S. Kimura, H. Kishimoto and T. Yoshino. 1996. Validity of the gerreid fish, Gerres macracanthus Bleeker,
1854, with designation of lectotype, and designation of a neotype for G. filamentosus Cuvier, 1829. Ichthyol.
Res. 43(4): 417–429.
Iwatsuki, Y., S. Kimura and T. Yoshino. 1999a. Description of Gerres chrysops sp. nov. from Thailand and redescription
of Gerres setifer (Hamilton, 1822) and G. decacanthus (Bleeker, 1865) (Perciformes: Gerreidae). Ichthyol.
Res. 46(1): 27–41.
Iwatsuki, Y., S. Kimura and T. Yoshino. 1999b. Redescription of Gerres baconensis (Evermann & Seale, 1907), G.
equulus Temminck & Schlegel, 1844 and G. oyena (Forsskål, 1775), included in the “G. oyena complex”, with
notes on other related species (Perciformes: Gerreidae). Ichthyol. Res. 46(4): 377–395.
Iwatsuki, Y., S. Kimura and T. Yoshino. 2001. Redescription of Gerres longirostris (Lacepède, 1801) and Gerres
oblongus Cuvier in Cuvier and Valenciennes, 1830, included in the Gerres longirostris complex (Perciformes:
Gerreidae). Copeia 2001(4): 954–965.
Iwatsuki, Y., S. Kimura and T. Yoshino. 2007. A review of the Gerres subfasciatus complex from the Indo-West Pacific,
with three new species (Perciformes: Gerreidae). Ichthyol. Res. 54(2): 168–185.
Family Haemulidae
Johnson, J.W., J.E. Randall and S.F. Chenoweth. 2001. Diagramma melanacrum, new species of haemulid fish from
Indonesia, Borneo and the Philippines with a generic review. Mem. Qd. Mus. 46(2): 657–676.
Iwatsuki, Y. and B.C. Russell. 2006. Revision of the genus Hapalogenys (Teleostei: Perciformes) with two new species
from Indo-West Pacific. Mem. Mus. Vict. 63(1): 29–46.
Iwatsuki, Y., U. Satapoomin and K. Amaoka. 2000. New species: Hapalogenys merguiensis (Teleostei; Perciformes)
from Andaman Sea. Copeia 2000(1): 129–139.
McKay, R.J. and U. Satapoomin. 1994. Pomadasys andamanensis, a new species of haemulid fish from Thailand.
Phuket mar. biol. Cent. Res. Bull. 59: 1–4.
Satapoomin, U and J.E. Randall. 2000. Plectorhinchus macrospilus, a new species of sweetlip (Perciformes: Haemulidae)
from the southwestern Thailand, Andaman Sea. Phuket mar. biol. Cent. Res. Bull. 63: 9–16.
Family Nemipteridae
75
The fishes of southwestern Thailand
Russell, B.C. 1986. Review of the western Indian Ocean species of Nemipterus Swainson 1839, with description of a
new species. Senckenberg. biol. 67(1/3): 19–35.
Russell, B.C. 1991. On the validity of Nemipterus furcosus (Valenciennes) (Nemipteridae). Cybium 15(1): 35–41.
Russell, B.C. and D. Golani. 1993. A review of the fish genus Parascolopsis (Nemipteridae) of the western Indian Ocean,
with description of a new species from the northern Red Sea. Israel J. Zool. 39: 337–347.
Family Polynemidae
Feltes, R.M. 1991. Revision of the polynemid fish genus Filimanus, with description of two new species. Copeia
1991(2): 302–322.
Motomura, H. 2002. Revision of the Indo-Pacific threadfin genus Polydactylus (Perciformes, Polynemidae) with a key
to the species. Bull. Natl. Sci. Mus., Ser. A. 28(3): 171–194.
Motomura, H., Y. Iwatsuki, S. Kimura and T. Yoshino. 2001. A new species, Polydactylus siamensis, from Thailand and
redescription of P. plebeius (Broussonet, 1782) with designation of a neotype (Perciformes: Polynemidae).
