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Phuket mar. biol. Cent. Res. Bull. 70: 1–6 (2011) ON THE IDENTITY OF THE MANGROVE CRAB, PARACLEISTOSTOMA ERIOPHORUM NOBILI, 1903 (CRUSTACEA: BRACHYURA: CAMPTANDRIIDAE) Peter K. L. Ng 1, 2, Cheryl G. S. Tan 2 and Rueangrit Promdam 3 1 Tropical Marine Science Institute and Raffles Museum of Biodiversity Research, National University of Singapore, 14, Science Drive 4, Singapore 117543, Republic of Singapore. 2 Department of Biological Sciences, National University of Singapore, 14, Science Drive 4, Singapore 119260, Republic of Singapore. 3 Reference Collection, Phuket Marine Biological Center, Phuket Marine Biological Center, P.O. Box 60, Phuket, 83000, Thailand. (r_promdam@yahoo.com) Corresponding author: P. K. L. Ng, e-mail: dbsngkl@nus.edu.sg ABSTRACT: The identity of the poorly known camptandriid mangrove crab Paracleistostoma eriophorum Nobili, 1903, is clarified. It is shown to be a senior synonym of Paracleistostoma tweediei Tan & Humpherys, 1995, and the taxonomy of the species is discussed, with the range of the species extended to Thailand. Notes on its ecology are also provided. INTRODUCTION The camptandriid genus Paracleistostoma De Man, 1895, is currently represented by eight species (Rahayu & Ng, 2003; Ng et al., 2008). Ng et al. (2008: 233, 234) commented that the poorly known Paracleistostoma eriophorum Nobili, 1903, was actually a senior synonym of Paracleistostoma tweediei Tan & Humpherys, 1995, but did not elaborate. We here redescribe and figure the type of Paracleistostoma eriophorum Nobili, 1903, and comment on its taxonomic relationships. MATERIALS AND METHODS The terminology follows that in Tan & Ng (1999). Measurements provided, in millimetres, are of the carapace length and width respectively. Specimens are deposited in the Museo Regionale di Scienze Naturali (MZUT), Sezione di Zoologia, Turin, Italy; Zoological Reference Collection (ZRC) of the Raffles Museum of Biodiversity Research, National University of Singapore, Singapore; Phuket Marine Biological Centre (PMBC), Phuket, Thailand; and Queensland Museum (QM), Brisbane, Australia. Paracleistostoma eriophorum Nobili, 1903 (Figs. 1, 2) Paracleistostoma eriophorum Nobili, 1903: 23. - Manning & Holthuis, 1981: 209 (list) - Ng et al., 2008: 233, 234. Paracleistostoma wardi - Yang, 1979: 39 (list). - Harminto, 1988: 88 (nec P. wardi Rathbun, 1926). - Tan & Ng, 1994: 83 (list). Paracleistostoma tweediei Tan & Humpherys, 1995: 251, figs. 1–3. Paracleistostoma tweediei Tan & Ng, 1995: 608. Material examined. - Holotype male of P. eriophorum (7.5 x 10.0 mm) (MZUT Cr1200, ex 1531), Buntal, Sarawak, East Malaysia, don. R. Shelford, 1902. Others - SINGAPORE: 1 male (holotype of P. tweediei) (7.9 x 10.2 mm) (ZRC 1987.57), 8 males (paratypes of P. tweediei) (5.3 x 6.7 mm, 6.4 x 8.3 mm, 6.6 x 8.4 mm (ZRC 1987.60–62), 7.4 x 9.4 mm, 7.5 x 9.7 mm, 7.5 x 10.0 mm, 8.3 x 10.7 mm, 8.8 x 11.5 mm (ZRC 1987.57–59), Lim Chu Kang mangrove, coll. P. K. L. Ng & S. Harminto, 13 September 1986; 1 male (7.2 x 9.3 mm) (ZRC 1987.1382), Woodlands, Singapore, coll. S, Harminto, 2 June 1987. PENINSULAR MALAYSIA: 18 males (ZRC 1998.1007), Matang mangroves, Perak, coll. C. G. S. Tan, 8 June 1995; 4 males (5.0–6.9 x 6.1– 8.8 mm) (ZRC 1998.998), Matang mangroves, Perak, coll. A. Ramli, M. Faizal, C. G. S. Tan, 10 June 1995. BRUNEI: 1 male (9.3 x 11.9 mm) (QM W19967), Batu Marang, coll. N. A. Mackenzie, 2 Phuket mar. biol. Cent. Res. Bull. Figure 1. Paracleistostoma eriophorum Nobili, 1903. A, holotype male (7.5 x 10.0 mm) (MZUT Cr1200, ex 1531), Sarawak; B, C, male (7.6 x 9.6 mm) (ZRC), Thailand. A, B, overall views; C, left G1 in situ. March 1993. THAILAND: 1 male (7.6 x 9.6 mm), 1 female (6.5 x 8.3 mm) (ZRC 2010.0071), 1 male (6.9 x 8.5 mm) (ZRC 2010.0072); 3 males (10.9 x 8.3 mm, 10.4 x 8.0 mm, 10.0 x 7.7 mm), 3 females (12.0 x 9.0 mm, 9.0 x 7.0 mm, 8.8 x 6.8 mm) (PMBC 23738), mangroves, Khanom District, Nakhon Srithammarat, Gulf of Thailand, coll. R. Promdam & A, Kaenphet, 23 June 2007. Type locality. - Buntal, Sarawak. Diagnosis (holotype male of P. eriophorum). Carapace surface smooth, slightly convex, broadly quadrangular, broader than long (1.2–1.3 times); regions indistinct, epigastric lobes moderately low; metagastric groove shallow. Front almost straight from dorsal view, upper surface slightly concave 3 Identity of the mangrove crab Figure 2. Paracleistostoma eriophorum Nobili, 1903. A, G–I, holotype male (7.5 x 10.0 mm) (MZUT Cr1200, ex 1531), Sarawak; B–F, male (7.9 x 10.2 mm) (ZRC 1987.57), holotype of P. tweediei Tan & Humpherys, 1995, Singapore. A, B, male abdomen; C, frontal view showing epistome; D, anterior part of male thoracic sternum; E, outer view of left chela; F, G, dorsal view of right carpus of cheliped; H, I, left G1 (setae not drawn). Scales: A, G = 0.5 mm; H, I = 1.0 mm. medially, anterolateral angles of front rounded; frontomedial projection triangular, apex rounded; orbital margins setose; supraorbital margin weakly granular, sinuous; infraorbital margin straight, finely granular along central third, inner infraorbital tooth well developed, distinct from rest of infraorbital margin; suborbital ridge defined by line of granules; posterolateral border and surfaces of carapace with dense setae; posterolateral margin lined with small granules, completely obscured by pubescence; posterior carapace margin straight. Anterior and posterior halves of epistome separated by median transverse ridge; postero medial-tooth broadly triangular, ending in small, pointed extremity, posterior margin on either side almost straight; sulcus separating epistome from anterior 4 Phuket mar. biol. Cent. Res. Bull. pterygostomian margin indistinct. Third maxilliped ischium broadly rectangular, outer distal margin with row of long setae along outer half; merus with well developed outer distal angle which is bilobed, outer surface uneven, inverted V shaped sulci on outer surface indistinct. Chelipeds subequal, enlarged; carpus subglobular, outer surface almost smooth, inner surface sparsely granular, inner dorsal edge finely beaded with granules, appearing subcarinate, inner ventral edge with large triangular granules, with short row of long setae. Ambulatory legs relatively long, densely setose; first and last pairs of ambulatory legs almost equal in length, first pair with setae only on dorsal surface of merus. Abdominal somites 2–5 completely fused, sutures not discernible, suture between somites 2 and 3 visible only as short, shallow grooves on either side of abdomen; somite 1 broader than somite 2, with transverse mdian ridge; somite 6 shorter than telson, lateral margin sinuous. G1 recurved, apex large, globular, with short but distinct submedian spine-like distal appendage. Remarks. - Nobili’s (1903) description of Paracleistostoma eriophorum was relatively detailed but he did not provide a figure. He distinguished P. eriophorum from P. depressum and P. cristatum by the unusually tomentose ambulatory legs and the presence of two carinae of long granules on the inferior and superior margins of the outer surface of the cheliped palm. Nobili (1903) also mentioned that the posterior carapace, including the median branchial region and the area around the intestinal region, bears a thick, short felt like tomentum, and the ambulatory legs were covered with long, woolly setae with the dactyli bearing sulci and setae. From his comparisons, Serène (1974:64) noted “... it is possible that eriophorum is a Leipocten or Baruna”. After examining the type, there is no doubt that P. eriophorum is a species of Paracleistostoma. As noted by Ng et al. (2008), P. eriophorum Nobili, 1903, is a senior synonym of P. tweediei Tan & Humpherys (1995). The type specimens of both species are of similar size, and agree closely in all characters. The holotype male of P. eriophorum is not in a good condition (Fig. 1A) with most of the appendages detached. Most of the setae on the carapace have also been lost, although one can discern where the main setose areas were (Fig. 1A). The distal part of the G1 of the holotype male is somewhat less swollen (Fig. 2H, I) compared to the recent material (Fig. 1C), but this is almost certainly because of its age and some dehydration. Paracleistostoma eriophorum appears to be close to P. depressum and is distinguished from all its congeners by the unique structure of the G1, which has the distal appendage modified to form a spine like process. It is one of the two Paracleistostoma species having the inner ventral edge of the carpus lined with sharp granules, the only other Paracleistostoma species exhibiting this condition being P. longimanum. Also, in P. eriophorum, the ridge below the infraorbital margin is formed by a raised line of granules. This condition is also found in P. wardi. Neither Nobili (1903) nor Tan & Humpherys (1991) described the epistome and the anterior sternal structure, and these are included in the present description (Fig. 2C, D). Only one female specimen of this species (from Thailand) is known. It closely resembles the male in all nonsexual features except that the chelae are even more slender, and the patches of setae on the posterolateral regions are less clearly demarcated, partially because there are also short setae on the dorsal parts of the carapace which are normally glabrous in males. The female abdomen is subcircular, covers the thoracic sternum, and all the somites and telson are mobile. Harminto (1988) recorded this species as P. wardi (Rathbun, 1926). His specimens were examined and compared with Australian specimens of P. wardi in the ZRC and were found to differ significantly from the latter species in terms of the structures of the G1, epistome and male cheliped carpus. Tan & Ng (1994) had incorrectly recorded this species as P. wardi in their list of mangrove Brachyura. The figure of the frontal region and abdomen provided by Tan & Humpherys (1995) is somewhat inaccurate in that the inner infraorbital tooth is not clearly shown (Tan & Humpherys, 1995: Fig. 2A) and the sutures between abdominal segments 3–5 are shown to be present as notches 5 Identity of the mangrove crab on either side of the abdomen (Tan & Humpherys, 1995: Fig. 2G). In fact, only the suture between somites 3 and 4 is visible as a notch on either side of the abdomen. The above features are refigured here (Fig. 2A, B). As discussed by Ng et al. (2008), there is a nomenclatural problem with the names Paracleistostoma tweediei Tan & Humpherys, 1995, and Paracleistostoma tweediei Tan & Ng, 1995. Ng et al. (2008: 234) noted that “Both are objective synonyms as they are based on the same holotype. The problem arose because C. G. S. Tan and P. K. L. Ng originally intended to publish the new species, and the paper was prepared for a regional symposium in 1994 (Tan & Ng, 1995). At the same time, P. J. F. Davie and A. Humpherys had independently also discovered the species. As it was clear that the two taxa in question were conspecific, P. J. F. Davie and P. K. L. Ng left it to C. G. S. Tan and A. Humpherys to finish the new species description on their own (Tan & Humpherys, 1995). In early 1995, P. K. L. Ng asked the editors of the symposium volume to have the paragraph on the new Paracleistostoma in their paper deleted, but although they agreed, the changes were not made, no proofs were sent, and the volume was published in 1995 with the problem paragraph still intact. This mistake nevertheless validates P. tweediei C. G. S. Tan & Ng, 1995. In any case, the symposium article came out later than the paper in the Raffles Bulletin of Zoology. Since the symposium volume did not have a publication date, under the Code, it should be regarded as published on 31 December 2005. This gives P. tweediei C. G. S. Tan & Humpherys, 1995, priority over P. tweediei C. G. S. Tan & Ng, 1995.” Ecology. - In Perak, Malaysia, Paracleistostoma eriophorum was collected from the Matang mangroves along with specimens of P. depressum and the varunid Metaplax elegans De Man, 1888. These were all found in burrows in soft, brown mud, close to the edge of the river. In the Khanom District, Thailand, P. eriophorum were found in mangrove habitats near rivers, with burrows typically occurring in the areas between nearby mangrove trees. Sympatric species there, include the sesarmids Episesarma palawanense (Rathbun, 1914), Parasesarma ungulatum (H. Milne Edwards, 1853), Perisesarma eumolpe (De Man, 1895) and P. indiarum (Tweedie, 1940). Distribution. - Buntal, Sarawak, East Malaysia; Brunei, Peninsular Malaysia, Singapore, and now for the first time from the eastern coast of Thailand. ACKNOWLEDGEMENTS The authors thank Giovanni Balma of the University of Turin for his help in obtaining the type of P. eriophorum. Part of the study was supported by research grants to the first author from the National University of Singapore. The third author was supported by the TOTAL Corporate Foundation, TOTAL E&P Thailand and the TRF/BIOTEC Special Program for Biodiversity Research and Training Grant BRT R_149023 and he would like to thank Arwut Kaenphet, staff of Walailak University, who worked closely with him during the field trips to the mangroves of Khanom. REFERENCES Harminto, S. 1988. Systematics of the family Ocypodidae Rafinesque, 1815, s. lat. (Crustacea: Decapoda: Brachyura) of Southeast Asia. Unpublished M.Sc. thesis, Department of Zoology, National University of Singapore. Manning, R.B. & L.B. Holthuis. 1981. West African Brachyuran Crabs (Crustacea: Decapoda). Smithson. Contr. Zool. 306: 1–379. Ng, P.K.L., D. Guinot & P.J.F. Davie. 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bull. Zool., Suppl. 17: 1–286. Nobili, G. 1903. Contributo alla fauna carcinologica di Borneo. Boll. Mus. Zool. Anat. comp. Torino. 18(447): 1–32, 3 figs. 6 Phuket mar. biol. Cent. Res. Bull. Rahayu, D.L. & P.K.L. Ng, 2003. The Camptandriidae of Irian Jaya, Indonesia, including the descriptions of two new species (Decapoda: Brachyura). J. Crust. Biol. 23(4): 951–962. Rathbun, M.J. 1926. Brachyuran crabs from Australia and New Guinea. Rec. Aust. Mus. 15(2): 177– 182, pls. 14–16. Serène, R. 1974. Note on the genera and species of the Camptandriinae Stimpson, 1858 (Decapoda, Brachyura: Ocypodidae). Treubia. 28(3): 59–117. Tan, C.G.S. & A. Humpherys. 1995. Paracleistostoma tweediei, a new species of camptandriine crab, from Singapore and Brunei mangroves (Crustacea: Decapoda: Brachyura: Ocypodidae). Raffles Bull. Zool. 43(1): 251–256. Tan, C.G.S. & P.K.L. Ng. 1995. The Camptandriinae of Singapore and Malaysia with description of one new species of Paracleistostoma De Man, 1895 (Crustacea: Decapoda: Brachyura: Ocypodidae). Proc 3rd ASEAN-Australia Symp. Living Coastal Resources Vol. 2: research papers. C. R. Wilkinson, S. Sudara & L. M. Chou (Eds.), Bangkok, pp. 607–611. Tan, C.G.S. & P.K.L. Ng. 1994. An annotated checklist of mangrove brachyuran crabs from Malaysia and Singapore. Hydrobiologia. 285: 75–84. Tan, C.G.S. & P.K.L. Ng. 1999. A revision of the genus Camptandrium Stimpson, 1858 (Crustacea: Decapoda: Brachyura: Camptandriidae). Raffles Bull. Zool. 47(1): 193–219. Yang, C. M. 1979. A list of Brachyura in the Zoological Reference Collection of the Department of Zoology. Unpublished checklist, Department of Zoology, University of Singapore, 60 pp. (mimeographed). Manuscript received: 20 October 2009 Accepted: 9 April 2010 Phuket mar. biol. Cent. Res. Bull. 70: 7–14 (2011) NEW RECORDS OF SPIDER CRABS OF THE GENERA CYRTOMAIA MIERS, 1886, AND PLATYMAIA MIERS, 1886 (DECAPODA: MAJOIDEA: INACHIDAE) FROM THE ANDAMAN SEA, THAILAND Rueangrit Promdam Department of Biology, Faculty of Science, Prince of Songkla University, Hatyai, Songkhla 90112, Thailand Email: r_promdam@yahoo.com ABSTRACT Two species of deep-water spider crabs, Cyrtomaia suhmii Miers, 1886, and Platymaia alcocki Miers, 1886, based on the materials of the BIOSHELF Project (1996–2000) are recorded for the first time from Thailand. It is also the first record of these two genera from the Andaman Sea coast of Thailand. Adult and juvenile forms of C. suhmii collected from the same locality confirm that C. curviceros Bouvier, 1915, is a junior synonym, the characters previously used to separate them explained by size and age. Key words: new records, spider crabs, Decapoda, Majoidea, Inachidae, Cyrtomaia, Platymaia, Thailand INTRODUCTION TAXONOMY Two rare spider crabs, Cyrtomaia suhmii Miers, 1886, and Platymaia alcocki Miers, 1886, were collected by the R.V. Chakratong Tongyai from the Andaman Sea during the BIOSHELF project surveys of the west coast of Thailand in 1996–2000. Based on Ng and Davie (2002), and Naiyanetr (2007), the species as well as the genera had not been previously reported from Thai waters. The present report serves to formally record these species, as well as to discuss variation in adult and juvenile specimens. Family Inachidae MacLeay, 1838 Genus Cyrtomaia Miers, 1886 Cyrtomaia suhmii Miers, 1886 (Figs. 1, 2A–F) MATERIALS AND METHODS Specimens examined are deposited in the Reference Collection of Phuket Marine Biological Center (PMBC). All measurements provided are pre-rostral lengths and carapace widths respectively. For details on all BIOSHELF stations, see Aungtonya et al. (2000). The classification used here follows Ng et al. (2008). Cyrtomaia suhmii Miers, 1886: 16, pl. 3, fig. 2, 2a–2c; Rathbun, 1918: 6; Richer de Forges and Guinot, 1990: 525; Richer de Forges and Ng, 2007: 56 (list); Ng et al., 2008: 111 (list). Cyrtomaia suhmi - Rathbun, 1893: 230; Griffin, 1974: 9; Guinot and Richer de Forges, 1982a: 1096; 1982b: 21, fig. 10, 11A–B, 23B; Griffin and Tranter, 1986b: 352, figs. 1, 2; Richer de Forges and Guinot, 1988: 42–43 fig. 2B, C, pl. 2F–G; Poore, 2004: 360; 361 fig. 108. Cyrtomaia Suhmi var. curviceros Bouvier, 1915b: 9–15, pls. 1. Cyrtomaja suhmi typica Serène and Lohavanijya, 1973: 46 (in key). 8 Phuket mar. biol. Cent. Res. Bull. Figure 1. Cyrtomaia suhmii, adult male (60.2 × 67.1 mm) (PMBC 19999), dorsal view of whole animal. Scale = 20 mm. Cyrtomaja suhmi curvicornis Serène and Lohavanijya, 1973: 45, 46 (in key). Cyrtomaia curviceros Sakai, 1976: 181; Guinot and Richer de Forges, 1982a: 1096; 1982b: 24, fig. 12A–D; Richer de Forges and Guinot, 1988: 42–43, fig. 2A; Ng et al., 2001: 13; 81 fig. 3h; Richer de Forges and Ng, 2007: 56 (list). not Cyrtomaia suhmi - Griffin and Brown. 1976: 252, fig. 6; Griffin and Tranter, 1986a: 30, fig. 91g [= C. griffini Richer de Forges and Guinot, 1990]. not Cyrtomaia suhmi typica Doflein, 1904 : 54– 55, pl. 19: figs. 1, 2 [= C. gaillardi Guinot and Richer de Forges, 1982]. Material examined PMBC 19999, 1 male (60.2 × 67.1 mm), BIOSHELF St. E8, 0832´N, 09604´E, Otter trawl, 488 m, coll. S. Bussarawit and C. Aungtonya, 06.02.1999; PMBC 20000, 2 juv. (11.8 × 11.5 mm, 8.6 × 8.2 mm), BIOSHELF St. J10, 0715´N, 09715´E, Agassiz trawl, 695–683 m, coll. C. Aungtonya & V. Vongpanich, 19.02.2000; PMBC 19929, 1 juv. (7.7 × 7.4 mm), 1 female (56.3 × 61.2 mm), BIOSHELF St. L10, 0645´N, 09723´E, Agassiz trawl, 707–651 m, coll. C. Aungtonya & V. Vongpanich, 21.02.2000 Diagnosis Carapace smooth, not granular or spinulate, with 2 long, subparallel protogastric spines. Pseudorostral spine divergent, arched. Basal antennal article cylindrical, armed with 3–5 spines (include internal spine). Orbital border smooth, intercalated orbital granule present (in young) or absent, without preocular spines (in adults). Ocular spine blunt. 9 New records on spider crabs Figure 2. Cyrtomaia suhmii: A–C, juvenile (11.8 × 11.5 mm) (PMBC 20000); D–F, adult male (60.2 × 67.1 mm) (PMBC 19999); A, D, dorsal view; B, E, anterior part of carapace showing rostrum; C, F, lateral view of carapace. Scale = 10 mm. Distribution Indo-West Pacific: Japan, Philippines, Indonesia, Australia, Thailand (Andaman Sea) and India. Remarks Cyrtomaia suhmii was described by Miers (1886) on the basis of an imperfect crushed young male specimen collected from Talaud Islands (Tulur 10 Phuket mar. biol. Cent. Res. Bull. Islands) (see Guinot & Richer de Forges 1982b: 22, fig. 10), Indonesia, by the Challenger Expedition. Subsequently, Cyrtomaia suhmii curviceros Bouvier, 1915, was described from Japan and recognized as a distinct species by Sakai (1976). The status of C. curviceros has long been uncertain. Griffin and Tranter (1986) treated both names as synonyms, while Richer de Forges and Guinot (1982a, 1988) argued that it may be a good species. Ng and Huang (1997) treated Taiwanese material of the species as C. curviceros (see also Ng et al., 2001), and until recently, was still regarded as a distinct species (see Richer de Forges and Ng, 2007). In their annotated checklist of extant brachyuran crabs of the world, Ng et al. (2008) treated C. suhmii curviceros as a synonym of C. suhmii but without comments. Examination of more material of other allied species like C. griffini Richer de Forges & Ng, 1990, has shown that the characters used to separate C. curviceros from C. suhmii are not valid at the species level (P. K. L. Ng, pers. comm., see also Richer de Forges and Ng, 2008). Specimens of various sizes were collected from the same Thai locality. The young specimens are similar to the original description and figure of C. suhmii by Guinot and Richer de Forges (1982b: 22, 23, fig. 10, 11A, B), especially with regards to the protogastric spines which are divergent (Miers, 1886: 16, pl. 3, fig. 2). On the other hand, large male specimens resemble C. curviceros (Fig. 1, 2D–F), although the number of the spines on the basal antennal article is slightly different. This difference, however, is not significant and can easily be explained by variation. The present series of specimens confirm that the differences which have been observed in the past between these two species are due to size and age, and that C. suhnii curviceros Bouvier, 1915, clearly is a junior synonym of C. suhmii Miers, 1886. Genus Platymaia Miers, 1886 Platymaia alcocki Rathbun, 1916 (Figs. 3, 4A–F) Platymaia wyville-thomsoni. - Wood-Mason and Alcock, 1891: 258; Alcock, 1895: 181– 182; Doflein, 1904: 59–69, 160–165, figs. 2–6; pls. 2, 20–23, 39, 43: figs. 5, 6; pl. Figure 3. Platymaia alcocki, adult male (71.0 × 77.5 mm) (PMBC 20367), dorsal view of whole animal. Scale = 20 mm. 11 New records on spider crabs Figure 4. Platymaia alcocki: A–C, adult male (71.0 × 77.5 mm); D–F, young male (25.9 × 30.4 mm) (PMBC 20367); A, D, dorsal view; B, E, anterior part of carapace showing rostrum; C, F, lateral view of carapace. Scale = 10 mm. 50: figs. 2, 5, 6 [not Platymaia wyvillethomsoni Miers, 1886]. Platymaia alcocki Rathbun, 1916: 530; 1918: 8; Serène and Lohavanijya, 1973: 48 (in key); Griffin, 1974: 27; Griffin and Tranter, 1986a: 48 (Pl. 4b; figs. 10a, b; 11c); 1986b: 357; Takeda and Webber, 2006: 194. Not Platymaia alcocki - Takeda and Miyake, 1969: 498–500, figs. 10a, b, 11a–c; Sakai, 1976: 12 Phuket mar. biol. Cent. Res. Bull. 176–177, fig. 94a; Dai and Yang, 1991: 120 [=Platymaia wyville-thomsoni Miers, 1886]. Material examined PMBC 20367, 4 male (71.0 × 77.5 mm, 25.9 × 30.4 mm, 20.6 × 22.7 mm, 20.2 × 22.8 mm), 5 female (19.4 × 21.4 mm, 19.0 × 21.4 mm, 19.0 × 22.1 mm, 18.7 × 21.2 mm, 18.4 × 20.3 mm), BIOSHELF St. J8, 0715´N, 09730´E, Agassiz trawl, 490–479 m, coll. S. C. Aungtonya & V. Vongpanich, 18.02.2000 Diagnosis Carapace tranversly subcircular, surface nearly smooth in adults, rostral spine subequal to interantennular spine, preocular spine absent; protogastric ridge with 2 tubercles, lacking spines (in adults); groove on each side of mesogastric region deep, clearly separating adjacent branchial regions; palm of male chelipeds more than twice as long as high, distally increasing in width; propodus of ambulatory legs flattened, 7.5 times width of fourth ambulatory leg. Remarks Platymaia alcocki can sometimes be misidentified with P. wywillethomsoni because young and juveniles of P. alcocki also have prominent spines on all the carapace regions (Alcock, 1895). Although all the spines on the dorsal carapace surface of adult P. alcocki are reduced to only tubercles (Fig. 4A–C), adult P. wywillethomsoni are still armed with one or two spines on each side of the protogastric regions even as adults (Griffin and Tranter, 1986a: 307, fig. 5b). The main diagnostic feature of P. alcocki is the deep groove on each side of mesogastric region, which separates the two adjacent branchial regions (Rathbun, 1916) (Fig. 4A, D). This character is useful even for young and adult specimens of P. alcocki and is clearly evident on the series of Thai specimens here. ACKNOWLEDGEMENTS The author would like to thanks Peter K. L. Ng of the Raffles Museum of Biodiversity Research, National University of Singapore, for reviewing the manuscript. Distribution Indian Ocean: Andaman Sea to eastern Africa REFERENCES Alcock, A. 1895. Materials for a carcinological fauna of India, No. 1: The Brachyura Oxyrhyncha. J. Asiat. Soc. Beng. 64: 157–291. Aungtonya, C., S. Thaipal and O. Tendal. 2000. A preliminary report on the Thai-Danish BIOSHELF surveys (1996–2000) of the west coast of Thailand, Andaman Sea. Phuket Mar. Biol. Cent. Res. Bull. 63: 53–76. Bouvier, E. L., 1915a. Décapodes Marcheurs (Reptantia) et Stomatopodes recueillis à I’île Maurice par M. Paul Carié. Bull. sc. Fr. Belg. (7)48(3): 178–318, figs. 1–42, pls. 4–7. Bouvier, E. L., 1915b. Étude sur un Cyrtomaia suhmi du Musée de Marseille. Annls. Mus. Hist. nat. Marseille. 15: 9–15, pl. 1. Dai, A.-Y. and S.-L. Yang. 1991. Crabs of China Seas. China Ocean Press, Beijing and Springer-Verlag, Berlin. 682 pp. Doflein, F. 1904. Brachyura. Wiss. Ergebn. Dt. Tiefsee Exped. ‘Valdivia’, 1898–1899, 1898–1899. 6: 1–314, figs 1–68, pls 1–58. Griffin, D.J.G. 1974. Spider crabs (Crustacea: Brachyura: Majidae) from the International Indian Ocean Expedition 1963–64. Smithson. Contr. Zool. 182: 1–35, figs 1–8, tables 1–6. Griffin, D.J.G. & D.E. Brown. 1976. Deepwater decapod Crustacea from eastern Australia: Brachyuran crabs. Rec. Aust. Mus. 30(11): 248–271, figs 1–10. 13 New records on spider crabs Griffin, D.J.G. & H.A. Tranter. 1986a. The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga Exped. Monogr. 39(C4): 1–335, text figs. 1–112, pls. 1–22. Griffin, D.J.G. & H.A. Tranter. 1986b. Some majid spider crabs from the deep Indo-West Pacific. Rec. Aust. Mus. 38(6): 351–371. Guinot, D., & B. Richer de Forges. 1982a. Nouvelles récoltes des genres Cyrtomaia Miers et Pleistacantha Miers (Crustacea, Decapoda, Brachyura). Bull. Mus. natn. Hist. nat., Paris. (3)4A(4), 1981 (1982): 1087–1124. Guinot, D., & B. Richer de Forges. 1982b. Révision du genre Indo-Pacifique Cyrtomaia Miers, 1886: Campagnes océanographiques du Challenger, de l’Albatross, du Siboga et du Vauban (Crustacea Decapoda Brachyura). Ann. Inst. Oceanogr., Paris. 58(1) 1982: 5–87. Miers, E.J. 1886. Report on the Brachyura collected by H.M.S. Challenger during the years 1873–76. In: Report on the Scientific Results of the Voyage of HMS Challenger during the years 1873– 76, Zool. 17(49): i–xii, 1–362, pls 1–29. Naiyanetr, P. 2007. Checklist of crustacean fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). 2nd ed. Office of Natural Resources and Environmental Policy and Planning. ONEP Biodiversity Series, vol. 19. 196 pp. Ng, P.K.L. and Davie, P.J.F. 2002. A checklist of the brachyuran crabs of Phuket and Western Thailand. Phuket Mar. Biol. Cent., Spec. Publ. 23(2): 369–384. Ng, P. K. L. and J. F. Huang, 1997. Unrecorded crabs (Crustacea: Decapoda: Brachyura) from Taiwan and Tungsha Islands, with description of a new genus and species of Xanthidae. Zool. Stud. 36(4): 261–276. Ng, P.K.L., Guinot, D and Davie, P. 2008. Systema Brachyuorum: Part I. An annotated checklist of extant Brachyuran crabs of the world. Raffles Bull. Zool., Suppl. 17: 1–286. Ng, P.K.L., C.H.Wang, P.H. Ho & H.T. Shih. 2001. An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Natl. Taiwan Mus. spec. Publ. Ser. 11: 1–86, 8 col. pls. Poore, G.C.B. 2004. Marine Decapod Crustacea of southern Australia: a guide to their identification. CSIRO Publishing. 574 pp. Rathbun, M.J. 1893. Descriptions of new genera and species of Crabs from the West Coast of North America and the Sandwich Islands. In: Scientific results of explorations by the U.S. Fish Commission Steamer Albatross. (No. 24). Proc. U.S. Natl. Mus. 16: 223–260. Rathbun, M.J. 1916. New Species of Crabs of the Families Inachidae and Parthenopidae. In: Scientific Results of the Philippine Cruise of the Fisheries Steamer Albatross, 1907–1910. (No. 34). Proc. U.S. Natl. Mus. 50: 527–559. Rathbun, M.J. 1918. Report on the spider crabs obtained by F.I.S. Endeavour on the coasts of Queensland, New South Wales, Victoria, South Australia and Tasmania. Zool. (biol.) Results Fish. Exp. Endeavour. 5(1): 1–29, pls. 1–15 Richer de Forges, B. & Guinot, D. 1988. Description de trois espèces de Cyrtomaia Miers, 1886, de Nouvelle Calédonie et des îles Chesterfield (Crustacea Decapoda Brachyura). Bull. Mus. natn. Hist, nat., Paris. 1988. 10(4)A(1): 39–55. Richer de Forges, B. & Guinot, D. 1990. A new Cyrtomaia, C. grifini, from Australia (Crustacea: Decapoda: Brachyura). Mem. Qld. Mus. 28: 523–530. Richer de Forges, B. & P.K.L. Ng. 2007. Notes on deep-sea spider crabs of the genus Cyrtomaia Miers, 1886, from the Philippines (Crustace: Decapoda: Brachyura: Majidae), with description of a new species. Raffles Bull. Zool., Suppl. 16: 55–65. Richer de Forges, B. and P. K. L. Ng, 2008. New records of deep-sea spider crabs of the genus Cyrtomaia Miers, 1886, from the Pacific Ocean, with description of a new species (Crustacea: Decapoda: Brachyura: Majidae). Zootaxa, 1861: 17–28. Sakai, T. 1976. Crabs of Japan and the adjacent seas. Kodansha, Tokyo: i–xxix, 1–773, text figs 1– 379, maps 1–3, pp. 1–16, pls. 1–251. 14 Phuket mar. biol. Cent. Res. Bull. Seréne, R. & P. Lohavanijaya. 1973. The Brachyura (Crustacea: Decapoda) collected by the NAGA Expedition, including a review of the Homolidae. Naga Rep. 4(4): 1–186, pls. 1–21. Takeda, M. & S. Miyake. 1969. Crabs from the East China Sea. III Brachygnatha Oxyrhyncha. J. Fac. Agric. Kyushu Univ. 15(4): 469–521, figs. 1–12, pls. 17–18. Takeda, M. & W.R., Webber. 2006. Crabs from the Kermadec Islands. In: Tomida, Y. et al. Proceedings of the 7th and 8th Symposia on Collection Building and Natural History Studies in Asia and the Pacific Rim. Natl. Sci. Mus. Monogr. 34: 191–237. Wood-Mason, J and A. Alcock. 1891. Natural history notes from H. M. Indian marine survey steamer Investigator, Commander R. F. Hoskyn, R.N., commanding. (No. 21). Note on the results of the last season’s deep-sea dredging. Ann. Mag. Nat. Hist. (6)7(39): 258–272, fig. 5. Manuscript received: 9 February 2011 Accepted: 1 August 2011 Phuket mar. biol. Cent. Res. Bull. 70: 15–27 (2011) NEW RECORDS OF THE BOBTAIL SQUID, EUPRYMNA HYLLEBERGI NATEEWATHANA, 1997 WITH DESIGNATION OF A NEOTYPE Charatsee Aungtonya1, Anuwat Nateewathana2, Ole Secher Tendal3 and Jaruwat Nabhitabhata4 1 Phuket Marine Biological Center, P.O. Box 60, Phuket 83000, Thailand 2 Department of Marine and Coastal Resources, Bangkok, Thailand 3 Zoological Museum, SNM, University of Copenhagen, Universitetsparken 15, DK-2100 Copenhagen, Denmark 4 Centre for Biodiversity of Peninsular Thailand (CBIPT), Faculty of Science, Prince of Songkla University, Hatyai, Songkhla 90112, Thailand Corresponding author: Charatsee Aungtonya Email: charatsee@gmail.com ABSTRACT: A total of 265 specimens, 118 males and 147 females, of the bobtail squid, Euprymna hyllebergi, are registered in the Reference Collection, Phuket Marine Biological Center, Thailand. Nearly all specimens were collected from Thai waters, the Andaman Sea and the Gulf of Thailand. The species is here reported as a new record for Myanmar waters and the distribution in the Indian Ocean has been extended to the west of the Andaman Islands. A male neotype is designated on a specimen collected from the original locality in order to replace the lost male holotype specimen. The neotype is deposited in the Reference Collection of Phuket Marine Biological Center, Thailand, as is the remaining female paratype, whereas selected specimens are deposited in National Science Museum Tokyo, Japan, U.S. National Museum, Washington D.C., USA and the Zoological Museum, SNM, University of Copenhagen, Denmark. Measurements of the neotype, paratype, and selected specimens are presented and discussed. Key word: bobtail squid, Euprymna hyllebergi, Thai waters, neotype INTRODUCTION Bobtail squids are small benthic cephalopods, belonging to the family Sepiolidae Leach, 1817, which contains about 14 genera and over 50 species. Sepiolid squids are distributed worldwide, ranging from tropical to temperate and sub-polar waters of all oceans (Nateewathana, 1997a). According to the most recent revision by Norman & Lu (1997) and a later paper on the genus by Nateewathana (1997b), 13 species have been referred to Euprymna Steenstrup, 1887. Six species, E. berryi Sasaki, 1929, E. hoylei Adam, 1986, E. hyllebergi Nateewathana, 1997, E. morsei (Verrill, 1881), E. scolopes Berry, 1913, and E. tasmanica (Peffer, 1884) are currently accepted in the genus. Two species, E. pusilla (Peffer, 1884) and E. schneehageni (Peffer, 1884), were placed as invalid while four species, E. albatrossae Voss, 1963, E. bursa (Pfeffer, 1884), E. phenax Voss, 1963, and E. stenodactyla (Grant, 1833), were treated as unresolved species (Norman & Lu, 1997). E. albatrossae, E. bursa, and E. phenax were known only from the types. The original type material of E. stenodactyla is lost, no further material has been found in Mauritius yet and the description of the species is poor (Norman & Lu, 1997). Furthermore, a nominal species, E. similis Sasaki, 1913, was considered a junior synonym of E. morsei (Norman & Lu, 1997). The species E. morsei, E. stenodactyla, E. berryi and E. hyllebergi, have been reported from Thai waters, the Andaman Sea and the Gulf of Thailand, by Chotiyaputta et al. (1992), Chotiyaputta (1993), Supongpan (1995), Nateewathana (1997b) and Nateewathana et al. (2001). 16 Phuket mar. biol. Cent. Res. Bull. According to Nateewathana et al. (2001) the occurrence of E. morsei, E. stenodactyla, and E. berryi in Thai waters was still uncertain since the voucher material could not be located. However, E. morsei and E. stenodactyla from the Andaman Sea and Myanmar waters have been registered at the PMBC Reference Collection in 2007. Future taxonomic study on those specimens is pending. All specimens of Euprymna hyllebergi deposited in the Reference Collection of Phuket Marine Biological Center, were checked and measured, but none were in accordance with the holotype. The first and the fourth authors contacted the cephalopod taxonomists who had previously studied the type material, but they all denied any knowledge about the fate of those types. We conclude that the holotype is lost. Euprymna hyllebergi is close to E. hoylei, but differs in the arrangement of suckers on the hectocotylized arm, the size and distribution of enlarged suckers on arms II–IV, and in the length of the tentacles (Nateewathana, 1997b: Fig. 3). E. hoylei is known from the tropical Western Pacific to North-West Australia (Norman & Lu, 1997). It is characterized by lacking enlarged suckers on arm II in mature males, having 3–4 enlarged suckers in dorsal and ventral rows of arm pair IV and ventral row only of arm pair III (Norman & Lu, 1997, Nesis, 1987). The suckers on arm II–IV of E. hyllebergi do not have any significantly or abruptly enlarged suckers (Nateewathana, 1997b). E. hyllebergi has a high number of stout papillae with a slitlike aperture in the distal half of the hectocotylus. The papillae of E. hyllebergi are crowded in rows of 4–6 suckers in the middle part, and reduced to two rows at the tip of the hectocotylus. The male characters are the most important features distinguishing the two species, E. hoylei and E. hyllebergi. Furthermore, the arrangement of the sessile suckers of the male is the only diagnostic feature to characterize the species of the genus, as well as other genera in the Family Sepiolidae. Hence it is necessary to select a male neotype of E. hyllebergi. The present report lists all specimens of this species in the Reference Collection, designates the male neotype of Euprymna hyllebergi in order to replace the lost male holotype and paratypes previously listed in Aungtonya et al. (2006, 2007), and describes specimens deposited in other museums. This study aims to facilitate future revision and taxonomic work on this genus and to update the known distribution of the species. MATERIALS AND METHODS The bobtail squids were obtained from fishing boats, research vessels, fish markets, and fish landings along the Andaman Sea coast of Thailand and the Gulf of Thailand as described by Nateewathana (1997b) and Nateewathana et al. (2001). Additional material for the present study was primarily obtained from the R.V. Chulaphorn of the Deep Sea Fishery Technology Research and Development Institute operating off the Andaman Sea coast of Thailand, and from fishing boats, fish markets and fish landings along the Andaman Sea coast of Thailand and the Gulf of Thailand. Taxonomic studies are based on Nateewathana (1997b). The specimen designated as the neotype and five lots of specimens were selected from the samples in the Reference Collection of Phuket Marine Biological Center. The neotype was, like the holotype, collected from small scale fishing areas in Trang Province, at a depth of less than 40 m off the Andaman Sea Coast of Thailand. Measurements follow those of Nateewathana (1997b). Definitions of measurements and indices are summarized in Table 1 and in Figure 1. The difference of the means of all the measurements and indices is compared with those listed by Nateewathana (1997b). Differences were considered significant when Specimens of the species, the newly designated neotype, and selected materials are deposited in the Reference Collection of Phuket Marine Biological Center, Thailand (PMBC), National Science Museum Tokyo, Japan (NSMT), U.S. National Museum, Washington, D.C., USA (USNM), and the Zoological Museum, SNM, University of Copenhagen, Denmark (ZMUC). 