Ichthyol. Res. 48(2): 117–126.
Motomura, H., Y. Iwatsuki and T. Yoshino. 2002. Revision of the Indo-Pacific polynemid fish genus Eleutheronema
(Teleostei: Perciformes). Ichthyol. Res. 49(1): 47–61.
Motomura, H., U. Satapoomin and Y. Iwatsuki. 2000. A new record of the threadfin, Filimanus perplexa Feltes, 1991
(Perciformes: Polynemidae), from the Andaman Sea, Thailand. Phuket mar. biol. Cent. Res. Bull. 63: 17–20.
Family Sciaenidae
Sasaki, K. 1992. Two new and two resurrected species of sciaenid genus Johnius (Johnius) from the west Pacific. Jap.
J. Ichthyol. 39(3): 191–199.
Sasaki, K. 1996. Sciaenid fishes of the Indian Ocean (Teleostei, Perciformes). Mem. Fac. Sci. Kochi Univ. Ser. D (Biol.)
16/17: 83–95.
Family Mullidae
Randall, J.E. 2004. Revision of the goatfish genus Parupeneus (Perciformes: Mullidae), with descriptions of two new
species. Indo-Pacific Fishes 36: 1–64, 16 pls.
Family Kuhliidae
Randall, J.E. and H.A. Randall. 2001. Review of the fishes of the genus Kuhlia (Perciformes: Kuhliidae) of the central
Pacific. Pac. Sci. 55(3): 227–256.
Family Pomacentridae
Allen, G.R. 1991. Damselfishes of the World. Mergus Publishers, Melle, Germany. 271 p.
Allen, G.R. and A.R. Emery. 1985. A review of the pomacentrid fishes of the genus Stegastes from the Indo-Pacific, with
descriptions of two new species. Indo-Pacific Fishes 3: 1–31, 3 pls.
Allen, G.R. and M.V. Erdmann. 2005. Chromis xouthos, a new species of damselfish (Pomacentridae) from the east
Andaman Sea and central Indian Ocean. Aqua, J. Ichthyol. Aquat. Biol. 10(3): 89–94.
Allen, G.R. and J.E. Randall. 2002. A review of the leucogaster species complex of the Indo-Pacific pomacentrid genus
Amblyglyphidodon, with descriptions of two new species. Aqua, J. Ichthyol. Aquat. Biol. 5(4): 139–152.
Randall, J.E. and G.R. Allen. 2005. Neopomacentrus sororius, a new species of damselfish from the Indian Ocean, with
description of a neotype for its sister species, N. azysron (Bleeker). Aqua, J. Ichthyol. Aquat. Biol. 10(2): 73–
80.
Family Labridae
Gomon, M.F. 2006. A revision of the labrid fish genus Bodianus with descriptions of eight new species. Rec. Aust. Mus.
Suppl. 30: 1–133.
Randall, J.E. 1996. Second revision of the labrid fish genus Leptojulis, with descriptions of two new species. IndoPacific Fishes. 24: 1–20, 3 pls.
Randall, J.E. 1999. Revision of the Indo-Pacific labrid fishes of the genus Pseudocheilinus, with descriptions of three
new species. Indo-Pacific Fishes. 28: 1–34, 2 pls.
Randall, J.E. 1999. Revision of the Indo-Pacific labrid fishes of the genus Coris, with descriptions of five new species.
Indo-Pacific Fishes. 29: 1–74, 22 pls.
Randall, J.E. 2000. Revision of the Indo-Pacific labrid fishes of the genus Stethojulis, with descriptions of two new
species. Indo-Pacific Fishes. 31: 1–42, 6 pls.
Randall, J.E. and Lubbock, R. 1981. Labrid fishes of the genus Paracheilinus, with descriptions of three new species
from the Philippines. Jap. J. Ichthyol. 28(1): 19–30, 2 pls.