17 New records of the bobtail squid Table 1. Definition of counts, measurements and indices (modified after Nateewathana, 1997b). ML Mantle Length MWI Mantle Width Index FLI FWI Fin Length Index Fin Width Index FBI HLI Fin Base Index Head Length Index HWI Head Width Index ALI Arm Length Index ASI Arm Sucker Index TtLI Tentacle Length Index ClLI HcLI Club Length Index Hectocotylized Length Index EDI Eye Diameter Index LnDI Lens Diameter Index dorsal mantle length measured from the anterior-most point of mantle to posterior tip, along dorsal mid-line. greatest straight-line (dorsal) width of mantle as a percentage of mantle length. greatest length of fins as a percentage of mantle length. greatest width (dorsally) across both fins as a percentage of mantle length. length of fin base as a percentage of mantle length. dorsal length of head measured from point of fusion of dorsal arms to anterior tip of nuchal locking cartilage as a percentage of mantle length. width of head across midpoint of eyes as a percentage of mantle length. length of arm measured from first basal (proximal-most) sucker to tip of arm as a percentage of mantle length. diameter of largest normal arm sucker on each designated arm as a percentage of mantle length. total length of tentacular stalk and club as a percentage of mantle length. length of club as a percentage of mantle length. length of modified portion of arm measured from proximal-most modified sucker to tip of arm as a percentage of total length of hectocotylized arm (left arm I). diameter of eye across bulbus as a percentage of mantle length. diameter of eyelens as a percentage of mantle length. RESULTS A total of 265 specimens, 118 males and 147 females, were registered in the Reference Collection (see Table 2). Nearly all of the specimens were collection from Thai waters, the Andaman Sea and the Gulf of Thailand, except for PMBC 11781 and PMBC 11775 which were collected from the Indian Ocean, west of the Andaman Islands and Myanmar waters, respectively. The latter specimens are the first records of E. hyllebergi from the Indian Ocean, west of the Andaman Islands and Myanmar waters, respectively. The neotype was selected from PMBC 11742 and registered as PMBC 21339 (Fig. 2). Three lots of NSMT-Mo 76424, USNM 1150474, and ZMUC CEP-25 have been selected from PMBC 11742, PMBC 11748 and PMBC 11747, respectively. Two females of PMBC 11734 have been selected and registered as NSMT-Mo 76425 and ZMUC CEP-51. The male neotype, PMBC accession number 21339, is deposited in the Reference Collection of Phuket Marine Biological Center, as is the remaining female paratype, PMBC 11731. Selected measurements and indices of the type materials and selected specimens of E. hyllebergi are presented in Table 3. Mean of indices (in percent), standard deviation of five males and four females are presented in Table 4. The t-test comparing the means of all the measurements and indices of the present material and those given by Nateewathana (1997b) are presented in Table 5. 18 Phuket mar. biol. Cent. Res. Bull. Figure 1. Diagrammatic illustration of measurements in sepiolids (modified after Nateewathana, 1997b). AL = arm length, C1L = club length, ED = eye diameter, FB = fin base, FL = fin length, FW = fin width, HL = head length, HW = head width, LD = lens diameter, ML = mantle length, MW = mantle width, TtL = tentacle length. Euprymna hyllebergi Nateewathana, 1997 (Figs. 2A–B, 3A–B, 4A–C, Table 2–5) Euprymna hyllebergi – Nateewathana, 1997b: 466– 474, figs. 2–5, Table 2. Neotype. PMBC 21339, 1 male, Kantang fish landing, Trang province, 14 Feb 1996 (Figs. 2– 3). Paratype. PMBC 11731, 1 female, Tha Lane, Phangnga Bay, push net, 23 July 1989 Selected specimens. NSMT-Mo 76424, 1 male, Kantang fish landing, Trang Province, 14 Feb 1996; NSMT-Mo 76425, 1 female, Chong Pra, Phangnga Bay, push net, 18 June 1988; USNM 1150474, 1 male and 1 female, Phuket fish market, Phuket Province, 23 Aug 1995; ZMUC CEP-25, 2 males, Krabi fish market, Krabi province, 17 Dec 1994; ZMUC CEP-51, 1 female, Chong Pra, Phangnga Bay, push net, 18 June 1988. Material examined. 265 specimens have been examined and registered (see Table 2). Description. Colour in alcohol yellowish; numerous large and conspicuous purplish brown chromatophores cover mantle, head and arms, more dense in middle part of dorsal and ventral sides. Mantle dome-shaped, thick, broad 19 New records of the bobtail squid anteriorly and rounded posteriorly, slightly longer than wide (Figs. 2–3, 4A–B); antero-dorsal margin fused with head (Figs. 2, 4A). Large portion of the tubular funnel visible, median ventral margin slightly concave (Figs. 2B, 4B–C). Fins rounded, almost circular in outline, separate and attached at middle of mantle; anterior margin to posterior margin nearly straight. Head broad, dorsoventrally flattened and compact, width slightly less than mantle width; eyes large, oval with almost circular pupil. Funnel long, tubular, free for most of its length, reaching to gap between ventral arms (Fig. 4B–C). Funnel valve triangular to rounded flaplike. Funnel organs with a roughly triangular dorsal pad and two oval ventral pads. Anterior end of dorsal pad with a low crest projected in midline. Funnel locking cartilage ovate, rounded at both ends with deep groove (Fig. 4C); mantle cartilage straight, ridge-like, two times longer than funnel cartilage. Arms long, subequal, rounded in cross section, tapering distally. Web C (between arms II and III) developed and D distinct (between arms III and IV). Web D extends up about one-fourth of arm length. Suckers almost equal-sized, subglobular, with a small, smooth aperture; pedicel thin. In females, arm suckers quadriserial but biserial at base and distally. In male, arm suckers globular A with smooth horny ring, rounded aperture on normal sucker, but more ovate on enlarged one. Left arm I of male hectocotylized, stout, shorter than other arms and curved outwards (Nateewathana, 1997b: figs. 3a, 4a–b). The size of arm suckers on different arms varied. Tentacles round, flattened on oral surface, edges of flat area angled, dorsal margin elevated into a low keel towards club. Club short, curled, and rounded with numerous rows of very small suckers. Gladius absent. Buccal membrane smooth without suckers. Type locality. Small scale fishing areas in Trang Province, in the depth less than 40 m, the Andaman Sea coast of Thailand. Distribution. Indian Ocean (Andaman Sea and Myanmar waters with range extension to the west of the Andaman Islands) and Gulf of Thailand. Remarks. The characters of all specimens match well with those of E. hyllebergi, as described and figured by Nateewathana (1997b). Description of beak, radula, ink sac, and spermatophores of the species are not repeated herein. The right tentacle in NSMT-Mo 76424 and the left tentacle in ZMUC CEP-51 are missing. B Figure 2: Euprymna hyllebergi Nateewathana, 1997 (PMBC 21339). Neotype, male: (A) dorsal and (B) ventral. 20 Phuket mar. biol. Cent. Res. Bull. A B Figure 3: Euprymna hyllebergi Nateewathana, 1997 (PMBC 11731). Paratype, female: (A) dorsal and (B) ventral. A B C Figure 4. Euprymna hyllebergi Nateewathana, 1997 (PMBC 21339). Neotype, male: (A) dorsal view; (B) ventral view; (C) funnel and funnel locking-cartilages. Scales = 2 cm (A–B) and 0.5 cm (C). Table 2. Details of specimens of Euprymna hyllebergi remaining in the Reference Collection of Phuket Marine Biological Center. No. PMBC No. 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 11727 11728 11729 11730 11731 11732 11733 11734 11735 11736 11737 11738 11739 11740 11741 11742 11744 11745 11746 11747 11748 11749 11750 11751 11752 11753 11755 11756 11757 11758 11759 11760 11761 5 1 HOLOTYPE specimen lost 6 11 PARATYPE 1 1 6 3 2 3 PARATYPE specimen lost 5 PARATYPE specimens lost 9 9 32 12 1 1 5 12 2 1 1 4 3 13 4 1 1 1 3 1 3 Female Locality Province Identifier 2 – Phuket A. Nateewathana Krabi Trang Trang Ranong Phangnga Trang Krabi Phangnga Phangnga Phangnga Trang Andaman Sea Krabi Phangnga Phangnga Trang Krabi Krabi Ranong Krabi Phuket Phuket Phangnga Phangnga Phuket Krabi Satul Phangnga Ranong Andaman Sea Phangnga Phuket Krabi A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana 1 2 5 3 1 1 4 6 1 1 3 2 1 3 5 2 3 15 6 4 5 1 2 3 6 1 1 2 7 6 17 6 1 1 1 7 1 1 1 2 7 3 1 1 1 1 Krabi fish market Kantang fish landing Kantang fish landing Kapur Tha Lane, Phangnga Bay Rok Island Krabi fish market Chong Pra, Phangnga Bay Raya Ring Island Tha Lane, Phangnga Bay Kantang fish landing 8°02´N, 98°30´E Southwest of Lanta Island Taeng Cape, Phangnga Bay Phangnga Bay Kantang fish landing Krabi fish market Krabi fish market – Krabi fish market Phuket fish market Tukkae Cape Raya Ring Island, Phangnga Bay Mak Island, Phangnga Bay Phuket fish market Southwest of Lanta Island Tamalang fish landing – Jam Island 7°48´N, 98°30´E Nakae Island Sapum Bay Dam Hok Island Remarks original recorded as Euprymna stenodactyla (Grant, 1833) by C.F.E. Roper in 1979 21 1607 No. of spec. Male New records of the bobtail squid 1 Type No. PMBC No. 22 Table 2. (continued) No. of spec. NEOTYPE 1 2 2 1 1 1 2 1 3 1 1 1 18 1 1 1 3 16 26 1 10 1 1 Male Female 58 23937 4 3 1 – – A. Nateewathana 59 25004 5 3 2 – – A. Nateewathana 2 5 2 10 17 1 6 4 1 1 Trang Phangnga Krabi – Krabi Andaman Sea Andaman Sea Phuket Krabi Ranong Trang Myanmar waters Pattani Trang Samut Prakan Indian Ocean Andaman Sea Trang Trang Andaman Sea Andaman Sea Andaman Sea Trang Identifier 11762 11763 11764 11765 11766 11767 11768 11770 11772 11773 11774 11775 11776 11777 11780 11781 11782 11785 11787 14420 19903 19904 21339 1 1 2 1 1 1 1 1 13 1 1 1 1 6 9 Trang fish market off Yao Yai Island East of Lanta Island – – 6°58´N, 98°48´E 6°57´N, 98°47´E Rang Yai Island – – Kantang fish landing 13°27´N, 97°00´E Pattani fish landing Kantang fish landing Samut Prakan fish landing 7°17´N, 90°00´E 7°04´N, 98°53´E Kantang fish landing Kantang fish landing 9°34´N, 97°49´E 7°51´N, 98°38´E 9°00´N, 97°54´E Kantang fish landing Province 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 1 1 1 2 1 Locality A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana A. Nateewathana T. Kubodera T. Kubodera A. Nateewathana Remarks specimen was selected from PMBC 11742 specimens found in the bottle of PMBC 11737, but not identical to original paratype specimens found in the bottle of PMBC 11739, but not identical to original paratype specimens Remarks: the number of specimens in some PMBC accession numbers is changed, compared to that published in Appendix 1 of Nateewathana (1997b), and some PMBC accession numbers were not included in that appendix. Phuket mar. biol. Cent. Res. Bull. Type Table 3. Measurements (mm) and indices (%) of neotype, paratype, and selected specimens of Euprymna hyllebergi from the Andaman Sea, Thailand. Paratype Registration No. PMBC 21339 PMBC 11731 NSMT-Mo 76424 NSMT-Mo 76425 Index\Sex Male Female Male Female Male ML (mm) MWI FLI FWI FBI HLI HWI ALII ALIII ALIIII ALIVI ASII ASIII ASIIII ASIVI TtLI CILI HcLI EDI LnDI 30.98 84.83 44.09 129.50 26.44 54.71 63.52 106.751 143.54 120.27 116.17 3.23 4.03 5.23 6.17 327.73 42.32 65.73 22.60 12.33 33.63 77.19 48.41 119.92 28.49 52.81 61.11 97.71 114.93 98.331 110.70 2.97 4.16 4.46 4.46 234.672 46.65 59.99 19.80 13.56 35.22 74.82 51.02 135.15 33.19 50.03 59.71 59.00 74.761 69.96 67.181 2.13 2.30 2.98 2.73 84.44 31.63 29.02 90.45 54.51 146.76 35.70 53.62 64.13 91.94 124.911 104.311 105.10 3.96 3.79 5.34 5.00 290.18 31.741 62.93 27.08 13.20 22.27 91.02 58.96 152.09 46.74 46.43 73.15 69.33 86.30 77.191 73.37 2.92 3.37 2.92 2.25 119.98 34.98 25.82 15.67 Selected specimens 24.11 12.12 USNM 1150474 Female 26.79 88.28 50.09 159.69 39.94 64.05 67.19 91.75 99.22 93.77 96.90 2.24 2.61 2.43 2.43 246.62 62.64 24.15 15.57 ZMUC CEP 25 ZMUC CEP 51 Male Female 27.21 83.83 52.11 132.82 32.97 53.55 68.83 88.06 110.66 101.14 98.46 2.94 2.94 4.41 4.59 285.34 42.08 58.01 30.10 15.40 28.65 81.85 48.94 134.90 37.63 58.71 64.96 99.34 124.15 108.34 101.22 4.01 3.66 5.34 4.54 229.60 41.64 60.94 26.35 14.52 28.42 76.18 57.74 135.50 34.24 57.35 57.74 84.87 102.39 91.87 88.53 2.53 2.43 3.20 2.81 169.11 39.94 0.00 25.97 13.62 New records of the bobtail squid Neotype Remarks: 1 = tip of the right arm broken; 2 = left tentacle due to the right one lost. The values in bold text are out of the range of the measurements and indices reported by Nateewathana (1997b). 23 24 Phuket mar. biol. Cent. Res. Bull. Table 4. Mean (X) of indices (%) and standard deviation of type materials and selected specimens of Euprymna hyllebergi from the Andaman Sea, Thailand. Male Female Index ML (mm) MWI FLI FWI FBI HLI HWI ALII ALIII ALIIII ALIVI ASII ASIII ASIIII ASIVI TtLI CILI HcLI EDI LnDI n Mean SD 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 29.90 83.63 49.61 132.78 32.24 54.68 64.51 96.76 123.64 106.48 106.33 3.42 3.72 4.96 4.95 273.50 40.89 61.52 25.19 13.80 2.48 4.81 3.96 9.70 4.72 2.35 2.81 7.18 12.67 8.56 7.17 0.53 0.48 0.48 0.71 41.21 5.50 2.95 4.02 1.19 Range n Mean SD 27.21–33.63 77.19–90.45 44.09–54.51 119.92–146.76 26.44–37.63 52.81–58.71 61.11–68.83 88.06–106.75 110.66–143.54 98.33–120.27 98.46–116.17 2.94–4.01 2.94–4.16 4.41–5.34 4.46–6.17 229.60–327.73 31.74–46.65 58.01–65.73 19.98–30.10 12.33–15.40 4 4 4 4 4 4 4 4 4 4 4 4 4 4 4 4 4 4 4 4 28.18 82.57 54.45 145.61 38.53 54.47 64.45 76.24 90.67 83.20 81.50 2.46 2.68 2.88 2.55 155.04 42.30 5.37 8.27 4.54 12.27 6.23 7.84 7.09 14.83 12.68 11.52 13.64 0.35 0.48 0.33 0.26 70.23 13.98 25.01 14.24 1.02 1.70 Range 22.27–35.22 74.82–91.02 50.09–58.96 135.15–159.69 33.19–46.74 46.43–64.05 57.74–73.15 59.00–91.75 74.76–102.39 69.96–93.77 67.18–96.90 2.13–2.92 2.30–3.37 2.43–3.20 2.25–2.81 84.44–246.62 31.63–62.64 24.11–25.97 12.12–15.67 Table 5. t-test of the mean of all measurements and indices by Nateewathana (1997b) and this study. Nateewathana (1997b) Male present study Index n Mean SD n Mean ML (mm) MWI FLI FWI FBI HLI HWI ALII ALIII ALIIII ALIVI ASII ASIII ASIIII ASIVI TtLI CILI HcLI EDI LnDI 20 20 20 20 20 20 20 20 20 20 20 20 20 20 20 20 20 20 20 20 27.20 81.20 49.10 131.10 31.50 52.80 65.20 103.70 141.00 118.20 118.60 3.50 4.40 5.10 5.30 282.80 48.40 63.40 23.30 14.40 4.20 11.30 7.10 13.80 4.00 8.50 7.10 20.00 15.70 14.10 18.00 0.50 0.60 0.70 0.60 47.10 7.10 8.30 5.60 2.80 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 29.90 83.63 49.61 132.78 32.24 54.68 64.51 96.76 123.64 106.48 106.33 3.42 3.72 4.96 4.95 273.50 40.89 61.52 25.19 13.80 SD T-test 2.48 4.81 3.96 9.70 4.72 2.35 2.81 7.18 12.67 8.56 7.17 0.53 0.48 0.48 0.71 41.21 5.50 2.95 4.02 1.19 No diff No diff No diff No diff No diff No diff No diff No diff Sig diff No diff No diff No diff Sig diff No diff No diff No diff Sig diff No diff No diff No diff 25 New records of the bobtail squid Table 5. (continued) Nateewathana (1997b) Female Index n ML (mm) MWI FLI FWI FBI HLI HWI ALII ALIII ALIIII ALIVI ASII ASIII ASIIII ASIVI TtLI CILI EDI LnDI 20 20 20 20 20 20 20 20 20 20 20 20 20 20 20 19 19 20 20 Mean 24.90 80.70 52.50 142.40 35.80 55.30 67.70 85.40 110.60 93.20 89.70 2.70 2.80 3.00 2.80 266.60 53.30 23.50 14.90 present study SD n Mean SD T-test 5.50 8.90 7.80 12.80 5.00 6.20 7.90 11.40 12.30 8.80 9.20 0.40 0.50 0.40 0.40 60.70 10.90 3.80 2.10 4 4 4 4 4 4 4 4 4 4 4 4 4 4 4 4 4 4 4 28.18 82.57 54.45 145.61 38.53 54.47 64.45 76.24 90.67 83.20 81.50 2.46 2.68 2.88 2.55 155.04 42.30 25.01 14.24 5.37 8.27 4.54 12.27 6.23 7.84 7.09 14.83 12.68 11.52 13.64 0.35 0.48 0.33 0.26 70.23 13.98 1.02 1.70 No diff No diff No diff No diff No diff No diff No diff No diff Sig diff No diff No diff No diff No diff No diff No diff Sig diff No diff No diff No diff DISCUSSION The Euprymna hyllebergi arm length formula is II > III > IV > I, but occasionally II > III = IV > I (Nateewathana, 1997b). The arm length formulas of the present material of males and females (Table 4, II > III > IV > I), matches those of Nateewathana (1997b) as well as those in the neotype and paratype (Table 3). Differences are, however, found in some of the selected specimens; four specimens show II > III > IV > I but three other specimens show II > IV > III > I (Table 3). These differences occur because the tips of the right arms were broken. All measurements and indices of the neotype are in the ranges reported by Nateewathana (1997b). Some measurements and indices in the paratype and selected specimens are, however, out of these ranges, i.e., Mantle Length (ML), Arm Length Index (ALI), Arm Sucker Index (ASI), Tentacle Length Index (TtLI), and Club Length Index (ClLI) (Tables 3–4). Furthermore, the t-test of the mean of all measurements and indices by Nateewathana (1997b) and in this study were considered significant for ALII in both male and female, for ASIII and ClLI in male only, and for TtLI in female (Table 5). These differences may occur because the suckers on arms easily fall off and are lost in preserved material and the tip of the first right arm in male specimen was broken. Therefore some errors occurred in estimation of the length of the first right arm, which was measured from the first basal (proxima-most) sucker to tip of arm as a percentage of mantle length. Furthermore, according to C.C. Lu (pers. comm., in Nateewathana, 1997b), the length of the tentacles in cephalopods may not be a usable taxonomic character since it varies greatly depending on how the specimen is fixed and preserved. Live fixed animals have short tentacles because of contraction and, on the other hand, specimens thawed from frozen condition have long tentacles. In order for this character to be of some 26 Phuket mar. biol. Cent. Res. Bull. use, specimens should be fixed under identical conditions. Specimens from the same batch may vary greatly (C.C. Lu, pers. comm., in Nateewathana, 1977b). The mantle length, the first arm length index, the tentacle length index, and club length index are the main characters/indices used for distinguishing the species from other members of the genus. Although the tentacle length is particularly important in recognizing this species, it is not necessarily an autopomorphy for the species. It is the combination of the arrangement of suckers on the hectocotylized arm, the size and distribution of enlarged suckers on arms II–IV, and in the length of the tentacles that distinguishes Euprymna hyllebergi. ACKNOWLEDGEMENTS We express our appreciation to Dr. Kongkiat Kittiwattanawong, Phuket Marine Biological Center (PMBC) for his kind assistance in t-test Analysis, and Dr. Janek von Byern, University of Vienna, Austria, for providing valuable comments on the draft of this manuscript. Sincere thanks are given to Ms. Kanjana Bussabong, the Reference Collection, Phuket Marine Biological Center, and Ms. Thitima Boonkaew, training student, PSU, for their kind assistance. Our sincere thanks also go to Mr. Rueangrit Promdam, (former staff of Reference Collection), and Mr. Patirat Singdum, the PMBC artist, for photographing and drawings. REFERENCES Adam, W. 1986. Contribution à la connaissance du genre Euprymna Steenstrup, 1887 (Mollusca: Cephalopoda). Bulletin de l’Institut Royal des Science Naturelles de Belgique, Biologie. 56: 131–136. Aungtonya, C., K. Bussabong, K. Wongkamhaeng, I. Eedkerd, and S. Ubonsuwan. 2006. Type specimens in the Reference Collection of Phuket Marine Biological Center. Reference Collection and Aquarium Contributions no. 4: 45 p. (in Thai). Aungtonya, C., S. Sungjeen, W. Tripanichkul, K. Bussabong, and P. Kongthaworn. 2007. Recent records of the Class Cephalopoda (Phylum Mollusca) in the Reference Collection of Phuket Marine Biological Center. Reference Collection and Aquarium Contributions no. 5: 69 p. (in Thai). Berry, S. S. 1913. Some new Hawaiian cephalopods. Proc. U. S. natn. Mus. 45: 563–566. Chotiyaputta, C. 1993. Cephalopod resources of Thailand. Pp. 71–80. In: Okutani,T., R.K. O’Dor & T. Kubodera (eds.). Recent Advances in Fisheries Biology. Tokai University Press, Tokyo. Chotiyaputta, C., T. Okutani, and S. Chaitiamvong. 1992. Systematic study of cephalopods in Thailand. A Research Report submitted to the National Research Council of Thailand under JSPS-NRCT Cooperation Project. 100 pp. (In Thai with English abstract). Grant, R. E. 1883. On a new species of Sepiola (Sepiola stenodactyla) from the Mauritius, presented by C. Telfair, Esq. Proceedings of the Zoological Society of London. 1: 42–43. Nateewathana, A. 1997a. Systematics of the Cephalopoda of the Andaman Sea, Thailand. Ph.D. Thesis. Institute of Biological Sciences, Faculty of Natural Sciences, University of Aarhus. 343 pp. Nateewathana, A. 1997b. The Sepiolidae (Cephalopoda) of the Andaman Sea, Thailand, with description of Euprymna hyllebergi sp. nov. Phuket Marine Biological Center Special Publication. 17: 465–481. Nateewathana, A., J. Nabhitabhata, and P. Nilaphat. 2001. A new record of bobtail squid, Euprymna hyllebergi Nateewathana, 1997 in the Gulf of Thailand. Phuket Marine Biological Center Special Publication. 25(2): 501–506. Nesis, K.N. 1987. Cephalopods of the world. T.F.H. Publications, Inc. Ltd., New Jersey. 351 pp. 27 New records of the bobtail squid Norman, M.D. & C.C. Lu. 1997. Redescription of the southern dumpling squid Euprymna tasmanica and a revision of the genus Euprymna (Cephalopoda: Sepiolidae). J. Mar. Biol. Ass. U.K. 77: 1109–1137. Pfeffer, G.J. 1884. Die Cephalopoden des Hamburger Naturhistorischen Museums. Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. 8(2): 63–90. Sasaki, M. 1913. Decapod cephalopods found in Japan: Sepiolidae. Zool. Mag., Tokyo. 25: 247–252. Steenstrup, J.J. 1887. Notae teuthologicae, no. 6. Species generic Sepiolae Maris Mediterranei. Oversi. K. Danske Vidensk Selsk Forh. Copenhagen. 1887: 47–66. Supongpan, M. 1995. Cephalopod resources in the Gulf of Thailand. pp. 191–203. In: Nabhitabhata, J. (ed). Biology and Culture of Cephalopods. Rayong Coastal Aquaculture Station, Contribution No. 18. (In Thai). Verrill, A.E. 1881. The cephalopods of the north-eastern coast of America: part 2. The smaller cephalopods, including the squids and the octopi, with other allied forms. Trans. Conn. Acad. Arts. Sci. 5: 259–446. Voss, G.L. 1963. Cephalopods of the Philippines. Smithsonian Institution Bulletin. 234: 1–180. Manuscript received: 17 August 2009 Accepted: 12 June 2011 28 Phuket mar. biol. Cent. Res. Bull. Phuket mar. biol. Cent. Res. Bull. 70: 29–77 (2011) THE FISHES OF SOUTHWESTERN THAILAND, THE ANDAMAN SEA – A REVIEW OF RESEARCH AND A PROVISIONAL CHECKLIST OF SPECIES Ukkrit Satapoomin Phuket Marine Biological Center, P.O. Box 60, Phuket 83000, Thailand Email: ukkrit@yahoo.com ABSTRACT: A brief review of recent ichthyological research and compiled results on the records of fishes from the Andaman Sea coast of Thailand are presented. A total of 1,746 species in 198 families of fishes are currently known from the area. The 10 most speciose families are the Gobiidae (227 species), Labridae (78), Pomacentridae (71), Serranidae (61), Apogonidae (60), Blenniidae (52), Carangidae (52), Scorpaenidae (49), Lutjanidae (39) and Chaetodontidae (37), and these together account for 42% of the total fish fauna. The ichthyofauna is dominated by reef-associated fishes (983 species) and pelagic/benthic fishes inhabiting offshore habitats (971 species). A zoogeographic analysis reveals a peculiarity of the fauna of this marginal-sea region, viz., being one of the areas of sympatry of Indian and Pacific Ocean fishes, as well as harbouring regional endemics. The PMBC Reference Collection currently contains examples of only about 63% of the fishes known from the area, indicating a need for further local sampling campaigns and/or international collaborative exploratory research programs in the Andaman Sea to build up the Reference Collection. Keywords: Thailand, Andaman Sea, fishes, checklist, diversity. INTRODUCTION The Andaman Sea is an approximately 1,200 km long x 650 km wide body of water to the southeast of the Bay of Bengal. It is bounded by the Gulf of Mataban in the north, the Malay Peninsula in the east, the Andaman and Nicobar Islands in the west and the northern tip of Sumatra in the south. At its southeastern corner, the Andaman Sea narrows to form the Strait of Malacca, which separates the Malay Peninsula from the island of Sumatra. Along an approximate 1,000 km length of the Andaman Sea coast of Thailand, a variety of marine and coastal habitats are present, viz., mangroves, mud-flats, seagrass beds, rocky shores, intertidal sand-flats, coral reefs, open pelagic zones, and complex seabed systems on the continental shelf. The integrity of different peculiar environmental settings and/or characteristics among those habitat types provides suitable refuges for a vast variety of aquatic biota, including fishes. Although, biogeographically, it belongs to the eastern Indian Ocean, it is strongly influenced by the western Pacific Ocean biota via the northwesterly through-flow in the Strait of Malacca. It is thus an area of distributional confluence of the biotas of both the Indian Ocean and the Pacific Ocean (Satapoomin, 2002a, 2007; Kimura and Satapoomin, 2009). Being a marginal sea separated from the main Indian Ocean by the Andaman-Nicobar island arcs, it has been subjected to a profound effect of sea-level lowering, especially during major glaciations when it became a separated, enclosed basin. Such isolation processes are believed to have promoted speciation and/or endemism of the fish fauna in the Andaman Sea (e.g., McManus, 1985; Randall, 1998; Randall and Satapoomin, 1999). The area is thus characterized by its high diversity and the unique faunal composition of the fishes. However, knowledge and information on the overall diversity of marine fishes in the area is still limited, because very few attempts had been made to compile a list of all known fish species from the area. The purposes of the present paper are to provide a brief review of recent ichthyological research, and to establish a comprehensive checklist of all known marine and brackish-water 30 Phuket mar. biol. Cent. Res. Bull. fishes from the Andaman Sea coast of Thailand. The checklist was compiled from various sources of information, including published literature, unpublished data made available under several sampling programs/projects, and the fish collection data repository of the Phuket Marine Biological Center (PMBC). It should be noted that the overall list of species presented herein is only provisional as a number of species records and identifications in previously published literature (particularly those of inventory, non-taxonomic works), cannot be verified. Nonetheless, this provisional checklist will serve as a significant baseline for further establishment of a conventional, annotated checklist or taxonomic review of the Andaman Sea fishes. METHODS The compilation of the species list was based largely on accumulating information from the earlier literature, including inventory works and/ or faunistic surveys, field guides, and taxonomic works. Some unpublished data, including recent unpublished survey results made by the author, collection data of the Royal Ontario Museum’s project on gobioid fishes of southwestern Thailand in 1993 (under auspices of Dr. Richard Winterbottom), and the PMBC fish collection data repository were also incorporated into the current checklist. Selected taxonomic papers pertaining to records of fishes from the Andaman Sea coast of Thailand used to compile the current checklist are listed in Appendix B. The systematic arrangement of families follows Nelson (2006). Species in each family are arranged alphabetically. The scientific names of fishes used in the cited literature have been updated where possible, following recent usage, and hence junior synonyms were omitted. Since the compilation was also based, in part, on literature that provided only a list of names, the author has only been able to check or correct misidentifications to a limited extent. In this regard, only correct names were retained, synonyms were allocated according to the latest usage, while doubtful species records were excluded. In addition to scientific names and classification, the checklist provides basic information on habitats and zoogeographic distribution of each species. The availability of voucher specimens lodged at the Reference Collection of the PMBC was also emphasized in order to draw the attention of ichthyologists who may wish to examine these specimens. The designation of habitat type(s) for each species was based largely on either quoted information from the compiled literature or from the author’s unpublished survey results and observations. Four principal habitat categories adopted herein included mangrove (Mg), coral reef (Cr), seagrass bed (Sg), and other offshore habitats (OShr). These categories can be considered as good representations of the major marine and coastal features along the southwestern Thailand coastline. Each of these habitat types, however, was more or less loosely defined in order to accommodate a range of associated minor biotopes of each habitat. The offshore category, in particular, was the most inclusive term as it encompassed a range of benthic- and pelagic-related habitats. This designation includes both near-shore and faroffshore fishes considered as either pelagic (i.e., coastal pelagic, offshore pelagic, epipelagic, and mesopelagic) or demersal (i.e., coastal demersal, benthopelagic, and deep demersal) species. The mangrove habitat was considered to include certain inter-connected zones like mud-flats and the upper reaches of the estuarine areas. Coral reef was defined to include several nearby and/or associated areas, namely intertidal rocky shore/rock platform, intertidal sand-flat, and subtidal bare-seafloor in front of the reef (so called as forereef). Most of the well defined zoogeographic distribution categories (see, e.g., Myers, 1991) were generally adopted or redefined with additional explanation, while some new categories were added with their definitions. The definitions for those distribution categories and their acronyms are as follows: Indo-Pacific (IP), widespread across the Indian Ocean and western portion of the Pacific with an eastern limit east of non-marginal areas of the Pacific Plate; Indo-west Pacific (IWP), a subset of the Indo-Pacific with northeastern limit on the Pacific Plate, marginally in the Carolines but not reaching the Marshalls; West Pacific (WP), limited to the area bound by the Andaman Island and Cocos-Keeling Island in the west to the margin of 31 The fishes of southwestern Thailand the Pacific Plate in the east, north to southern Japan, south to western and eastern Australia; West central Pacific (WCP), west Pacific and Pacific Plate distributions and may include the East Pacific; Indo-Australian Archipelago (IA), a subset of the West Pacific, north to southern Taiwan, east to the Solomons, south to the Great Barrier Reef; Indo-Malayan Archipelago (IM), a subset of IndoAustralian which is limited to the areas within bounds of Malay Peninsula, Sumatra, Java, New Guinea and Philippines; Indian Ocean (IO), widespread throughout the Indian Ocean; Northern Indian Ocean (NIO), the extended range of eastern Indian Ocean to encompass the northwestern limits which include the west coast of India, Pakistan, Oman and Persian Gulf; Eastern Indian Ocean (EIO), the eastern section of the Indian Ocean westwards to Laccadive, Maldive and Chagos archipelagoes; the eastern extremity of the Indian Ocean (EEIO), a narrow stripe or transition area along the joint margin of Indian-Australian and Asian lithospheric plates, including the Andaman Sea, Sumatra, Java, Bali, Flores, CocosKeeling/Christmas Islands and northwestern Australian Shelf; Andaman Sea province (AS), the area assigned to accommodate species having restricted distribution only to the Andaman Sea or ranging southwards only to western Sumatra; Circumtropical (CIR), tropical seas worldwide; and Cosmopolitan (COS), tropical and temperate seas worldwide. The boundary of some distribution categories is presented (Fig. 1) for clarification of the definitions of those areas. RESULTS AND DISCUSSION 1. Brief review of ichthyological research The fish fauna of the Andaman Sea coast of Thailand was poorly documented until relatively recent times. There were several well-known western ichthyologists/naturalists (e.g., Cuvier, Valenciennes, Bleeker, Günther, Fowler, Regan, Boulenger, etc.) who reported, listed or described Thai fishes between 1800 and the 1950s, but their works were restricted only to either the Gulf of Thailand (marine) or inland (freshwater) fishes (see Monkolprasit et al. (1997) for detailed historical review). The first marine fish of the Andaman Sea coast of Thailand described by a westerner was a reef-associated goby, Pipidonia quinquecincta Smith, 1931 [valid as Gobiopsis quinquecincta (Smith)]. During his term of service (1923–1935) as the Adviser in Fisheries to the Department of Fisheries of Thailand (formerly Siamese Bureau of Fisheries), Smith published a number of articles pertaining to freshwater fishes and fishes of the Gulf of Thailand. Later, in 1945, he published a monograph entitled “The Freshwater Fishes of Siam, or Thailand” (Smith, 1954). Suvatti (1950) was the first Thai ichthyologist to publish a list of Thai fishes. Several subsequent studies on marine fishes in Thai waters by the Thais were confined to groups of particular interest, for instance Pleuronectiformes (by Punpoka, 1964), Lutjanidae (Banasopit, 1968), Serranidae (Banasopit, 1969; Monkolprasit, 1983), Chaetodontidae (Banasopit, 1971), Leiognathidae (Monkolprasit, 1973), Elasmobranchii (Monkolprasit, 1984), and Labridae (Monkolprasit, 1989). The foremost comprehensive compiled checklist of the fishes of Thailand, which included both freshwater and marine fishes, was published by Monkolprasit et al. in 1997. It listed 171 fish families with a total of 1,741 species, about 30% (516 species) of which can be considered as freshwater fishes. The records of marine fishes in the checklist were compiled from the literature published before 1990, except for two papers (one on mangrove fishes by Monkolprasit (1994) and one on ammodytid fishes by Ida et al. (1994)). That checklist recorded 710 species representing 129 families of fishes from the Andaman Sea. A few collections of reef fishes from the Similan Islands were made during the R/V Te Vega Cruise 2, eastern Indian Ocean cruise (from Singapore to Colombo, Ceylon), in late 1963 (http:/ /tevega.stanford.edu/cruises.htm). These collections were deposited at the U.S. National Museum of Natural History, and some were designated as types of new fish species described subsequently (i.e., Melichthys indicus Randall & Klausewitz, 1973, Gymnothorax pseudoherrei Böhlke, 2000, and Helcogramma lacuna Williams & Howe, 2003). In February 1979, Randall (Bishop Museum, Hawaii) and Allen (Western Australian Museum) made a few collections and photographed reef fishes around Phuket and the Similan Islands (see Allen et al., 2005). Although 32 Phuket mar. biol. Cent. Res. Bull. Figure 1. Approximate range of some zoogeographic distribution categories. A: Indo-Pacific (IP), Indo-west Pacific (IWP), West central Pacific (WCP), and West Pacific (WP); B: Indo-Australian archipelago (IA) and Indo-Malayan Archipelago (IM); C: Northern Indian Ocean (NIO), Eastern Indian Ocean (EIO), and Eastern extremity of Indian Ocean (EEIO). no comprehensive list of fishes resulted from that effort, the collected materials had been used in describing several new species (i.e., Leptojulis chrysotaenia Randall & Ferraris, 1981, Chromis flavipectoralis Randall, 1988, Pomacentrus adelus Allen, 1991, P. azuremaculatus Allen, 1991, P. polyspinus Allen, 1991, P. similis Allen, 1991, Neopomacentrus sororius Randall & Allen, 2005, Plesiops auritus Mooi, 1995, Plectorhinchus macrospilus Satapoomin & Randall, 2000, Ctenochaetus truncatus Randall & Clements, 2001, Scorpaenopsis possi Randall & Eschmeyer, 2001, Helcogramma lacuna Williams & Howe, 2003, and Opistognathus cyanospilotus Smith-Vaniz, 2009). The most important collections of reef fishes for the area were made in November 1993 under the collaborative project between the Royal Ontario Museum (ROM), Canada and the Phuket Marine Biological Center (PMBC). From well over 35 sampling stations using the ichthyocide rotenone 33 The fishes of southwestern Thailand around Phuket and adjacent islands, nearly eighteen thousand specimens representing about 570 species in 75 families of fishes were collected and documented. It was also the first time that significant information on small cryptobenthic reef fishes for the area was obtained. The collected materials have been used in the descriptions of 13 new species to date (including 7 species in Gobiidae and 1 in each of the following families: Apogonidae, Serranidae, Pomacentridae, Opistognathidae, Tripterygiidae, and Soleidae), with several other descriptions underway. Satapoomin (1993) established a first comprehensive checklist of coral reef fishes of the west coast of Thailand by compiling information from both available literature (dated back to 1950) and the author’s survey results carried out during 1990–1992. The list revealed a total of 597 species belonging to 66 families of reef fishes in the area. Allen et al. (2005) provided an extensive list, consisting of 764 species in 70 families, of coral reef fishes being recorded and/or known for the area. The most recent list of coral reef fishes, with 