Randall, J.E. and M.M. Smith. 1982. A review of the labrid fishes of the genus Halichoeres of the western Indian Ocean,
with descriptions of six new species. Ichthyol. Bull. J.L.B. Smith Inst. Ichthyol. 45: 1–26, 6 pls.
Family Scaridae
Satapoomin, U., R.H. Kuiter and J.E. Randall. 1994. First Record of the parrotfish Scarus viridifucatus from Thailand
(the Andaman Sea) and Indonesia. Phuket mar. biol. Cent. Res. Bull. 59: 5–9.
76
Phuket mar. biol. Cent. Res. Bull.
Westneat, M., U. Satapoomin and J.E. Randall. 2007. Scarus maculipinna, a new species of parrotfish (Perciformes,
Scaridae) from eastern Indian Ocean. Zootaxa. 1628: 59–68.
Family Ammodytidae
Ida, H., P. Sirimontaporn and S. Monkolprasit. 1994. Comparative morphology of the fishes of the family Ammodytidae,
with description of two new genera and two new species. Zool. Stud. 33(4): 251–277.
Family Tripterygiidae
Holleman, W. 2005. A review of the triple fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the western Indain
Ocean, with descriptions of four new species. Smithiana Bulletin. 5: 1–25, 2 pls.
Holleman, W. 2006. Fishes of the Helcogramma steinitzi species group (Blennioidei: Tripterygiidae) from the Indain
Ocean, with descriptions of two new species. Aqua, J. Ichthyol. Aquat. Biol. 11(3): 89–104.
Williams, J.T. and J.C. Howe. 2003. Seven new species of the triplefin genus Helcogramma (Tripterygiidae) from the
Indo-Pacific. Aqua, J. Ichthyol. Aquat. Biol. 7(4): 151–176.
Family Blenniidae
Bath, H. 1992. Revision der gattung Praealticus Schultz & Chapman 1960 (Pisces: Blenniidae). Senckenberg. biol. 72(4/
6): 237–316.
Carlson, B.A. 1980. A new Indo-Pacific fish of the genus Cirripectes (Blenniidae, Salariini). Pac. Sci. 34(4): 407–414.
Smith-Vaniz, W.F. 1976. The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae). The Academy of Natural
Sciences of Philadelphia. Monograph 19. vii + 196 p.
Smith-Vaniz, W.F., U. Satapoomin and G.R. Allen. 2001. Meiacanthus urostigma, a new fangblenny from the northeastern
Indian Ocean, with discussion and examples of mimicry in species of Meiacanthus (Teleostei: Blenniidae:
Nemophini). Aqua, J. Ichthyol. Aquat. Biol. 5(1): 25–43.
Springer, V.G. 1988. The Indo-Pacific blenniid fish genus Ecsenius. Smithson. Contr. Zool. 465. 134 p., 14 pls.
Springer, V.G. and M.F. Gomon. 1975. Revision of the blenniid fish genus Omobranchus with descriptions of three new
species and notes on other species of the tribe Omobranchini. Smithson. Contr. Zool. 177. 135 p.
Springer, V.G. and J.E. Randall. 1999. Ecsenius polystictus, new species of blenniid fish from Mentawai Islands,
Indonesia, with notes on other species of Ecsenius. Revue fr. Aquariol. 26(1–2): 39–48.
Springer, V.G. and J.T. Williams. 1994. The Indo-west Pacific blenniid fish genus Istiblennius reappraised: A revision of
Istiblennius, Blenniella, and Paralticus, new genus. Smithson. Contr. Zool. 565. 193 p.
Williams, J.T. 1988. Revision and phylogenetic relationships of the blenniid fish genus Cirripectes. Indo-Pacific Fishes.
17: 1–78, 7 pls.
Family Callionymidae
Fricke, R. 1983. Revision of the Indo-Pacific genera and species of the dragonet family Callionymidae. Theses Zoologicae
Vol. 3. 774 p.
Family Gobiidae
Akihito and Meguro, K. 2000. Review of the Gobiid genus Cristatogobius found in Japan with description of a new
species. Ichthyol. Res. 47(3): 249–261.