888 species in 85 families, was published by Satapoomin (2007). There are a limited number of studies on ichthyofauna of either mangroves or seagrass beds in the Andaman Sea coast of Thailand, most of which were confined to particular sampling sites (e.g., Boonruang et al., 1994; Monkolprasit, 1994; Janekitkarn and Monkolprasit, 1994). Satapoomin and Poovachiranon (1997) provided the first comprehensive list of the fish fauna from several major mangrove and seagrass bed sites along the coast of southwestern Thailand. That study revealed a total of 280 species representing 75 families of fishes, which included 232 species (69 families) and 149 species (51 families) found in mangroves and seagrass beds, respectively. In 2002, a study by Fish Team of the Trang Project (2002) provided a list, together with illustrations, of 134 species in 49 families of fishes inhabiting a mangrove estuary at Sikao, southwestern Thailand. Under the JSPS (Japan Society for the Promotion of Science) Multilateral Project on Coastal Marine Science, Matsuura and Kimura (2005) published a guide book with taxonomic accounts and illustrations of 128 species in 46 families of fishes inhabiting mangroves, seagrass beds and surrounding area of Libong Island, Trang Province. The most recent list of the fish fauna in the seagrass beds of the Andaman Sea coast of Thailand, with 230 species in 62 families, was published by Satapoomin (2006). The first publication on demersal and/or pelagic fishes of the Andaman Sea coast of Thailand was published by Lohakarn (1968). This included taxonomic accounts of 109 species in 57 families. Fishes collected by bottom trawls during the 5th Thai-Danish Expedition to the Andaman Sea in 1966 (the first Thai-Danish marine expedition) were reported by Boonyanate and Hylleberg (1993) with a total records of 80 species in 41 families. Other early studies on demersal fishes in the area were mostly focused on fisheries aspects and, in most cases, faunal composition was reported mainly as higher taxonomic ranks rather than individual species (e.g., Pokapunt and Sinoda, 1978; Boonragsa, 1988; Chantawong, 1996). Some studies on experimental operations of certain types of fishing gears reported not more than 50 fish species each (e.g., Wongratana, 1982b, 1983; Ruengpan, 1988). A more comprehensive list of demersal fishes collected by otterboard trawl nets and deep sea shrimp traps along the Andaman Sea coast of Thailand at depths of about 30–420 m was published by Wongratana (1982a), with 256 species in 91 families. Also, Pokapunt and Tantivala (1987) reported 182 species of demersal fishes (in 66 families) in the area from surveys using otterboard trawls at water depths less than 90 m. Deep-sea fishes of the Andaman Sea were poorly known. The results of surveys during 1975 and 1976 on fishes inhabiting the continental shelf within the depth range of 200– 550 m was reported by Pokapunt et al. (1983), with taxonomic accounts of 31 species in 20 families. Under the Scientific Cooperation Programme between Denmark and Thailand during 1996–2000, the Biodiversity of the Andaman Sea Shelf (BIOSHELF) Project was undertaken by the staff of the PMBC together with their Danish counterparts. The benthic fauna, including fishes, was sampled at depths of 40–900 m. Preliminary results revealed a total of 55 families in 20 orders of fishes, most of which were deep-water fishes that have not yet been identified (Aungtonya et 34 Phuket mar. biol. Cent. Res. Bull. al., 2000). Under the JSPS project, a comprehensive guide book entitled “Fishes of Andaman Sea, West Coast of Southern Thailand” covered 778 species in 106 families of fishes (Kimura et al., 2009). The Reference Collection of the PMBC is one of the major sources of information on marine biodiversity of the Andaman Sea coast of Thailand. The collection of fishes, in particular, was largely established as a result of the FAO/DANIDA Seminar on Fish Taxonomy in Southeast Asia held at the PMBC in 1972. Apart from the Andaman Sea fishes, a number of specimens obtained from the Gulf of Thailand, as well as from neighboring countries (i.e., India, Malaysia, Singapore, Indonesia, Philippines, and Hong Kong) were brought in during the seminar and subsequently deposited in the PMBC Reference Collection. Some deposited material of fishes from Australia was donated by the Australian Museum under a past exchange programme. One lot of fishes collected from Myanmar during the first joint Myanmar-Thai Fishery Exploratory Survey in 1990 was also deposited in the Reference Collection (Nateewathana et al., 1983). A revised checklist of fishes in the PMBC Reference Collection was published by Nateewathana et al. (1983), comprising a total of 629 species in 119 families. The collection has gradually increased following a number of subsequent researches (e.g., Sirimontraporn et al., 1993; Satapoomin and Poovachiranon, 1997; Satapoomin, 1999, 2002b) and intensive sampling programmes (e.g., a collaborative project between ROM and PMBC in 1993 and the PMBC’s past and ongoing projects, namely Coral Reef Management Project (1995– 1999), Seagrass Resources Management and Dugong Conservation Project (1997–2000), and Assessment of Coastal Marine Resources Project (since 2004)). According to the recent PMBC data repository (http://pmbc.go.th/pmbc-rc/ Taxonomy.php/), the most reliable records of fishes (excluding doubtful records and/or misidentifications) deposited in the Reference Collection total at least 1,200 species belonging to 170 families. The Andaman Sea fishes, in particular, are well represented with a total number of 1,093 species, comprising 1, 39, and 1,053 species in Myxini, Chondrichthyes, and Actinopterygii, respectively. 2. Faunal composition and zoogeography The present checklist contains 1,746 species, representing 696 genera, 198 families, 40 orders, and 3 classes, of marine and coastal fishes from the Andaman Sea coast of Thailand (Appendix A). The figure is relatively high as compared to those reported from several other localities in the Indian Ocean. For example, 533 species were reported from the Cocos (Keeling) Islands (Allen and Smith-Vaniz, 1994), 603 from the Laccadive Archipelago (Jones and Kumaran, 1980), 622 from Christmas Island (Hobbs et al., 2010), 784 from the Chagos Archipelago (Winterbottom and Anderson, 1999), 880 from Seychelles and adjacent areas (Smith and Smith, 1969), 885 from the Reunion Island (Letourneur et al., 2004), 930 species from Oman (Randall, 1995), 1,078 from the Red Sea (Golani and Bogorodsky, 2010), 1,090 from the Maldives (Anderson et al., 1998), and 1,800 (as an estimate) from southern Africa (Heemstra and Heemstra, 2004). There may be the real differences in fish diversity among those localities, but it should be noted that the number of species recorded probably reflect biases or differences in any or all of the following aspects: sampling effort, habitat/depth representation, and geographical coverage. The 10 most speciose families (Gobiidae, Labridae, Pomacentridae, Serranidae, Apogonidae, Blenniidae, Carangidae, Scorpaenidae, Lutjanidae and Chaetodontidae) collectively account for about 42% of the total fish fauna. The cumulative percentage contribution when the top 20 families are taken into account (Table 1) is over half the total (56%), suggesting that the fauna is not predominated by only a few families but by a set of several families. This seems to reflect the diversity of habitats encompassed by the current faunal list. Each habitat usually contains a unique composition of the predominant fish families, except for the Gobiidae, which is the most diverse group in nearly all coastal habitats (e.g., coral reefs, seagrass beds and mangroves – Satapoomin, 2006, 2007; Fish Team of the Trang Project, 2002). The current checklist, which includes species from 35 The fishes of southwestern Thailand Table 1. Twenty most speciose families of fishes of southwestern Thailand, the Andaman Sea. Family Number of genera Number of species % of total fish fauna Gobiidae Labridae Pomacentridae Serranidae Apogonidae Blenniidae Carangidae Scorpaenidae Lutjanidae Chaetodontidae Acanthuridae Muraenidae Syngnathidae Carcharhinidae Nemipteridae Engraulidae Lethrinidae Leiognathidae Tetraodontidae Scaridae 62 30 18 17 14 21 20 23 10 7 5 7 11 7 3 5 5 8 8 6 227 78 71 61 60 52 52 49 39 37 32 32 26 25 25 24 22 21 21 20 13.0 4.5 4.1 3.5 3.4 3.0 3.0 2.8 2.2 2.1 1.8 1.8 1.5 1.4 1.4 1.4 1.3 1.2 1.2 1.1 Subtotal 287 974 55.8 Other 176 families 409 772 44.2 Total 696 1,746 100.0 various habitats, thus evens out the significance contribution of certain predominant families in the analysis. The relative abundance of these families is similar, although with variable ranking, to those in studies of inshore fishes at many other IndoPacific localities, for example, Reunion Island (Letourneur et al., 2004), Maldive Islands (Randall and Anderson, 1993), Chagos Archipelago (Winterbottom and Anderson, 1999), northwestern Australia (Allen, 1993), Calamianes Islands, Palawan Province (Allen, 2000), southern Taiwan (Shao et al., 2008), Yaku-shima Island, southern Japan (Motomura et al., 2010), and Mariana Islands (Myers and Donaldson, 2003). The Gobiidae, in particular, was usually the leading family when it was adequately sampled. Other less diverse families include 25 families with a range of species numbers of 10–19 species, 27 families with 5–9 species, 59 families with 2–4 species, and the remaining 66 families each with single species. With respect to habitat association, the fish fauna consists overwhelmingly of reef fishes, which comprise about 56% (983 species) of the total species. The coastal species from mangroves and seagrass beds represent about 17% (296 species) and 14% (246 species) of the total fauna, respectively. The other major component, which is loosely categorized to accommodate fishes inhabiting a variety of offshore habitats (spanning pelagic to benthic zones at various depths), comprises about 55% (971 species) of the total 36 Phuket mar. biol. Cent. Res. Bull. fauna. The figures accounting for the proportion of species shared between habitats vary considerably and are relatively low, i.e., 42% between mangroves and seagrass beds, 20.5% between offshore habitats and coral reefs, 16.8% between offshore habitats and seagrass beds, 15.6% between offshore habitats and mangroves, 9.4% between coral reefs and seagrass beds, and 5.3% between coral reefs and mangroves. The species shared among the 4 biotopes account for only 2.3% of the total species recorded. Species shared between or among habitats, however, comprise a number of species which are strongly dependent on more than a single habitat type, and often exhibit a segregated habitat-use pattern with respect to their different life-history stages (e.g., Mumby et al., 2004; Shibuno et al., 2008). This seems to suggest an inter-system connection in fish populations. Because unique assemblages of species are restricted to each particular habitat type, while others depend upon the existence of multiple habitat-types through connectivity, special attention must be paid to holistic approaches of multi-ecosystems management and conservation if the overall diversity of the fishes is to be maintained. At present, there is growing concern over the science of connectivity because understanding of the linkage among ecosystems is considered as a key to the success of conservation and management of marine and coastal areas, particularly when emphasizing the design of marine protected areas (MPAs) and MPA network (Botsford et al., 2009; Sale et al., 2010). The zoogeographic analysis of species with adequately known distribution ranges is presented in Table 2. The analysis reveals that the fish fauna consists primarily of species which are widely distributed either in the Indo-Pacific or Indowest Pacific (50.7%). Approximately 17.1% of the fauna are found within portions of the western Pacific and northeastern Indian Ocean (i.e., NIO– WCP, NIO–WP, NIO–IA, NIO–IM, EIO–WCP, EIO–WP, EIO–IA, and EIO–IM). Similarly 15.5% of species have a distribution range in the western Pacific or portions of this region (i.e., WCP, WP, IA, and IM). This clearly confirms that eastern most boundary of the Indian Ocean including the Andaman Sea overlaps the biogeographic region of the West Pacific. About 12% of the fauna has a primarily Indian Ocean range, with lesser values (Table 3) for either the fauna confined to the eastern extremity of the Indian Ocean (about 1%) or the fauna restricted only to the Andaman Sea province (1.7%). The remaining species (4.9%) have panoceanic ranges, including circumtropical and cosmopolitan distributions. The presence of fauna having both an Indian Ocean distribution range and also a western Pacific distribution range in the area can be considered as compelling evidence of a distribution confluence of both Indian Ocean and Pacific Ocean faunas in the region. The boundary where sympatry of the eastern (= Pacific Ocean) and western (= Indian Ocean) populations is most evident seems to encompass Christmas Island and the Java–Bali region (see, Randall, 1998; Allen and Adrim, 2003; Hobbs et al., 2010). The strong influence of the Indian Ocean fauna in the area can be further highlighted by considering the number of sibling species that involve Indian and Pacific Ocean species pairs, and some geographic variants that contain different Indian and Pacific Ocean colour morphs. While inconsistent considerations among ichthyologists over taxonomic status (i.e., variants, subspecies, or full species) of the latter exist, concrete morphological and/or genetic evidence is required to ascertain their taxonomic status (Randall, 1998; Hastings and Springer, 2009). As exemplified in Table 4, which includes 68 sibling species-pairs (in 17 families) and 18 species (in 10 families) possessing geographic variation in colour morphs, the Andaman Sea fauna is predominantly represented by the Indian Ocean forms. Since establishment of the PMBC Reference Collection about 25 years ago, a great deal of knowledge has accumulated and well over 190 new species (see also Aungtonya et al., 2006) have been described from the Andaman Sea. At present, the Reference Collection maintains type materials of 21 fish species, including 9 holotypes and 27 lots of paratypes (Table 5). Reliable records of Andaman Sea fishes currently deposited at the Reference Collection account for about 63% of the total fish fauna. The fish collection thus still does not include a comprehensive representation of the Andaman Sea fish fauna. In order to maintain its significant role in supporting biodiversity research of the ichthyofauna of this particular 37 The fishes of southwestern Thailand region, the fish collection needs to be upgraded by additional sampling so that the missing 27% of species can be made available as voucher or reference materials for future researchers. Also worth mentioning here are the figures reflecting biotope representation of the fishes catalogued in the Reference Collection, which vary considerably, i.e., 64% of total offshore fishes and 65% of the reef-associated fishes, but 81% of the mangrove fishes and 88% of the seagrass fishes. These figures can form a basis for targeting future sampling programs for the Reference Collection. Small and cryptic reef fishes and particularly those inhabiting the deep-reef habitats are in need of extensive sampling. For fishes dwelling among various offshore habitats, deep demersal, benthopelagic, and mesopelagic fishes are among the groups that have not been adequately sampled. Deep sea exploratory research programs are generally costly and particularly require specific techniques, equipment as well as taxonomic expertise. The establishment of such programs is likely to require international collaborative efforts and funding. Although the present checklist provides the most extensive species records for the area, it should be considered as a provisional rather than a complete faunal list. Also, there are a number of identifications that still need verification through examination of voucher specimens. Furthermore, the sampling efforts for several fish groups and/ or in certain habitats remain deficient. The present provisional checklist, however, can at least help to draw the attention of ichthyologists who are interested in fish fauna of the Andaman Sea to where there are knowledge gaps. Also, it can serve as a baseline for a comprehensive taxonomic review and/or annotated checklist of the Andaman Sea fishes. It is hoped that, joint research or collaborative efforts will be possible in the near future to accomplish such goals. While the current limited effort and interest, especially with respect to taxonomy, has slowed down the progress in this research area, a concomitant obstacle regarding decreasing personnel trained in ichthyology in Thailand seems to be more critical and needs urgent action. Table 2. Zoogeographic analysis of the fish fauna of southwestern Thailand, the Andaman Sea. Distribution Indo-Pacific (IP) Indo-West Pacific (IWP) West central Pacific (WCP) West Pacific (WP) Indo-Australian Archipelago (IA) Indo-Malayan Archipelago (IM) Indian Ocean (IO) Northern Indian Ocean (NIO) Eastern Indian Ocean (EIO) Northern Indian Ocean to West central Pacific (NIO–WCP) Northern Indian Ocean to West Pacific (NIO–WP) Northern Indian Ocean to Indo-Australian Archipelago (NIO–IA) Northern Indian Ocean to Indo-Malayan Archipelago (NIO–IM) Eastern Indian Ocean to West central Pacific (EIO–WCP) Eastern Indian Ocean to West Pacific (EIO–WP) Eastern Indian Ocean to Indo-Australian Archipelago (EIO–IA) Eastern Indian Ocean Indo-Malayan Archipelago (EIO–IM) Eastern extremity of Indian Ocean (EEIO) Andaman Sea (AS) Circumtropical (CIR) Cosmopolitan (COS) No. species % 417 448 27 137 48 53 99 10 46 2 37 21 29 12 112 28 50 15 28 38 46 24.5 26.3 1.6 8.0 2.8 3.1 5.8 0.6 2.7 0.1 2.2 1.2 1.7 0.7 6.6 1.6 3.0 0.9 1.7 2.2 2.7 38 Phuket mar. biol. Cent. Res. Bull. Table 3. List of fishes currently known to have restricted distribution in the Andaman Sea and adjacent areas: A) Andaman Sea endemics, including the species ranges southward to western Sumatra; B) The species found in Andaman Sea and some other localities along eastern edge of the eastern Indian Ocean (= eastern extremity of Indian Ocean (EEIO)) Family: Species A)Andaman Sea endemics Myxinidae: Eptatretus indrambaryai Wongratana, 1983 Proscylliidae: Proscyllium magnificum Last & Vongpanich, 2004 Rajidae: Anacanthobatis sp.1 Acropomatidae: Acropoma argentistigma Okamoto & Ida, 2002 Aploactinidae: Cocotropus echinatus (Cantor, 1850) Apogonidae: Apogon sp.1 Archamia ataenia Randall & Satapoomin, 1999 Blenniidae: Blenniella leopardus (Fowler, 1904) Meiacanthus urostigma Smith-Vaniz, Satapoomin & Allen, 2001 Bythitidae: Alionematichthys phuketensis Møller & Schwarzhans, 2008 Gobiidae: Callogobius andamanensis Menon & Chatterjee, 1974 Scartelaos cantoris (Day, 1871) Trimma fucatum Winterbottom & Southcott, 2007 Trimma sanguinellus Winterbottom & Southcott, 2007 Haemulidae: Hapalogenys merguiensis Iwatsuki, Satapoomin & Amaoka, 2000 Plectorhinchus macrospilus Satapoomin & Randall, 2000 Pomadasys andamanensis McKay & Satapoomin, 1994 Labridae: Cirrhilabrus joanallenae Allen, 2000 Halichoeres kallochroma (Bleeker, 1853) Monacanthidae: Paraluteres sp.1 Ophichthidae: Allips concolor McCosker, 1972 Phallostethidae: Phenacostethus posthon Roberts, 1971 Plesiopidae: Plesiops thysanopterus Mooi, 1995 Pomacentridae: Chromis xouthos Allen & Erdmann, 2005 Pomacentrus polyspinus Allen, 1991 Serranidae: Cephalopholis polyspila Randall & Satapoomin, 2000 Meganthias filiferus Randall & Heemstra, 2007 Distribution Andaman Sea Andaman Sea Andaman Sea Andaman Sea Andaman Sea West coast of Thailand Andaman Sea–W. Sumatra Andaman Sea–W. Sumatra Andaman Sea–W. Sumatra West coast of Thailand Andaman Sea Andaman Sea West coast of Thailand Andaman Sea–W. Sumatra Andaman Sea West coast of Thailand West coast of Thailand Andaman Sea–W. Sumatra Andaman Sea–W. Sumatra Andaman Sea Andaman Sea Andaman Sea N. Sumatra (Andaman Sea) Andaman Sea–W. Sumatra West coast of Thailand Andaman Sea–W. Sumatra Andaman Sea 39 The fishes of southwestern Thailand Table 3. (continued) Family: Species Soleidae: Aseraggodes satapoomini Randall & Desoutter-Meniger, 2007 Sternoptychidae: Polyipnus asper Harold, 1994 Tripterygiidae: Helcogramma lacuna Williams & Howe, 2003 B) Limited to eastern extremity of Indian Ocean (EEIO) Urolophidae: Urolophus javanicus (Martens, 1864) Blenniidae: Ecsenius lubbocki Springer, 1988 Ecsenius paroculus Springer, 1988 Praealticus triangulus (Chapman, 1951) Labridae: Cirrhilabrus cyanopleura (Bleeker, 1851) Malacanthidae: Hoplolatilus luteus Allen & Kuiter, 1989 Opistognathidae: Opistognathus cyanospilotus Smith-Vaniz, 2009 Polynemidae: Filimanus perplexa Feltes, 1991 Pomacentridae: Pomacentrus alleni Burgess, 1981 Pomacentrus azuremaculatus Allen, 1991 Pomacentrus xanthosternus Allen, 1991 Pseudochromidae: Pseudochromis andamanensis Lubbock, 1980 Scaridae: Chlorurus troschelii (Bleeker, 1853) Scarus maculipinna Westneat, Satapoomin & Randall, 2007 Siganidae: Siganus magnificus (Burgess, 1977) Distribution West coast of Thailand Andaman Sea West coast of Thailand Andaman Sea, Java Sea (W. Java) Andaman Sea, Bali Andaman Sea., W. Sumatra, Java Sea (Seribu Is.& N. Java) Andaman Sea, Java Sea (N. Java) Andaman Sea, W. Sumatra, Christmas Is., Java Sea (Pulau Putri & N. Java), Bali Andaman Sea, Bali, Flores Andaman Sea, Bali Andaman Sea, W. Sumatra, S. Java, Bali Andaman Sea, Christmas Is., Java Sea (Sunda Strait & Pulau Putri) Andaman Sea, Java Sea (Seribu Is.) Andaman Sea, W. Sumatra, Java Sea (Seribu Is.), Bali Andaman Sea, W. Sumatra, Bali, Timor Sea (Ashmore Reef & Cartier Is., N. Australia) Andaman Sea, Java Sea (Seribu Is.) Andaman Sea, W. Sumatra, Java Sea (Pulau Putri) Andaman Sea, W. Sumatra, Java Sea (W. Java) 40 Phuket mar. biol. Cent. Res. Bull. Table 4. Examples of Indian and Pacific Ocean species pairs and geographic variants that the Andaman Sea fishes represented by the Indian Ocean forms. A) Sibling species: Andaman Sea fish (vs. its Pacific sibling) Acanthuridae Acanthurus tristis (A. pyroferus) Ctenochaetus truncatus (C. strigosus) Naso elegans (N. lituratus) Zebrasoma desjardinii (Z. veliferum) Apogonidae Archamia ataenia (A. zosterophora) Blenniidae Aspidontus tractus (A. taeniatus) Atrosalarias fuscus (A. holomelas) Blenniella cyanostigma (B. caudolineata) Blenniella leopardus (B. interrupta) Meiacanthus urostigma (M. ditrema) Plagiotremus phenax (P. laudandus) Chaetodontidae Chaetodon andamanensis (C. plebeius) Chaetodon triangulum (C. baronessa) Chaetodon falcula (C. ulietensis) Chaetodon gardineri (C. selene) Chaetodon interruptus (C. unimaculatus) Chaetodon madagaskariensis (C. mertensii) Chaetodon melannotus (C. ocellicaudus) Chaetodon trifasciatus (C. lunulatus) Hemitaurichthys zoster (H. polylepis) Heniochus pleurotaenia (H. varius) Gobiidae Amblygobius semicinctus (A. phalaena) Trimma flammeum (T. macrophthalma) Labridae Anampses lineatus (A. melanurus) Cirrhilabrus cyanopleura (C. lyukyuensis) Coris cuvieri (C. gaimard) Bodianus diana (B. dictynna) Gomphosus caeruleus (G. varius) Halichoeres cosmetus (H. claudia) Halichoeres kallochroma (H. leucurus) Halichoeres leucoxanthus (H. chrysus) Halichoeres vrolokii (H. melanurus) Halichoeres zeylonicus (H. hartzfeldii) Paracheilinus mccoskeri (P. carpenteri) Pseudojuloides kaleidos (P. cerasinus) Mullidae Parupeneus trifasciatus (P. crassilabris) Pinguipedidae Parapercis maculata (P. pulchella) Pomacanthidae Centropyge eibli (C. vroliki) Genicanthus caudovittatus (G. melanospilos) Pomacentridae Amphiprion akallopisos (A. sandarcinos) Amphiprion ephippium (A. melanopus) Amphiprion sebae (A. polymnus) Chromis dimidiata (C. iomelas) Chromis opercularis (C. xanthura) Dascyllus carneus (D. reticulatus) Neopomacentrus sororius (N. azysron) Pomacentrus similis (P. coelestis) Pomacentrus xanthosternus (P. simsiang) Ptereleotridae Ptereleotris arabica (P. hanae) Scaridae Chlorurus capistratoides (C. japanensis) Chlorurus rhakoura (C. oedema) Chlorurus sordidus (C. spilurus) Chlorurus strongylocephalus (C. microrhinos) Chlorurus troschelii (C. bleekeri) Hipposcarus harid (H. longiceps) Scarus maculipinna (S. flavipectoralis) Scarus russelii (S. schlegeli) Scarus scaber (S. oviceps) Scarus viridifucatus (S. spinus) Scorpaenidae Pterois miles (P. volitans) Serranidae Cephalopholis polyspila (C. microprion) Pogonoperca ocellata (P. punctata) Pseudanthias ignitus (P. dispar) Rabualichthys stigmaticus (R. altipinnis) Siganidae Siganus puelloides (S. puellus) Siganus stellatus (S. punctatus) Syngnathidae Doryramphus multiannulatus (D. pessuliferus) Tetraodontidae Canthigaster smithae (C. epilampra) 41 The fishes of southwestern Thailand Table 4. (continued) B) Geographic variants: Andaman Sea fish possessing Indian Ocean colour morph Dasyatidae *(4) Dasyatis kuhlii Acanthuridae *(7) Paracanthurus hepatus Balistidae *(5, 7) Balistapus undulatus Caesionidae Caesio cuning Labridae *(9) Epibulus insidiator *(5, 7) Halichoeres hortulanus *(5) Labroides dimidiatus *(7) Thalassoma jansenii Lutjanidae *(1, 7) Lutjanus russelli Nemipteridae *(6, 7) Scolopsis bilineatus Scolopsis vosmeri Pomacanthidae *(7, 8) Pomacanthus imperator *(5, 7) Pygoplites diacanthus Pomacentridae *(2) Chrysiptera rollandi *(7) Neoglyphidodon nigroris *(6) Pomacentrus chrysurus *(2, 8) Pomacentrus pavo Serranidae *(3) Cephalopholis urodeta *Note: Recognition of allopatric colour morphs in certain species can be referred to (1) Allen and Talbot, 1985; (2) Allen, 1991; (3) Randall and Heemstra, 1991; (4) Randall, 1995; (5) Randall, 1998; (6) Satapoomin, 2000; (7) Kuiter and Tonozuka, 2001; (8) Lieske and Myers, 2001; and (9) Carlson et al., 2008. Table 5. List of fishes for which their type materials are deposited at the Reference Collection of the Phuket Marine Biological Center. Number in bracket denotes the number of catalogued lots for paratypes. Family: Species Apogonidae: Archamia ataenia Randall & Satapoomin, 1999 Blenniidae: Meiacanthus urostigma Smith-Vaniz, Satapoomin & Allen, 2001 Gobiidae: Cryptocentrus crocatus Wongratana, 1975 [valid as Myersina crocatus (Wongratana, 1975)] Gladiogobius rex Shibukawa & Allen, 2007 Myersina adonis Shibukawa & Satapoomin, 2006 Periophthalmus walailakae Darumas & Tantichodok, 2002 Trimma fucatum Winterbottom & Southcott, 2007 Trimma sanguinellus Winterbottom & Southcott, 2007 Haemulidae: Hapalogenys merguiensis Iwatsuki, Satapoomin & Amaoka, 2000 Plectorhinchus macrospilus Satapoomin & Randall, 2000 Pomadasys andamanensis McKay & Satapoomin, 1994 Nemipteridae: Nemipterus aurora Russell, 1993 Nemipterus thosaporni Russell, 1991 Type materials Paratypes (1) Paratypes (2) Holotype Holotype, Paratypes (3) Holotype, Paratypes (1) Paratypes (2) Paratypes (1) Paratypes (1) Paratypes (2) Paratypes (1) Holotype Paratypes (1) Paratypes (1) 42 Phuket mar. biol. Cent. Res. Bull. Table 5. (continued) Family: Species Type materials Opistognathidae: Opistognathus variabilis Smith-Vaniz, 2009 Proscylliidae: Proscyllium magnificum Last & Vongpanich, 2004 Pomacentridae: Pomacentrus azuremaculatus Allen, 1991 Pomacentrus polyspinus Allen, 1991 Pomacentrus similis Allen, 1991 Scaridae: Scarus maculipinna Westneat, Satapoomin & Randall, 2007 Serranidae: Cephalopholis polyspila Randall & Satapoomin, 2000 Meganthias filiferus Randall & Heemstra, 2007 ACKNOWLEDGEMENTS This paper was initially prepared as a contribution to the concluding conference of the 10-year JSPS Multilateral Core University Programme on “Coastal Marine Science” in Tokyo during 26–29 October 2010, but I finally could not make the trip. It was then presented as a plenary talk at the seminar on “Biodiversity of Southern Thailand in Focus: Status and Research Direction” at Walailak University, Nakorn Sri Thammarat, Thailand, during 10–12 November 2010. I am grateful for the generous support made available by the JSPS for me to be able to join several fish workshops conducted over the years, which ultimately gave me a wonderful opportunity to learn more about the fish faunas of SE Asia. Special Paratypes (2) Holotype, Paratypes (3) Holotype Holotype Paratypes (1) Holotype, Paratypes (1) Paratypes (4) Holotype thanks are due to Dr. Keiichi Matsuura [KM] of the National Museum of Nature and Science, Tokyo, for his considerable effort in leading the fish diversity project under the JSPS multilateral programme and encouraging me to write up this paper. I am very grateful to Dr. Richard Winterbottom [RW] of the Royal Ontario Museum for sharing the research outcomes from the 1993 expedition to Phuket and inspiring my interest in small cryptobenthic reef fishes. Both KM and RW kindly gave their time for reviewing and improving the first draft of this manuscript. 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Nat. Hist. Bull. Siam Soc. 31(1): 1–7. Manuscript received: 10 January 2011 Accepted: 1 August 2011 47 The fishes of southwestern Thailand Appendix A Checklist of marine fishes of southwestern Thailand, the Andaman Sea Abbreviations: 1. Habitat types – Cr (coral reefs); Mg (mangroves); Sg (seagrass beds); OShr (other coastal and offshore areas) 2. Distribution categories – IP (Indo-Pacific); IWP (IndoWest Pacific); WCP (west central Pacific); WP (west Pacific); IA (Indo-Australian Archipelago); IM (IndoMalayan Archipelago); IO (Indian Ocean); NIO (northern Indian Ocean); EIO (eastern Indian Ocean); EEIO (eastern extremity of Indian Ocean); AS (Andaman Sea); CIR (circumtropical); COS (cosmopolitan) [see methods section for definitions] 3. Voucher specimens – PMBC (denotes availability of voucher specimens at the Reference Collection of the Phuket Marine Biological Center) CLASS MYXINI Order Myxiniformes Myxinidae Eptatretus indrambaryai Wongratana, 1983: OShr; [AS]; PMBC CLASS CHONDRICHTHYES Order Chimaeriformes Chimaeridae Hydrolagus sp.: OShr; PMBC Order Orectolobiformes Parascylliidae Parascyllium sp.: OShr; PMBC Hemiscylliidae Chiloscyllium griseum Müller & Henle, 1838: OShr; [NIOIM]; PMBC Chiloscyllium hasselti Bleeker, 1852: Sg, OShr; [IM]; PMBC Chiloscyllium indicum (Gmelin, 1788): OShr; [IWP]; PMBC Chiloscyllium plagiosum (Bennett, 1830): OShr; [IWP] Chiloscyllium punctatum Müller & Henle, 1838: Cr, Sg, OShr; [EIO-WP] Stegostomatidae Stegostoma fasciatum (Hermann, 1793): Cr, OShr; [IP]; PMBC Ginglymostomatidae Nebrius ferrugineus (Lesson, 1830): Cr, OShr; [IWP] Rhincodontidae Rhincodon typus Smith, 1828: Cr, OShr; [COS] Order Lamniformes Alopiidae Alopias pelagicus Nakamura, 1935: OShr; [CIR] Alopias superciliosus (Lowe, 1840): OShr; [CIR] Alopias vulpinus (Bonnaterre, 1788): OShr; [COS] Lamnidae Isurus oxyrinchus Rafinesque, 1810: OShr; [COS] Order Carcharhiniformes Scyliorhinidae Atelomycterus marmoratus (Bennett, 1830): OShr; [IWP] Cephaloscyllium umbratile Jordan & Fowler, 1903: OShr; [WP]; PMBC Halaelurus buergeri (Müller & Henle, 1838): OShr; [WP] Halaelurus hispidus (Alcock, 1891): OShr; [EIO-WP]; PMBC Proscylliidae Proscyllium magnificum Last & Vongpanich, 2004: OShr; [AS]; PMBC Triakidae Iago omanensis (Norman, 1939): OShr; [IO]; PMBC Mustelus manazo Bleeker, 1854: OShr; [IWP]; PMBC Mustelus mosis Hemprich & Ehrenberg, 1899: OShr; [IO]; PMBC Hemigaleidae Chaenogaleus macrostoma (Bleeker, 1852): OShr; [IWP] Hemigaleus microstoma Bleeker, 1852: OShr; [EIO-WP]; PMBC Hemipristis elongatus (Klunzinger, 1871): OShr; [IWP]; PMBC Carcharhinidae Carcharhinus albimarginatus (Rüppell, 1837): Cr, OShr; [IP] Carcharhinus altimus (Springer, 1950): OShr; [COS] Carcharhinus amblyrhynchoides (Whitley, 1934): OShr; [IWP] Carcharhinus amblyrhynchos (Bleeker, 1856): OShr; [IP] Carcharhinus amboinensis (Müller & Henle, 1839): OShr; [IWP] Carcharhinus brachyurus (Günther, 1870): OShr; [COS] Carcharhinus brevipinna (Müller & Henle, 1839): OShr; [CIR] Carcharhinus dussumieri (Valenciennes, 1839): OShr; [NIO-WP] Carcharhinus falciformis (Müller & Henle, 1839): OShr; [CIR] Carcharhinus leucas (Valenciennes, 1839): OShr; [CIR] Carcharhinus limbatus (Valenciennes, 1839): OShr; [COS] Carcharhinus longimanus (Poey, 1861): OShr; [COS] Carcharhinus macloti (Müller & Henle, 1839): OShr; [IWP]; PMBC Carcharhinus melanopterus (Quoy & Gaimard, 1824): Cr, OShr; [IP]; PMBC Carcharhinus obscurus (LeSueur, 1818): OShr; [COS] Carcharhinus plumbeus (Nardo, 1827): OShr; [CIR]; PMBC Carcharhinus sealei (Pietschmann, 1916): OShr; [IWP] Carcharhinus sorrah (Valenciennes, 1839): OShr; [IWP] Galeocerdo cuvier (Péron & LeSueur, 1822): OShr; [CIR] Loxodon macrorhinus Müller & Henle, 1839: OShr; [IWP] Negaprion acutidens (Rüppell, 1837): OShr; [IP] Rhizoprionodon acutus (Rüppell, 1837): OShr; [CIR]; PMBC Rhizoprionodon oligolinx Springer, 1964: OShr; [IWP] 48 Phuket mar. biol. Cent. Res. Bull. Scoliodon laticuadus Müller & Henle, 1839: OShr; [IWP] Triaenodon obesus (Rüppell, 1835): Cr, OShr; [IP] Sphyrnidae Eusphyra blochii (Cuvier, 1817): OShr; [NIO-WP]; PMBC Sphyrna lewini (Griffith & Smith, 1834): OShr; [COS]; PMBC Sphyrna mokarran (Rüppell, 1837): OShr; [CIR] Sphyrna zygaena (Linnaeus, 1758): OShr; [CIR]; PMBC Order Hexanchiformes Hexanchidae Heptranchias perlo (Bonnaterre, 1788): OShr; [COS] Order Squaliformes Squalidae Etmopterus brachyurus Smith & Radcliffe, 1912: OShr; [WP] Squalus acanthias Linnaeus, 1758: OShr; [COS]; PMBC Squalus mitzukurii Jordan & Snyder, 1903: OShr; [CIR]; PMBC Order Squantiformes Squatinidae Squatina sp.1: OShr Order Torpediniformes Narcinidae Narcine brunnea Annandale, 1909: OShr; [NIO-IM]; PMBC Narcine indica Henle, 1834: OShr; [WP] Narcine maculata (Shaw, 1804): OShr; [WP]; PMBC Narcine prodosalis Bessednov, 1966: OShr; [EIO-WP] Narcine timlei (Bloch & Schneider, 1801): OShr; [EIOWP] Narke dipterygia (Bloch & Schneider, 1801): OShr; [NIOWP] Temera hardwickii Gray, 1831: OShr; [IM]; PMBC Order Pristiformes Pristidae Pristis microdon Latham, 1794: OShr; [CIR]; PMBC Order Rajiformes Rhinidae Rhina ancylostoma (Bloch & Schneider, 1801): Cr, OShr; [IP]; PMBC Rhynchobatidae Rhynchobatus palpebratus Compagno & Last, 2008: Cr, OShr; [IA (presently known from west coast of Thailand and northern Australia)] Rhinobatidae Rhinobatos granulatus Cuvier, 1829: OShr; [NIO-IM]; PMBC Rhinobatos thouin (Anonymous in Lacepède, 1798): OShr; [IWP] Rhinobatos typus Bennett, 1830: OShr; [EIO-WP] Rajidae Anacanthobatis sp.1 : OShr, PMBC Raja sp.: OShr Order Myliobatiformes Plesiobatidae Plesiobatis daviesi Wallace, 1967: OShr; [IP]; PMBC Urolophidae Urolophus javanicus (Martens, 1864): OShr; [EEIO] Dasyatidae Dasyatis kuhlii (Müller & Henle, 1841): Cr, OShr; [IWP]; PMBC Dasyatis zugei (Müller & Henle, 1841): OShr; [EIO-WP] Himantura fai Jordan & Seale, 1906: OShr; [IWP] Himantura gerrardi (Gray, 1851): Sg, OShr; [IP]; PMBC Himantura imbricata (Bloch & Schneider, 1801): Mg, Sg, OShr; [IWP]; PMBC Himantura jenkinsii (Annadale, 1909): Cr, OShr; [EIO] Himantura undulata (Bleeker, 1852): OShr; [EIO-WP] Himantura walga (Müller & Henle, 1841): Sg, OShr; [EIOIM]; PMBC Pastinachus sephen (Forsskål, 1775): Cr, OShr; [IWP] Taeniura lymma (Forsskål, 1775): Cr; [IWP]; PMBC Taeniura meyeni Müller & Henle, 1841: Cr; [IP] Urogymnosus asperrimus (Bloch & Schneider, 1801): Cr; [IP] Gymnuridae Gymnura poecilura (Shaw, 1804): Mg, OShr; [IP]; PMBC Myliobatidae Aetobatus narinari (Euphrasen, 1790): Cr, OShr; [COS] Aetomylaeus maculatus (Gray, 1832): OShr; [EIO-WP] Aetomylaeus milvus (Valenciennes, 1841): OShr; [IWP] Aetomylaeus nichofii (Bloch & Schneider, 1801): OShr; [IWP]; PMBC Aetomylaeus vespertilio (Bleeker, 1852): OShr; [IWP] Manta birostris (Donndorff, 1798): Cr, OShr; [CIR] Mobula kuhlii (Valenciennes, in Müller & Henle, 1841): OShr; [IWP] Rhinoptera javanica Müller & Henle, 1841: OShr; [IWP]; PMBC Rhinoptera sewelli Misra, 1946: OShr; [EIO] CLASS ACTINOPTERYGII Order Elopiformes Elopidae Elops hawaiiensis Regan, 1909: Mg, Sg, OShr; [WCP]; PMBC Megalopidae Megalops cyprinoides (Broussonet, 1782): Mg, OShr; [IP]; PMBC Order Albuliformes Albulidae Albula glossodonta (Forsskål, 1775): OShr; [IP] Albula oligolepis Iwatsuki & Hidaka, 2008: OShr; [IO]; PMBC Notacanthidae Notacanthus sp.: OShr; PMBC Order Anguilliformes Anguillidae Anguilla bicolor bicolor McClelland, 1844: Mg; [IWP]; PMBC Anguilla marmorata Quoy & Gaimard, 1824: OShr; [IP]; PMBC Chlopsidae 49 The fishes of southwestern Thailand Kaupichthys sp.: Cr Muraenidae Echidna delicatula (Kaup, 1856): OShr; [EIO-WP]; PMBC Echidna nebulosa (Ahl, 1789): Cr; [IP]; PMBC Echidna polyzona (Richardson, 1844): Cr; [IP]; PMBC Enchelycore bayeri (Schultz, 1953): Cr; [IP] Gymnomuraena zebra (Shaw, 1797): Cr; [IP]; PMBC Gymnothorax annulatus (Smith & Bohlke, 1997): OShr; [IM] Gymnothorax buroensis (Bleeker, 1857): Cr; [IP] Gymnothorax chilospilus Bleeker, 1865: Cr; [IP]; PMBC Gymnothorax favagineus (Bloch & Schneider, 1801): Cr; [IWP]; PMBC Gymnothorax fimbriatus (Bennett, 1832): Cr; [IP]; PMBC Gymnothorax flavimarginatus (Rüppell, 1828): Cr; [IP] Gymnothorax javanicus (Bleeker, 1859): Cr; [IP]; PMBC Gymnothorax margaritophorus Bleeker, 1864: Cr; [IP] Gymnothorax melatremus Schultz, 1953: Cr; [IP] Gymnothorax meleagris (Shaw & Nodder, 1795): Cr; [IP] Gymnothorax moluccensis (Bleeker, 1864): Cr; [IWP] Gymnothorax monochrous (Bleeker, 1856): Cr; [IP] Gymnothorax permistus (Smith, 1962): Cr; [IO]; PMBC Gymnothorax pindae Smith, 1962: Cr; [IP]; PMBC Gymnothorax pseudoherrei Böhlke, 2000: Cr; [IWP] Gymnothorax pseudothyrsoideus (Bleeker, 1852): Cr; [NIO-WP]; PMBC Gymnothorax reticularis Bloch, 1795: OShr; [IO]; PMBC Gymnothorax richardsoni (Bleeker, 1852): Cr; [IWP]; PMBC Gymnothorax rueppelliae (Mcclelland, 1845): Cr; [IP] Gymnothorax thyrsoideus (Richardson, 1844): Cr, OShr; [EIO-WCP]; PMBC Gymnothorax undulatus (Lacepède, 1803): Cr; [IP]; PMBC Gymnothorax zonipectis Seale, 