Darumas, U. and P. Tantichodok. 2002. A new species of mudskipper (Gobiidae: Oxucercinae) from southern Thailand.
Phuket mar. biol. Cent. Res. Bull. 64: 101–107.
Hoese, D.F. and H.K. Larson. 1994. Revision of the Indo-Pacific gobiid fish genus Valenciennea, with descriptions of
seven new species. Indo-Pacific Fishes. 23: 1–71, 6 pls.
Lachner, E.A. and S.J. Karnella. 1980. Fishes of the Indo-Pacific fish genus Eviota with descriptions of eight new species
(Teleostei: Gobiidae). Smithson. Contr. Zool. 315. 127 p.
Lachner, E.A. and J.F. McKinney. 1978. A revision of the Indo-Pacific fish genus Gobiopsis with descriptions of four
new species (Pisces: Gobiidae). Smithson. Contr. Zool. 262. 52 p.
Larson, H.K. 1999. A review of the mangrove goby genus Hemigobius (Gobioidei, Gobiidae, Gobionellinae). The Beagle,
Records of the Museums and Art Galleries of the Northern Territory. 15: 23–42.
Larson, H.K. 1999. Allocation to Calamiana and redescription of the fish species Apocryptes variegates and Vaimosa
mindora (Gobioidei: Gobiidae: Gobionellinae), with description of a new species. Raffles Bull. Zool. 47(1):
257–281.
Larson, H.K. 2001. A revision of the gobiid fish genus Mugilogobius (Teleostei: Gobioidei). Rec. West. Aust. Mus.
Suppl. No. 62: 1–233.
Larson, H.K. 2005. A revision of the gobiid genus Stigmatogobius (Teleostei: Gobiidae), with descriptions of two new
species. Ichthyol. Explor. Freshwaters. 16(4): 347–370.
Larson, H.K. 2009. Review of the gobiid fish genera Eugnathogobius and Pseudogobiopsis (Gobioidei: Gobiidae:
Gobionellinae), with descriptions of three new species. Raffles Bull. Zool. 57(1): 127–181.
Murdy, E.O. 1985. A review of the gobiid fish genera Exyrias and Macrodontogobius, with description of a new species
of Exyrias. Indo-Pacific Fishes. 10: 1–14, 2 pls.
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The fishes of southwestern Thailand
Murdy, E.O. 1989. A taxonomic revision and cladistic analysis of the oxudercine gobies (Gobiidae: Oxudercinae). Rec.
Aust. Mus. Suppl. 11: 1–93.
Murdy, E.O. and D.F. Hoese. 1985. Revision of the gobiid fish genus Istigobius. Indo-Pacific Fishes 4: 1–41, 3 pls.
Polunin, N.V.C. and R. Lubbock. 1979. Five new prawn-associated gobies (Teleostei: Gobiidae) of the genus Amblyeleotris.
Bull. Br. Mus. Nat. Hist. (Zool.). 36(4): 239–249.
Randall, J.E. 2001. Five new Indo-Pacific gobiid fishes of the genus Coryphopterus. Zool. Stud. 40(3): 206–225.
Randall, J.E. and D.W. Greenfield. 2001. A preliminary review of the Indo-Pacific gobiid fishes of the genus Gnatholepis.
Ichthyol. Bull. J.L.B. Smith Inst. Ichthyol. 69: 1–17, 2 pls.
Randall, J.E. and D.F. Hoese. 1985. Revision of the Indo-Pacific dartfishes, genus Ptereleotris (Perciformes: Goioidei)
Indo-Pacific Fishes. 7: 1–36, 4 pls.
Shibukawa, K. and G.R. Allen. 2007. Review of the cheek-spine goby genus Gladiogobius (Actinopterigii, Perciformes,
Gobiidae), with descriptions of two new species from the Indo-Pacific. Bull. Natl. Mus. Nat. Sci., Ser. A,
33(4): 193–206.