1906: Cr; [IP]; PMBC Rhinomuraena quaesita Garman, 1888: Cr; [IWP] Scuticaria tigrina (Lesson, 1828): Cr; [IP] Uropterygius concolor Rüppell, 1838: Cr; [IP]; PMBC Uropterygius inornatus Gosline, 1958: Cr; [WCP] Uropterygius xanthopterus Bleeker, 1859: Cr; [IP] Ophichthidae Allips concolor McCosker, 1972: Cr, Mg; [AS] Apterichthus sp.: OShr; PMBC Brachysomophis crocodilinus (Bennet, 1833): Cr, OShr; [IP] Echelus uropterus (Temminck & Schlegel, 1846): OShr; [IWP]; PMBC Muraenichthys gymnopterus (Bleeker, 1853): OShr; [EIOWP] Myrichthys colubrinus (Boddaert, 1781): Cr; [IP]; PMBC Neenchelys sp.: OShr; PMBC Ophichthus evermanni Jordan & Richardson, 1909: OShr; [WP] Ophichthus sp.: OShr; PMBC Pisonodophis cancrivorus (Richardson, 1844): Mg, Sg, OShr; [IWP]; PMBC Scolecenchelys gymnota (Bleeker, 1854): Sg; [IWP] Scolecenchelys macroptera (Bleeker, 1857): Sg, OShr; [IWP]; PMBC Xyrias revulsus Jordan & Snyder, 1901: OShr; [IWP] Colocongridae Coloconger scholesi Chan, 1967: OShr; [IWP]; PMBC Coloconger sp.: OShr; PMBC Muraenesocidae Congresox talabon (Cuvier, 1829): OShr; [EIO-IM]; PMBC Congresox talabonoides (Bleeker, 1853): OShr; [IWP]; PMBC Muraenosox bagio (Hamilton, 1822): OShr; [IWP] Muraenosox cinereus (Forsskål, 1775): OShr; [IWP]; PMBC Congridae Ariosoma anago (Temminck & Schlegel, 1846): OShr; [IWP]; PMBC Bathycongrus guttulatus (Günther, 1887): OShr; [IP] Bathyuroconger sp.: OShr; PMBC Conger cinereus Rüppell, 1828: OShr; [IP]; PMBC Gnathophis sp.: OShr; PMBC Gorgasia preclara Böhlke & Randall, 1981: Cr; [EIO-IA] Heteroconger hassi (Klausewitz & Eibl-Eibesfeldt, 1959): Cr; [IP] Rhynchoconger squaliceps (Alcock, 1894): OShr; [EIO] Uroconger lepturus (Richardson, 1845): OShr; [IWP]; PMBC Nettastomatidae Nettastoma sp.: OShr Order Clupeiformes Pristigasteridae Ilisha elongata (Bennett, 1830): OShr; [EIO-IM]; PMBC Ilisha kampeni (Weber & Beaufort, 1913): Sg, OShr; [EIOIM] Ilisha megaloptera (Swainson, 1839): Mg, OShr; [EIOIM]; PMBC Ilisha melastoma (Schneider, 1801): Mg, OShr; [EIO-WP]; PMBC Ilisha sirishai Seshagiri Rao, 1975: OShr; [NIO-IM] Opisthopterus tardoore (Cuvier, 1829): Mg, OShr; [NIOIM]; PMBC Pellona ditchela Valenciennes, 1847: Mg, OShr; [IWP]; PMBC Raconda russeliana Gray, 1831: OShr; [EIO-IM] Engraulidae Coilia dussumieri Valenciennes, 1848: OShr; [EIO-IM]; PMBC Coilia macrognathos Bleeker, 1852: OShr; [IM] Coilia neglecta Whitehead, 1968: OShr; [EIO-IM] Enchasicholina punctifer Fowler, 1938: OShr; [IP] Encrasicholina devisi (Whitley, 1940): OShr; [IWP]; PMBC Encrasicholina heteroloba (Rüppell, 1837): OShr; [IWP]; PMBC Setipinna breviceps (Cantor, 1850): OShr; [EIO-IM] Setipinna taty (Valenciennes, 1848): Mg, OShr; [EIO-IM] Setipinna tenuifilis (Valenciennes, 1848): OShr; [EIO-WP] 50 Phuket mar. biol. Cent. Res. Bull. Stolephorus andhraensis Babu Rao, 1966: OShr; [EIOIA]; PMBC Stolephorus baganensis Hardenberg, 1933: Mg, OShr; [EIO-IM] Stolephorus chinensis (Günther, 1880): Mg, OShr; [IM] Stolephorus commersonii Lacepède, 1803: Mg, Sg, OShr; [IWP]; PMBC Stolephorus dubiosus Wongratana, 1983: OShr; [EIO-IM] Stolephorus indicus (van Hasselt, 1823): Mg, Sg, OShr; [IWP]; PMBC Stolephorus insularis Hardenberg, 1933: Mg, OShr; [NIOWP] Stolephorus waitei Jordan & Seale, 1926: OShr; [EIO-IA]; PMBC Thryssa baelama (Forster, 1775): OShr; [IWP] Thryssa dussumieiri (Valenciennes, 1848): OShr; [NIOIM] Thryssa hamiltonii (Gray, 1835): Mg, Sg, OShr; [NIOIA]; PMBC Thryssa kammalensis (Bleeker, 1849): Mg, OShr; [IM] Thryssa mystax (Schneider, 1801): OShr; [NIO-IM]; PMBC Thryssa setirostris (Broussonet, 1782): Mg, Sg, OShr; [IWP]; PMBC Thryssa spinidens (Jordan & Seale, 1925): OShr; [EIO] Chirocentridae Chirocentrus dorab (Forsskål, 1775): Mg, Sg, OShr; [IWP]; PMBC Chirocentrus nudus Swainson, 1839: OShr; [IWP] Clupeidae Amblygaster clupeoides Bleeker, 1849: OShr; [EIO-WP] Amblygaster leiogaster (Valenciennes, 1847): OShr; [IWP] Amblygaster sirm (Walbaum, 1792): OShr; [IWP]; PMBC Anodontostoma chacunda (Hamilton, 1822): Mg, Sg, OShr; [NIO-WP]; PMBC Anodontostoma thailandiae Wongratana, 1983: OShr; [EIOIM] Dussumieria acuta Valenciennes, 1847: OShr; [NIO-IM]; PMBC Dussumieria elopsoides Bleeker, 1849: Mg, OShr; [IWP] Escualosa thoracata (Valenciennes, 1847): Mg, Sg, OShr; [NIO-IA]; PMBC Herklotsichthys quadrimaculatus (Rüppell, 1837): Cr, OShr; [IP]; PMBC Hilsa kelee (Cuvier, 1829): Mg, Sg, OShr; [IWP] Nematalosa galatheae Nelson & Rothman, 1973: OShr; [EIO-IM] Nematalosa nasus (Bloch, 1795): OShr; [EIO-WP] Sardinella albella (Valenciennes, 1847): Mg, Sg, OShr; [IWP]; PMBC Sardinella fimbriata (Valenciennes, 1847): OShr; [EIO-IA]; PMBC Sardinella gibbosa (Bleeker, 1849): OShr; [IWP]; PMBC Sardinella lemura Bleeker, 1853: OShr; [WP] Spratelloides delicatulus (Bennett, 1831): Cr, OShr; [IP]; PMBC Spratelloides gracilis (Temminck & Schlegel, 1846): Cr, Mg, Sg, OShr; [IP] Tenualosa reevesii (Richardson, 1846): OShr; [WP] Tenualosa toli (Valenciennes, 1847): OShr; [NIO-IM]; PMBC Order Gonorynchiformes Chanidae Chanos chanos (Forsskål, 1775): Cr, Mg, OShr; [IP] Order Siluriformes Plotosidae Plotosus canius Hamilton, 1822: Mg, Sg, OShr; [EIO-IA]; PMBC Plotosus lineatus (Thunberg, 1787): Cr, Mg, Sg, OShr; [IWP]; PMBC Ariidae Arius arius (Hamilton, 1822): OShr; [EIO-IM]; PMBC Arius caelatus Valenciennes, 1840: Mg; [EIO-IM] Arius gagora (Hamilton, 1822): OShr; [EIO] Arius oetik Bleeker, 1846: OShr; [WP] Arius venosus Valenciennes, 1840: OShr; [EIO-IM] Nemapteryx nenga (Hamilton, 1822): OShr; [EIO-IM] Nemuta thalassinus (Rüppell, 1837): OShr; [IWP]; PMBC Osteogeneiosus militaris (Linnaeus, 1758): OShr; [EIOIM] Plicofollis nella Bleeker, 1846: OShr; [WP] Bagridae Mystus gulio (Hamilton, 1822): Mg; [EIO-IM]; PMBC Order Argentiniformes Alepocephalidae Alepocephalus bicolor Alcock, 1891: OShr; [IWP]; PMBC Order Stomiiformes Sternoptychidae Argyropelecus affinis Garman, 1899: OShr; [CIR]; PMBC Argyropelecus sladeni Regan, 1908: OShr; [CIR]; PMBC Polyipnus asper Harold, 1994: OShr; [AS]; PMBC Sternoptyx obscura Garman, 1899: OShr; [IWP]; PMBC Stomiidae Astronesthes lucifer Gilbert, 1905: OShr; [IP] Chauliodus sloanei Bloch & Schneider, 1801: OShr; [COS] Eustomias sp.: OShr; PMBC Order Ateleopodiformes Ateleopodidae Ateleopus indicus Alcock, 1891: OShr; [EIO]; PMBC Order Aulopiformes Paraulopidae Paraulopus brevirostris (Fourmanoir, 1981): OShr; [WP] Synodontidae Harpadon nehereus (Hamilton, 1822): OShr; [IWP] Saurida elongata (Temminck & Schlegel, 1846): OShr; [WP]; PMBC Saurida gracilis (Quoy & Gaimard, 1824): Cr, OShr; [IP]; PMBC Saurida longimanus Norman, 1939: OShr; [NIO]; PMBC Saurida micropectoralis Shindo & Yamada, 1972: Mg, OShr; [WP]; PMBC Saurida nebulosa Valenciennes, 1849: Cr, Mg, Sg, OShr; [IP]; PMBC 51 The fishes of southwestern Thailand Saurida tumbil (Bloch, 1795): OShr; [IWP]; PMBC Saurida undosquamis (Richardson, 1848): Cr, OShr; [IWP]; PMBC Synodus dermatogenys Fowler, 1912: Cr, OShr; [IP]; PMBC Synodus hoshinonis Tanaka, 1917: OShr; [IWP]; PMBC Synodus indicus (Day, 1873): OShr; [IWP]; PMBC Synodus jaculum Russell & Cressey, 1979: Cr; [IP]; PMBC Synodus rubromarmoratus Russell & Cressey, 1979: Cr; [WP] Synodus variegatus (Lacepède, 1801): Cr, OShr; [IP]; PMBC Trachinocephalus myops (Bloch & Schneider, 1801): Mg, Sg, OShr; [CIR]; PMBC Chlorophthalmidae Chlorophthalmus corniger Alcock, 1894: OShr; [EIO-WP] Alepisauridae Alepisaurus ferox Lowe, 1833: OShr; [COS] Paralepididae Lestolepis luetkeni (Ege, 1933): OShr; [IP] Maculisudis longipinnis Kotthaus, 1967: OShr Paralepis philippinus Fowler, 1934: OShr Order Myctophiformes Neoscopelidae Neoscopelus macrolepidotus Jonhson, 1863: OShr; [COS] Neoscopelus microchir Matsubara, 1943: OShr; [CIR]; PMBC Myctophidae Benthosema pterotum (Alcock, 1890): OShr; [IWP] Diaphus coeruleus (Kluzinger, 1871): OShr; [IWP] Diaphus splendidus (Brauer, 1904): OShr; [CIR] Diaphus thiollierei Fowler, 1934: OShr; [IWP] Order Lampriformes Lophotidae Lophotus lecepede Giorna, 1809: OShr; [CIR]; PMBC Trachipteridae Trachipterus jacksonensis (Ramsay, 1881): OShr; [COS] Zu cristatus (Bonelli, 1820): OShr; [CIR]; PMBC Order Polymixiiformes Polymixiidae Polymixia berndti Gilbert, 1905: OShr; [IP] Order Gadiformes Bregmacerotidae Bregmaceros mcclellandi Thompson, 1840: OShr; [EIO]; PMBC Macrouridae Coelorinchus argentatus Smith & Radcliffe, 1912: OShr; [IWP] Coelorinchus macrorhynchus Smith & Radcliffe, 1912: OShr; [WP] Coelorinchus radcliffei Gilbert & Hubbs, 1920: OShr; [WP] Coryphaenoides asprellus (Smith & Radcliff, 1912): OShr; [IO]; PMBC Malacocephalus laevis (Lowe, 1843): OShr; [COS] Moridae Physiculus roseus Alcock, 1891: OShr; [EIO-WP] Carapidae Encheliophis homei (Richardson, 1846): Sg; [IWP]; PMBC Onuxodon fowleri (Smith, 1955): Cr; [IP] Onuxodon parvibranchium (Fowler, 1927): Cr; [IP]; PMBC Pyramodon ventralis Smith & Radcliffe, 1913: OShr; [IP] Order Ophidiiformes Ophidiidae Glyptophidium argenteum Alcock, 1889: OShr; [IM]; PMBC Glyptophidium macropus Alcock, 1894: OShr; [NIO] Hypopleuron caninum Smith & Radcliffe, 1913: OShr; [IWP]; PMBC Neobythites fasciatus Smith & Radcliffe, 1913: OShr; [WP] Bythitidae Alionematichthys phuketensis Møller & Schwarzhans, 2008: Cr; [AS] Alionematichthys piger (Alcock, 1890): Cr; [WCP] Alionematichthys riukiuensis (Aoyaki, 1954): Cr; [WP] Brosmophyciops pautzkei Schultz, 1960: Cr; [IWP]; PMBC Dinematichthys iluocoeteoides Bleeker, 1855: Cr; [IWP]; PMBC Order Batrachoidiformes Batrachoididae Allenbatrachus grunniens (Linnaeus, 1758): Mg, OShr; [NIO-IM]; PMBC Order Lophiiformes Lophiidae Lophiodes lugubris (Alcock, 1894): OShr; [EIO] Lophiodes mutilus (Alcock, 1893): OShr; [IWP]; PMBC Antennariidae Antennarius coccineus (Cuvier, 1831): Cr; [IP] Antennarius commerson (Latreille, 1804): Cr, OShr; [IP]; PMBC Antennarius hispidus (Bloch & Schneider, 1801): Cr, OShr; [IWP] Antennarius maculatus (Desjardins, 1840): Cr; [IP] Antennarius nummifer (Cuvier, 1817): Cr; [IP]; PMBC Antennarius pictus (Shaw & Nodder, 1794): Cr; [IP]; PMBC Chaunacidae Chaunax pictus Lowe, 1846: OShr; [COS] Ogcocephalidae Halieutaea coccinea Alcock, 1889: OShr; [IO] Halieutaea indica Annandale & Jenkins, 1910: OShr; [IWP] Halieutaea stellata (Vahl, 1797): OShr; [WP]; PMBC Halieutopsis micropa (Alcock, 1891): OShr; [IWP] Order Mugiliformes Mugilidae Chelon macrolepis (Smith, 1846): Mg; [IP]; PMBC Chelon parsia (Hamilton, 1822): Mg; [EIO] Chelon subviridis (Valenciennes, 1836): Mg, Sg; [IWP]; PMBC Crenimugil crenilabris (Forsskål, 1775): Cr; [IP] Ellochelon vaigiensis (Quoy & Gaimard, 1825): Cr, Mg, Sg; [IP]; PMBC Moolgarda cunnesius (Valenciennes, 1836): Mg, Sg; [IWP]; PMBC Moolgarda engeli (Bleeker, 1859): Mg; [IP] 52 Phuket mar. biol. Cent. Res. Bull. Moolgarda pedaraki (Valenciennes, 1836): Mg, Sg; [IWP]; PMBC Moolgarda perusii (Valenciennes, 1836): Mg, Sg; [IWP]; PMBC Moolgarda seheli (Forsskål, 1775): Cr, Mg; [IP]; PMBC Oedalechilus labiosus (Valenciennes, 1836): Cr; [IP]; PMBC Paramugil parmatus (Cantor, 1849): Mg; [IM]; PMBC Order Atheriniformes Phallostethidae Neostethus bicornis Regan, 1916: Mg; [IM] Neostethus lankesteri Regan, 1916: Mg; [IM]; PMBC Phenacostethus posthon Roberts, 1971: Mg; [AS] Atherinidae Atherinomorus duodecimalis (Valenciennes, 1835): Cr, Mg, Sg, OShr; [IP]; PMBC Atherinomorus lacunosus (Forster & Schneider, 1801): OShr; [EIO-WP]; PMBC Atherinomorus pinguis (Lacepède, 1803): Sg, OShr; [IWP] Atherion elymus Jordan & Starks, 1901: Cr, OShr; [WP]; PMBC Hypoatherina temminckii (Bleeker, 1853): Cr, OShr [IP]; PMBC Hypoatherina valenciennei (Bleeker, 1853): Mg, Sg, OShr; [WP]; PMBC Order Beloniformes Andrianichthyidae Oryzias javanicus (Bleeker, 1854): Mg; [IM]; PMBC Exocoetidae Cheilopogon arcticeps (Günther, 1866): OShr; [WP] Cheilopogon spilopterus (Valenciennes, 1846): OShr; [WP]; PMBC Exocoetus volitans (Linnaeus, 1758): OShr; [COS]; PMBC Hirundichthys speculiger (Valenciennes, 1847): OShr; [CIR] Parexocoetus brachypterus (Richardson, 1846): OShr; [IP]; PMBC Hemiramphidae Dermogenys siamensis Fowler, 1934: Mg; [IM] Hemiramphus far (Forsskål, 1775): Cr, Sg, OShr; [IWP]; PMBC Hyporhamphus balinensis (Bleeker, 1858): OShr; [IWP] Hyporhamphus dussumieri (Valenciennes, 1846): Sg, OShr; [IP]; PMBC Hyporhamphus limbatus (Valenciennes, 1846): Mg, Sg, OShr; [NIO-WP] Hyporhamphus melanopterus Collette & Parin, 1978: Mg, OShr; [IM]; PMBC Hyporhamphus quoyi Valenciennes, 1847: Mg, Sg, OShr; [IWP]; PMBC Hyporhamphus unicuspis Collette & Parin, 1978: OShr; [NIO] Zenarchopterus buffornis (Valenciennes, 1847): Mg, Sg, OShr; [EIO-IA]; PMBC Zenarchopterus dispar (Valenciennes, 1847): Mg, Sg, OShr; [IWP]; PMBC Zenarchopterus dunckeri Mohr, 1926: Mg, Sg, OShr; [WP]; PMBC Belonidae Ablennes hians (Valenciennes, 1846): Cr, OShr; [COS]; PMBC Strongylura incisa (Valenciennes, 1846): OShr; [EIO-WP]; PMBC Strongylura leiura (Bleeker, 1850): OShr; [IWP]; PMBC Strongylura strongylura (van Hasselt, 1823): Mg, Sg, OShr; [NIO-IA]; PMBC Tylosurus acus melanotus (Bleeker, 1850): Cr, OShr; [IP] Tylosurus crocodilus crocodilus (Peron & Lesueur, 1821): Cr, Mg, Sg, OShr; [CIR] Order Cyprinodontiformes Aplocheilidae Aplocheilus panchax Day, 1875: Mg; [EIO-IM]; PMBC Order Beryciformes Diretmidae Diretmoides veriginae Kotlyar, 1987: OShr; [EIO-WP]; PMBC Monocentridae Monocentris japonica (Houttuyn, 1782): OShr; [IWP]; PMBC Trachichthyidae Gephyroberyx darwinii (Johnson, 1866): OShr; [COS]; PMBC Berycidae Beryx splendens Lowe, 1834: OShr; [COS] Centroberyx druzhinini (Busakhin, 1981): OShr; [IWP]; PMBC Holocentridae Myripristis adustus (Bleeker, 1853): Cr, OShr; [IP] Myripristis berndti (Jordan & Evermann, 1903): Cr, OShr; [IP]; PMBC Myripristis botche Cuvier, 1829: Cr, OShr; [IWP]; PMBC Myripristis chryseres Jordan & Evermann, 1903: Cr, OShr; [IP] Myripristis hexagona (Lacepède, 1802): Cr, OShr; [IWP]; PMBC Myripristis kuntee Cuvier, 1831: Cr, [IP] Myripristis murdjan (Forsskål, 1775): Cr; [IP] Myripristis violacea Bleeker, 1851: Cr; [IP] Myripristis vittatus Valenciennes, 1831: Cr; [IP] Neoniphon sammara (Forsskål, 1775): Cr, OShr; [IP]; PMBC Ostichthys japonicus (Cuvier, 1829): OShr; [WP]; PMBC Sargocentron caudimaculatum (Rüppell, 1835): Cr; [IP]; PMBC Sargocentron cornutum (Bleeker, 1853): Cr; [WP]; PMBC Sargocentron diadema (Lacepède, 1802): Cr; [IP] Sargocentron melanospilos (Bleeker, 1858): Cr; [IWP] Sargocentron rubrum (Forsskål, 1775): Cr, OShr; [IWP]; PMBC Sargocentron spiniferum (Forsskål, 1775): Cr; [IP] Order Zeiformes Grammicolepididae Xenolepidichthys dalgleishi Gilchrist, 1922: OShr; [CIR] Zeidae Cyttopsis rosea (Lowe, 1843): OShr; [COS]; PMBC 53 The fishes of southwestern Thailand Zenopsis nebulosa (Temminck & Schlegel, 1845): OShr; [IP] Order Gasterosteiformes Pegasidae Eurypegasus draconis (Linnaeus, 1766): Cr, OShr; [IP]; PMBC Pegasus volitans Linnaeus, 1758: Cr, Sg, OShr; [IWP]; PMBC Solenostomidae Solenostomus cyanopterus Bleeker, 1852: Cr; [IP] Solenostomus paradoxus (Pallas, 1770): Cr, Sg; [IWP]; PMBC Syngnathidae Bhanotia fasciolata (Dumeril, 1870): Cr, OShr; [IA]; PMBC Choeroichthys brachysoma (Bleeker, 1855): Cr; [IP]; PMBC Choeroichthys sculptus (Günther, 1870): Cr, OShr; [IP] Corythoichthys amplexus Dowson & Randall, 1906: Cr, OShr; [IWP] Corythoichthys flavofasciatus (Rüppell, 1838): Cr; [IP] Corythoichthys haematopterus (Bleeker, 1851): Cr, OShr; [IWP]; PMBC Corythoichthys schultzi Herald, 1953: Cr; [IP]; PMBC Doryrhamphus janssi (Herald & Randall, 1972): Cr, OShr; [WP]; PMBC Dunckerocampus multiannulatus (Regan, 1903): Cr; [IO] Halicampus grayi Kaup, 1856: OShr; [NIO-WP]; PMBC Halicampus mataafae (Jordan & Seale, 1906): Cr, OShr; [IP] Halicampus spinirostris (Dawson & Allen, 1981): Cr; [EIO-WP] Hippichthys cyanospilus (Bleeker, 1854): Sg, OShr; [IWP]; PMBC Hippichthys heptagonus (Bleeker, 1853): Sg, OShr; [IWP]; PMBC Hippichthys penicillus (Cantor, 1849): Mg, Sg, OShr; [NIOWP]; PMBC Hippichthys spicifer (Rüppell, 1840): Mg, OShr; [IP]; PMBC Hippocampus comes Cantor, 1850: Cr, Sg; [IM]; PMBC Hippocampus kelloggi Jordan & Snyder, 1901: OShr; [IWP]; PMBC Hippocampus kuda Bleeker, 1852: Mg, Sg, OShr;[NIOWCP]; PMBC Hippocampus spinosissimus Weber, 1913: OShr; [EIO-IA]; PMBC Hippocampus trimaculatus Leach, 1814: OShr; [EIO-WP]; PMBC Phoxocampus belcheri (Kaup, 1856): Cr, OShr; [IWP] Syngnathoides biaculeatus (Bloch, 1785): Sg, OShr; [IWP]; PMBC Trachyrhamphus bicoarctatus (Bleeker, 1857): Cr, Sg, OShr; [IWP]; PMBC Trachyrhamphus longirostris Kaup, 1856: OShr; [IWP]; PMBC Trachyrhamphus serratus (Temminck & Schlegel, 1850): OShr; [EIO-WP]; PMBC Aulostomidae Aulostomus chinensis (Linnaeus, 1766): Cr; [IP]; PMBC Fistulariidae Fistularia commersonii Rüppell, 1835: Cr, Sg; [IP]; PMBC Fistularia petimba Lacepède, 1803: OShr; [CIR]; PMBC Centriscidae Aeoliscus strigatus (Günther, 1860): Cr, Sg, OShr; [IWP] Centriscus cristatus De Vis, 1885: OShr; [WP] Centriscus scutatus Linnaeus, 1758: Cr, Sg, OShr; [IWP]; PMBC Order Sybranchiformes Sybranchidae Macrotrema caligans (Cantor, 1849): Mg; [IM] Order Scorpaeniformes Dactylopteridae Dactyloptena macracantha (Bleeker, 1854): OShr; [EIOWP]; PMBC Dactyloptena orientalis (Cuvier, 1829): Cr, OShr; [IP]; PMBC Dactyloptena perterseni (Nystrom, 1887): OShr; [IWP]; PMBC Scorpaenidae Ablabys taenianotus (Cuvier, 1829): Cr; [EIO-WP] Apistus carinatus (Bloch & Schneider, 1801): OShr; [IWP] Choridactylus multibarbus Richardson, 1848: OShr; [IWP] Dendrochirus biocellatus (Fowler, 1938): Cr; [IP] Dendrochirus brachypterus (Cuvier, 1829): Cr, OShr; [IP]; PMBC Dendrochirus zebra (Cuvier, 1829): Cr, Sg, OShr; [IWP]; PMBC Inimicus cuvieri (Gray, 1832): Cr, OShr; [IM] Lioscorpius longiceps Günther, 1880: OShr; [WP] Minous coccineus Alcock, 1890: OShr; [IWP] Minous inermis Alcock, 1889: OShr; [IO] Minous monodactylus (Bloch & Schneider, 1801): OShr; [IWP]; PMBC Minous trachycephalus (Bleeker, 1854): OShr; [EIO-WP] Neomerinthe amplisquamiceps (Fowler, 1938): OShr; [WP] Paracentropogon longispinis (Bleeker, 1829): Cr, OShr; [EIO-WP] Parapterois heterurus (Bleeker, 1856): OShr; [IWP] Parascorpaena aurita (Rüppell, 1838): Cr, OShr; [IWP]; PMBC Parascorpaena mossambica (Peters, 1855): Cr; [IP] Pontinus macrocephalus (Sauvage, 1882): OShr; [WP]; PMBC Pterois antennata (Bloch, 1787): Cr, OShr; [IP]; PMBC Pterois miles (Bennett, 1828): Cr, Sg, OShr; [IO]; PMBC Pterois mombasae (Smith, 1957): Cr, OShr; [IO] Pterois radiata Cuvier, 1829: Cr, OShr; [IP]; PMBC Pterois russellii Bennett, 1831: Sg, OShr; [IWP]; PMBC Rhinopias eschmeyeri Conde, 1977: Cr; [IWP] Rhinopias frondosa (Günther, 1891): Cr, OShr; [IWP] Richardsonichthys leucogaster (Richardson, 1848): OShr; [IWP] 54 Phuket mar. biol. Cent. Res. Bull. Scorpaenodes albaiensis (Evermann & Seale, 1907): Cr; [IWP]; PMBC Scorpaenodes guamensis (Quoy & Gaimard, 1824): Cr, OShr; [IP]; PMBC Scorpaenodes hirsutus (Smith, 1957): Cr; [IP] Scorpaenodes minor (Smith, 1958): Cr; [IP] Scorpaenodes parvipinnis (Garrett, 1863): Cr, OShr; [IP]; PMBC Scorpaenodes scaber (Ramsay & Ogilby, 1886): Cr; [EIOWCP] Scorpaenodes varipinnis Smith, 1957: Cr; [IWP] Scorpaenopsis diabolus