Shibukawa, K. and U. Satapoomin. 2006. Myersina adonis, a new species of shrimp-associated goby (Pisces: Perciformes:
Gobiidae) from the Andaman Sea. Bull. Natl. Sci. Mus., Ser. A, 32(1): 29–37.
Shibukawa, K., T. Suzuki, H. Senou and K. Yano. 2005. Records of three shrimp-goby species (Teleostei, Perciformes,
Gobiidae) from the Ryukyu Archipelago, Japan. Bull. Natl. Sci. Mus., Ser. A, 31(4): 191–204.
Satapoomin, U. and R. Winterbottom. 2002. Redescription of the gobioid fish Cryptocentrus pavoninoides (Bleeker,
1849), with notes on sexual dichromatism in shrimp gobies. Aqua, J. Ichthyol. Aquat. Biol. 5(2): 53–64.
Winterbottom, R. 2002. A redescription of Cryptocentrus crocatus Wongratana, a redefinition of Myersina Herre
(Acanthopterygii; Gobiidae), a key to the species, and comments on relationships. Ichthyol. Res. 49(1): 69–
75.
Winterbottom, R. and M. Burridge. 1993. Revision of the species of Priolepis possessing a reduced transverse pattern
of cheek papillae and no predorsal scales (Teleostei: Gobiidae). Can. J. Zool. 71: 494–514.
Winterbottom, R. and A.S. Harold. 2005. Gobiodon prolixus, a new species of gobiid fish (Teleostei: Perciformes:
Gobiidae) from the Indo-west Pacific. Proc. Biol. Soc. Wash. 118(3): 582–589.
Winterbottom, R. and L. Southcott. 2007. Two new species of the genus Trimma (Percomorpha: Gobiidae) from western
Thailand. Aqua, Int. J. Ichthyol. 13(2): 69–76.
Family Siganidae
Woodland, D.J. 1990. Revision of the fish family Siganidae with descriptions of two new species and comments on
distribution and biology. Indo-Pacific Fishes. 19: 1–136, 11 pls.
Family Acanthuridae
Randall, J.E. 1993. Acanthurus tristis, a valid Indian Ocean surgeonfish (Perciformes: Acanthuridae). J.L.B. Smith Inst.
Ichthyol. Spec. Publ. 54: 1–8.
Randall, J.E. and K.D. Clements. 2001. Second revision of the surgeonfish genus Ctenochaetus (Perciformes: Acanthuridae),
with descriptions of two new species. Indo-Pacific Fishes. 32: 1–33, 6 pls.
Family Sphyraenidae
Doiuchi, R. and T. Nakabo. 2005. The Sphyraena obtusata group (Perciformes: Sphyraenidae) with a description of a
new species from southern Japan. Ichthyol. Res. 52(2): 132–151
Order Pleuronectiformes
Menon, A.G.K. 1977. A systematic monograph of the tongue soles of the genus Cynoglossus Hamilton-Buchanan
(Pisces: Cynoglossidae). Smithson. Contr. Zool. 238. 129 p.
Randall, J.E. and M. Desoutter-Meniger. 2007. Review of the soles of the genus Aseraggodes (Pleuronectiformes:
Soleidae) from the Indo-Malayan region, with descriptions of nine new species. Cybium. 31(3): 301–331.
Randall, J.E. and J.W. Johnson. 2007. Revision of the soleid fish genus Pardachirus. Indo-Pacific Fishes. 39: 1–22, 4 pls.
Order Tetraodontiformes
Hutchins, J.B. 1997. Review of the monacanthid fish genus Paramonacanthus, with descriptions of three new species.
Rec. West. Aust. Mus. Suppl. No. 54. 57 p.
Leis, J.M. 2006. Nomenclature and distribution of the species of the porcupinefish family Diodontidae. Mem. Mus.
Vict. 63(1): 77–90.
Randall, J.E. and W. Klausewitz. 1973. A review of the triggerfish genus Melichthys, with description of a new species
from the Indian Ocean. Senckenberg. biol. 54(1/3): 57–69.
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