Cuvier, 1829: Cr, OShr; [IP]; PMBC Scorpaenopsis longispina Randall & Eschmeyer, 2001: Cr; [IO] Scorpaenopsis neglecta Heckel, 1837: Sg, OShr; [EIO-WP]; PMBC Scorpaenopsis oxycephala (Bleeker, 1849): Cr, OShr; [IWP]; PMBC Scorpaenopsis possi Randall & Eschmeyer, 2001: Cr; [IP] Scorpaenopsis ramaraoi Randall & Eschmeyer, 2001: Cr, OShr; [NIO-WP] Scorpaenopsis vesosa (Cuvier, 1829): OShr; [IWP] Sebastapistes fowleri (Pietschmann, 1934): Cr; [IP] Sebastapistes mauritiana (Cuvier, 1829): Cr, OShr; [IP] Sebastapistes strongia (Cuvier, 1829): Cr; [IP] Setarches guentheri Johnson, 1862: OShr; [CIR] Synanceia horrida (Linnaeus, 1766): Cr, OShr; [EIO-WP]; PMBC Synanceia verrucosa (Smith, 1956): Cr, OShr; [IP] Taenianotus triacanthus Lacepède, 1802: Cr; [IP] Trachicephalus uranoscopus (Bloch & Schneider, 1801): Mg, Sg, OShr; [EIO-WP], PMBC Vespicula trachinoides (Cuvier & Valenciennes, 1829): Mg, Sg, OShr; [WP]; PMBC Caracanthidae Caracanthus unipinna (Gray, 1831): Cr; [IWP]; PMBC Aploactinidae Acanthosphex leurynnis (Jordan & Seale, 1905): Mg, Sg, OShr; [EIO-IM]; PMBC Cocotropus echinatus (Cantor, 1850): Mg, Sg; [AS] Cocotropus larvatus Poss & Allen, 1987: OShr; [WP] Paraploactis obbesi (Weber, 1913): Cr, OShr; [WP] Sthenopus mollis Richardson, 1848: OShr; [WP] Xenaploactis cautes Poss & Eschmeyer, 1980: OShr; [IM (presently known from AS and Gulf of Thailand)] Triglidae Lepidotrigla punctipectoralis Fowler, 1938: OShr; [IWP] Lepidotrigla spiloptera Günther, 1880: OShr; [IWP]; PMBC Peristediidae Peristedion liorhynchus Günther, 1871: OShr; [IWP]; PMBC Peristedion moluccense Bleeker, 1850: OShr; [WP]; PMBC Satyrichthys adeni (Lloyd, 1907): OShr; [IWP]; PMBC Satyrichthys sp.: OShr; PMBC Bembridae Brachybembras sp.: OShr Plathycephalidae Cociella punctata (Cuvier, 1829): Cr, Mg, Sg, OShr; [IWP]; PMBC Elates ransonneti (Steindachner, 1877): OShr; [IA]; PMBC Grammoplites scaber (Linnaeus, 1758): Mg, Sg, OShr; [NIO-IM]; PMBC Inegocia japonica (Tilesius, 1812): Mg, Sg, OShr; [EIOWP]; PMBC Kumococius rodericensis Cuvier in Cuvier & Valenciennes, 1829: OShr; [NIO-IA]; PMBC Platycephalus indicus (Linnaeus, 1758): Mg, Sg, OShr; [IWP]; PMBC] Rogadias pristiger (Cuvier, 1829): OShr; [IWP]; PMBC Rogadias serratus (Cuvier, 1829): OShr; [IWP]; PMBC Sorsogona melanoptera (Knapp & Wongratana, 1987); OShr; [NIO] Sorsogona tuberculata (Cuvier, 1829): OShr; [NIO-IA]; PMBC Suggrundus macracanthus (Bleeker, 1869): OShr; [EIOIA]; PMBC Sunagocia carbunculus (Valenciennes, 1833): Mg, Sg, OShr; [EIO-IM]; PMBC Sunagocia otaitensis (Cuvier, 1829): Cr, OShr; [IP]; PMBC Thrysanophrys celebicus (Bleeker, 1854): Cr; [IWP] Thrysanophrys chitonae Schultz, 1966: Cr, OShr; [IP] Hoplichthyidae Hoplichthys citrinus Gilbert, 1905: OShr; [WP] Order Perciformes Ambassidae Ambassis buruensis Bleeker, 1857: Mg; [WP]; PMBC Ambassis dussumieri Cuveir, 1828: Sg; [IWP] Ambassis gymnocephalus (Lacepède, 1802): Mg; [IWP]; PMBC Ambassis interruptus Bleeker, 1852: Mg, Sg; [IA]; PMBC Ambassis macracanthus Bleeker, 1849: Mg, Sg; [EIO-IM]; PMBC Ambassis nalua (Hamilton, 1822): Mg, Sg; [EIO-IA]; PMBC Ambassis urotaenia Bleeker, 1852: Mg, Sg; [IWP]; PMBC Ambassis vachellii Richardson, 1846: Mg, Sg; [IWP]; PMBC Latidae Lates calcalifer (Bloch, 1790): Mg; [NIO-WP]; PMBC Acropomatidae Acropoma argentistigma Okamoto & Ida, 2002: OShr; [AS] Synagrops japonicus (Döderlein, 1883): OShr; [IP]; PMBC Synagrops malayanus Weber, 1913: OShr; [WCP] Synagrops philippinensis (Günther, 1880): OShr; [EIOWP]; PMBC Serranidae Aethaloperca rogaa (Forsskål, 1775): Cr, OShr; [IP]; PMBC Anyperodon leucogrammicus (Valnciennes, 1828): Cr, OShr; [IP]; PMBC 55 The fishes of southwestern Thailand Cephalopholis argus Bloch & Schneider, 1801: Cr, OShr; [IP]; PMBC Cephalopholis aurantia (Valenciennes, 1828): OShr; [IWP] Cephalopholis boenak (Bloch, 1790): Cr, Sg, OShr; [IWP]; PMBC Cephalopholis formosa (Shaw & Nodder, 1812): Cr, OShr; [EIO-WP]; PMBC Cephalopholis leopardus (Lacepède, 1801): Cr; [IP] Cephalopholis miniata (Forsskål, 1775): Cr, OShr; [IP]; PMBC Cephalopholis polyspila Randall & Satapoomin, 2000: Cr, OShr; [AS]; PMBC Cephalopholis sexmaculata (Rüppell, 1830): Cr; [IP] Cephalopholis sonnerati (Valenciennes, 1828): Cr, OShr; [IP]; PMBC Cephalopholis urodeta (Bloch & Schneider, 1801): Cr, OShr; [IP]; PMBC Chelidoperca margaritifera Weber, 1913: OShr; [WP]; PMBC Cromileptes altivelis (Valenciennes, 1828): Cr, OShr; [WP]; PMBC Diploprion bifasciatum Cuvier, 1828: Cr, OShr; [EIO-WP]; PMBC Epinephelus amblycephalus (Bleeker, 1857): OShr; [WP]; PMBC Epinephelus areolatus (Forsskål, 1775): Cr, OShr; [IWP]; PMBC Epinephelus bleekeri (Vaillant, 1877): Cr, Mg, Sg, OShr; [NIO-IA]; PMBC Epinephelus caeruleopunctatus (Bloch, 1790): Cr, OShr; [IWP]; PMBC Epinephelus coioides (Hamilton, 1822): Cr, Mg, Sg, OShr: [IWP]; PMBC Epinephelus epistictus (Temminck & Schlegel, 1842): OShr; [IWP]; PMBC Epinephelus erythrurus (Valenciennes, 1828): Cr, Sg, OShr; [NIO-IM]; PMBC Epinephelus fasciatus (Forsskål, 1775): Cr, OShr; [IP]; PMBC Epinephelus flavocaeruleus (Lacepède, 1801): Cr, OShr; [IO] Epinephelus fuscoguttatus (Forsskål, 1775): Cr, OShr; [IP]; PMBC Epinephelus heniochus Fowler, 1904: OShr; [WP] Epinephelus hexagonatus (Bloch & Schneider, 1801): Cr; [IP] Epinephelus lanceolatus (Bloch, 1790): Cr, OShr; [IP]; PMBC Epinephelus latifasciatus (Temminck & Schlegel, 1842): OShr; [IWP] Epinephelus longispinis (Kner, 1864): Cr, OShr; [IO] Epinephelus macrospilos (Bleeker, 1855): Cr; [IP]; PMBC Epinephelus malabaricus (Bloch & Schneider, 1801): Cr, Mg, OShr; [IWP]; PMBC Epinephelus merra Bloch, 1793: Cr, OShr; [IP]; PMBC Epinephelus morrhua (Valenciennes, 1833): OShr; [IP]; PMBC Epinephelus octofasciatus Griffin, 1926: OShr; [IP]; PMBC Epinephelus ongus (Bloch, 1790): Cr, OShr; [IWP]; PMBC Epinephelus polyphekadion (Bleeker, 1849): Cr, OShr; [IP]; PMBC Epinephelus quoyanus (Valenciennes, 1830): Cr, Sg, OShr; [WP]; PMBC Epinephelus radiatus (Day, 1867): OShr; [IWP]; PMBC Epinephelus sexfasciatus (Valenciennes, 1828): Mg, Sg, OShr; [IA]; PMBC Epinephelus spilotoceps Schultz, 1953: Cr; [IP] Epinephelus tauvina (Forsskål, 1775): Cr, OShr; [IP]; PMBC Epinephelus tukula Morgan, 1959: Cr, OShr; [IWP] Epinephelus undulosus (Quoy & Gaimard, 1824): Cr, OShr; [IWP]; PMBC Grammistes sexlineatus (Thunberg, 1792): Cr; [IP] Grammistops ocellatus Schultz, 1953: Cr; [IP] Liopropoma africanum (Smith, 1954): Cr; [IO]; PMBC Liopropoma susumi (Jordan & Seale, 1906): Cr; [IP]; PMBC Meganthias filiferus Randall & Heemstra, 2007: OShr; [AS]; PMBC Plectropomus areolatus (Rüppell, 1830): Cr, OShr; [IP] Plectropomus laevis (Lacepède, 1801): Cr, OShr; [IP] Plectropomus pessuliferus Fowler, 1901: Cr, OShr; [IWP] Pogonoperca ocellata Günther, 1859: Cr, OShr; [IO]; PMBC Pseudanthias evansi (Smith, 1954): Cr; [IO]; PMBC Pseudanthias ignitus (Randall & Lubbock, 1981): Cr; [EIO] Pseudanthias rubrizonatus (Randall, 1983): Cr; [WP] Pseudanthias squamipinnis (Peters, 1855): Cr; [IWP]; PMBC Pseudogramma polyacanthum (Bleeker, 1856): Cr; [IP]; PMBC Rabaulichthys stigmaticus Randall & Pyle, 1989: Cr, OShr; [IO] Variola albimarginata Baissac, 1953: Cr, OShr; [IWP] Variola louti (Forsskål, 1775): Cr, OShr; [IP]; PMBC Ostracoberycidae Ostracoberyx dorygenys Fowler, 1934: OShr; [IWP] Pseudochromidae Amsichthys knighti (Allen, 1987): Cr; [WP] Blennodesmus scapularis Günther, 1871: Cr; [IA]; PMBC Pseudochromis andamanensis Lubbock, 1980: Cr; [EEIO]; PMBC Pseudochromis caudalis Boulenger, 1898: Cr; [IO] Pseudochromis coccinicauda Tickell in Day, 1888: Cr; [EIO] Pseudochromis fuscus Müller & Troschel, 1849: Cr; [EIOIA]; PMBC Pseudochromis tapeinosoma Bleeker, 1853: Cr; [IA]; PMBC Pseudoplesiops rosae Schultz, 1953: Cr; [EIO-WCP]; PMBC Plesiopidae Calloplesiops altivelis (Steindachner, 1903); Cr; [IP] Plesiops auritus Mooi, 1995: Cr; [EIO] 56 Phuket mar. biol. Cent. Res. Bull. Plesiops caeruleolineatus Rüppell, 1835: Cr; [IP]; PMBC Plesiops corallicola Bleeker, 1853: Cr; [WCP]; PMBC Opistognathidae Opistognathus cyanospilotus Smith-Vaniz, 2009: Cr; [EEIO] Opistognathus nigromarginatus Rüppell, 1830: Cr, OShr; [IWP]; PMBC Opistognathus rosenbergii (Bleeker, 1856): Cr, OShr; [EIO-WP]; PMBC Opistognathus variabilis Smith-Vaniz, 2009: Cr; [EIO-WP]; PMBC Priacanthidae Heteropriacanthus cruentatus (Lacepède, 1801): Cr, OShr; [COS]; PMBC Priacanthus blochii Bleeker, 1853: OShr; [IWP]; PMBC Priacanthus hamrur (Forsskål, 1775): Cr, OShr; [IP]; PMBC Priacanthus macracanthus Cuvier, 1829: Cr, OShr; [WP]; PMBC Priacanthus sagittarius Starnes, 1988: OShr; [IWP] Priacanthus tayenus Richardson, 1846: OShr; [NIO-WP]; PMBC Pristigenys niphonia (Cuvier, 1829): OShr; [IWP] Apogonidae Apogon abrogramma Fraser & Lachner, 1985: Cr; [IWP] Apogon angustatus (Smith & Radcliffe, 1911): Cr; [IP]; PMBC Apogon apogonides (Bleeker, 1856): Cr; [IWP] Apogon aureus (Lacepède, 1802): Cr, OShr; [IWP] Apogon bryx Fraser, 1998: OShr; [IWP] Apogon campbelli Smith, 1949: Cr; [IO] Apogon compressus (Smith & Radcliffe, 1911): Cr; [WP]; PMBC Apogon cookii Macleay, 1881: Cr; [IWP]; PMBC Apogon cyanosoma Bleeker, 1853: Cr; [IWP]; PMBC Apogon endekataenia Bleeker, 1852: Cr; [IA]; PMBC Apogon fasciatus (White, 1790): OShr; [IWP]; PMBC Apogon fleurieu (Lacepède, 1802): Cr, OShr; [IWP] Apogon frenatus Valenciennes, 1832: Cr; [IP]; PMBC Apogon hyalosoma Bleeker, 1852: Mg, Sg; [WP]; PMBC Apogon kallopterus Bleeker, 1856: Cr; [IP]; PMBC Apogon kalosoma (Bleeker, 1852): Cr; [EIO-WP] Apogon lateralis Valenciennes, 1832: Cr, Mg, Sg; [IWP]; PMBC Apogon moluccensis Valenciennes, 1832: Cr, Sg; [IA]; PMBC Apogon nigrofasciatus Lachner, 1953: Cr; [IP]; PMBC Apogon novaeguinae Valenciennes, 1832: Mg, Sg; [IWP]; PMBC Apogon ocellicaudus Allen, Kuiter & Randall, 1994: Cr; [IA] Apogon pleuron Fraser, 2005: Sg, OShr; [EIO-WP]; PMBC Apogon poecilopterus Cuvier, 1828: OShr; [EIO-WP] Apogon semiornatus Peters, 1876: Cr; [IWP] Apogon smithi (Kotthaus, 1970): OShr; [IWP]; PMBC Apogon sp.1: Cr; [AS] Apogon striatodes Gon, 1996: OShr; [IM]; PMBC Apogon striatus (Smith & Radcliffe, 1912): OShr; [NIOWP]; PMBC Apogon taeniophorus Regan, 1908: Cr; [IP]; PMBC Apogon talboti Smith, 1961: Cr; [IWP] Apogon thermalis Cuvier, 1829: Cr, Sg; [IWP]; PMBC Apogon trimaculatus Cuvier, 1828: Cr, OShr; [WP]; PMBC Apogon truncatus Bleeker, 1854: OShr; [IWP]; PMBC Apogonichthyoides nigripinnis (Cuvier, 1828): OShr; [EIOIA]; PMBC Apogonichthyoides sialis (Jordan & Thompson, 1914): Cr; [EIO-WP] Apogonichthys sp.: OShr Archamia ataenia Randall & Satapoomin, 1999: Cr; [AS]; PMBC Archamia bleekeri Günther, 1859: Cr; [IWP] Archamia fucata (Cantor, 1850): Cr, Sg, OShr; [IP]; PMBC Archamia macroptera (Cuvier, 1828): Cr; [EIO-WP] Cheilodipterus artus Smith, 1961: Cr; [IP]; PMBC Cheilodipterus macrodon (Lacepède, 1802): Cr; [IP]; PMBC Cheilodipterus quinquelineatus Cuvier, 1828: Cr; [IP]; PMBC Foa fo (Jordan & Seale, 1906): Sg, OShr; [IWP]; PMBC Fowleria aurita (Valenciennes, 1831): Cr; [IWP]; PMBC Fowleria isostigma (Jordan & Seale, 1960): Cr; [IWP]; PMBC Fowleria vaiulae (Jordan & Seale, 1906): Cr; [IWP]; PMBC Fowleria variegata (Valenciennes, 1832): Cr, Mg, Sg; [IWP]; PMBC Gymnapogon urospilotus Lachner, 1953: Cr; [WCP] Neamia octospina Smith & Radcliffe, 1912: Cr; [IWP] Nectamia fusca (Quoy & Gaimard, 1824): Cr; [IWP]; PMBC Nectamia luxuria Fraser, 2008: Cr; [EIO-WP]; PMBC Nectamia savayensis (Günther, 1872): Cr; [IP] Pseudamia hayashii Randall, Lachner, & Fraser, 1985: Cr; [IP] Pseudamia sp.1: Cr Rhabdamia cypselurus Weber, 1909: Cr; [IWP]; PMBC Rhabdamia gracilis (Bleeker, 1856): Cr; [IWP]; PMBC Siphamia tubifer Weber, 1909: Cr; [WP] Zoramia leptacantha (Bleeker, 1856): Cr; [IWP]; PMBC Zoramia perlita (Fraser & Lachner, 1985): Cr; [IM] Sillaginidae Sillaginopsis panijus (Hamilton, 1822): OShr; [EIO]; PMBC Sillago aeolus Jordan & Evermann, 1902: Cr, Mg, Sg, OShr; [IWP]; PMBC Sillago chondropus Bleeker, 1849: OShr; [IWP]; PMBC Sillago intermedius Wongratana, 1977: OShr; [EIO-IM] Sillago lutea McKay, 1985: OShr; [EIO-IA] Sillago sihama (Forsskal, 1775): Mg, Sg, OShr; [IWP]; PMBC Malacanthidae 57 The fishes of southwestern Thailand Branchiostegus sawakinensis Amirthalingam, 1969: OShr; [IWP]; PMBC Hoplolatilus cuniculus Randall & Dooley, 1974: Cr; [IP] Hoplolatilus luteus Allen & Kuiter, 1989: Cr, OShr; [EEIO]; PMBC Malacanthus brevirostris Guichenot, 1848: Cr, OShr; [IP]; PMBC Malacanthus latovittatus (Lacepède, 1801): Cr; [IP] Lactariidae Lactarius lactarius (Bloch & Schneider, 1801): OShr; [NIO-WP]; PMBC Coryphaenidae Coryphaena equiselis Linnaeus, 1758: OShr; [COS]; PMBC Coryphaena hippurus Linnaeus, 1758: OShr; [COS]; PMBC Rachycentridae Rachycentron canadum (Linnaeus, 1766): Cr, Sg, OShr; [CIR]; PMBC Echeneidae Echeneis naucrates Linnaeus, 1758: Cr, OShr; [COS]; PMBC Phtheirichthys lineatus (Menzies, 1791): OShr; [COS] Remora remora (Linnaeus, 1758): OShr; [COS] Carangidae Alectis ciliaris (Bloch, 1788): Cr, OShr; [CIR]; PMBC Alectis indica (Rüppell, 1830): Cr, Sg, OShr; [IWP]; PMBC Alepes djedaba (Forsskål, 1775): Mg, OShr; [IWP]; PMBC Alepes kleinii (Bloch, 1793): Mg, Sg, OShr; [NIO-WP]; PMBC Alepes melanoptera (Swainson, 1839): OShr; [NIO-IM]; PMBC Alepes vari (Cuvier, 1833): Cr, Mg, OShr; [IWP] Atropus atropus (Schneider, 1801): OShr; [NIO-WP] Atule mate (Cuvier, 1833): Cr, Mg, Sg, OShr; [IP]; PMBC Carangoides armatus (Rüppell, 1830): Mg, Sg, OShr; [IWP]; PMBC Carangoides caeruleopinnatus (Rüppell, 1830): Cr, Sg, OShr; [IWP]; PMBC Carangoides chrysophrys (Cuvier, 1833): OShr; [IWP] Carangoides equula (Temminck & Schlegel, 1844): Cr, OShr; [IP] Carangoides ferdua (Forsskål, 1775): Cr, OShr; [IP]; PMBC Carangoides fulvoguttatus (Forsskål, 1775): OShr; [IWP]; PMBC Carangoides gymnostethus (Cuvier, 1833): OShr; [IWP]; PMBC Carangoides hedlandensis (Whitley, 1933): Cr, OShr; [IWP]; PMBC Carangoides malabaricus (Bloch & Schneider, 1801): OShr; [IWP]; PMBC Carangoides oblongus (Cuvier, 1833): OShr; [IWP]; PMBC Carangoides orthogrammus (Jordan & Gilbert, 1882): Cr, OShr; [IP] Carangoides plagiotaenia Bleeker, 1857: Cr, OShr; [IP]; PMBC Carangoides praeustus (Bennett, 1830): Mg, Sg, OShr; [NIO-IM]; PMBC Carangoides talamparoides Bleeker, 1852: OShr; [NIOIA] Caranx heberi (Bennett, 1830): Cr, OShr; [NIO-WP] Caranx ignobilis (Forsskål, 1775): Cr, Mg, Sg, OShr; [IP]; PMBC Caranx melampygus Cuvier, 1833: Cr, OShr; [IP]; PMBC Caranx papuensis Alleyne & Macleay, 1877: Cr, OShr; [IP] Caranx sexfasciatus Quoy & Gaimard, 1824: Cr, Mg, Sg, OShr; [IP]; PMBC Decapterus kurroides Bleeker, 1855: OShr; [IWP]; PMBC Decapterus macarellus (Cuvier, 1833): OShr; [CIR]; PMBC Decapterus macrosoma Bleeker, 1851: Cr, OShr; [IP]; PMBC Decapterus maruadsi (Temminck & Schlegel, 1843): OShr; [WP]; PMBC Decapterus russelli (Rüppell, 1830): Cr, OShr; [IWP]; PMBC Elagatis bipinnulata (Quoy & Gaimard, 1825): Cr, OShr; [CIR]; PMBC Gnathanodon speciosus (Forsskål, 1775): Cr, OShr; [IP]; PMBC Megalaspis cordyla (Linnaeus, 1758): Cr, OShr; [IP]; PMBC Naucrates ductor (Linnaeus, 1758): OShr; [CIR]; PMBC Parastromateus niger (Bloch, 1795): OShr; [IWP]; PMBC Scomberoides commersonianus Lacepède, 1801: Cr, Mg, Sg, OShr; [IWP]; PMBC Scomberoides lysan (Forsskål, 1775): Cr, Mg, Sg, OShr; [IP]; PMBC Scomberoides tala (Cuvier, 1832): OShr; [EIO-WP]; PMBC Scomberoides tol (Cuvier, 1832): Cr, Mg, Sg, OShr; [IWP]; PMBC Selar boops (Cuvier, 1833): Cr, OShr; [IWP]; PMBC Selar crumenophthalmus (Bloch, 1793): Cr, OShr; [CIR]; PMBC Selaroides leptolepis (Cuvier, 1833): Cr, Sg, OShr; [NIOWP]; PMBC Seriola rivoliana Valenciennes, 1833: OShr; [CIR]; PMBC Seriolina nigrofasciata (Rüppell, 1829): Cr, OShr; [IWP]; PMBC Trachinotus bailloni (Lacepède, 1801): Cr, OShr; [IP]; PMBC Trachinotus blochii (Lacepède, 1801): Cr, OShr; [IP] Trachinotus botla (Shaw, 1803): OShr; [IO]; PMBC Ulua mentalis (Cuvier, 1833): Cr, OShr; [IWP]; PMBC Uraspis helvola (Forster, 1801): OShr; [IP]; PMBC Uraspis uraspis (Günther, 1860): OShr; [IWP]; PMBC Menidae Mene maculata (Bloch & Schneider, 1801): OShr; [IWP]; PMBC Leiognathidae Equulites berbis (Valenciennes, 1835): OShr; [IWP]; PMBC 58 Phuket mar. biol. Cent. Res. Bull. Equulites elongatus (Günther, 1860): OShr; [IWP]; PMBC Equulites leuciscus (Günther, 1860): Mg, Sg, OShr; [IWP]; PMBC Equulites oblongus (Valenciennes, 1835): Mg, OShr; [IWP]; PMBC Equulites stercorarius (Everman & Seale, 1907): Mg, Sg, OShr; [WP]; PMBC Eubleekeria jonesi (James, 1971): Mg, Sg, OShr; [EIOWP]; PMBC Gazza dentex (Valenciennes, 1835): OShr; [IO]; PMBC Gazza minuta (Bloch, 1797): Mg, Sg, OShr; [IWP]; PMBC Gazza rhombea Kimura, Yamashita & Iwatsuki, 2000: OShr; [EIO-WP]; PMBC Karalla daura (Cuvier, 1829): Mg, OShr; [IWP]; PMBC Leiognathus equulus (Forsskål, 1775): Mg, Sg, OShr; [IWP]; PMBC Leiognathus fasciatus (Lacepède, 1803): Mg, OShr; [IWP]; PMBC Leiognathus longispinis (Valenciennes, 1835): Mg, Sg, OShr; [EIO-IA]; PMBC Nuchequula blochii (Valenciennes, 1835): Mg, OShr; [NIOWP]; PMBC Nuchequula gerroides (Bleeker, 1851): Mg, Sg, OShr; [EIOWP]; PMBC Photopectoralis bindus (Valenciennes, 1835): Mg, OShr; [IWP]; PMBC Secutor hanedai Moshizuki & Hayashi, 1989: Mg, Sg, OShr; [IM]; PMBC Secutor indicius Monkolprasit, 1973: OShr; [IM]; PMBC Secutor interruptus (Valenciennes, 1835): Mg, OShr; [EIOWP] Secutor insidiator (Bloch, 1787): Mg, Sg, OShr; [IWP]; PMBC Secutor megalolepis Moshizuki & Hayashi, 1989: Mg, Sg, OShr; [IA]; PMBC Bramidae Brama dussumieri Cuvier, 1831: OShr; [CIR]; PMBC Lutjanidae Aphareus furca (Lacepède, 1801): OShr; [IP]; PMBC Aphareus rutilans Cuvier, 1830: Cr, OShr; [IP]; PMBC Aprion virescens Valenciennes, 1830: Cr, OShr; [IP]; PMBC Lipocheilus carnolabrum (Chan, 1970): OShr; [NIO-WP]; PMBC Lutjanus argentimaculatus (Forsskål, 1775): Cr, Mg, Sg, OShr; [IP]; PMBC Lutjanus bengalensis (Bloch, 1790): Cr, OShr; [IO] Lutjanus biguttatus (Valenciennes 1830): Cr, Mg, Sg, OShr; [EIO-IA]; PMBC Lutjanus bohar (Forsskål, 1775): Cr, OShr; [IP]; PMBC Lutjanus decussatus (Cuvier, 1828): Cr, OShr; [EIO-IA]; PMBC Lutjanus ehrengergii (Peters, 1869): OShr; [IWP] Lutjanus erythropterus Bloch, 1790: Cr, OShr; [NIO-IA] Lutjanus fulviflamma (Forsskål, 1775): Cr, Mg, Sg, OShr; [IWP]; PMBC Lutjanus fulvus (Schneider, 1801): Cr, OShr; [IP]; PMBC Lutjanus gibbus (Forsskål, 1775): Cr, OShr; [IP]; PMBC Lutjanus johnii (Bloch, 1792): Cr, Mg, Sg, OShr; [IWP]; PMBC Lutjanus kasmira (Forsskål, 1775): Cr, OShr; [IP]; PMBC Lutjanus lemniscatus (Valenciennes, 1828): Cr, Mg, OShr; [EIO-IA]; PMBC Lutjanus lunulatus (Park, 1797): Cr, Mg, Sg; [NIO-IA]; PMBC Lutjanus lutjanus Bloch, 1790: Cr, OShr; [IWP]; PMBC Lutjanus madras (Valenciennes, 1831): Cr, Mg, OShr; [IWP]; PMBC Lutjanus malabaricus Schneider, 1801: Cr, OShr; [NIOWP]; PMBC Lutjanus monostigma (Cuvier, 1828): Cr, OShr; [IP]; PMBC Lutjanus quinquelineatus Bloch, 1790: Cr, OShr; [NIOWP]; PMBC Lutjanus rivulatus (Cuvier, 1828): Cr, OShr; [IP]; PMBC Lutjanus rufolineatus (Valenciennes, 1830): OShr; [WP] Lutjanus russelli (Bleeker, 1849): Cr, Mg, Sg, OShr; [IWP]; PMBC Lutjanus sebae (Cuvier, 1828): Cr, OShr; [IWP]; PMBC Lutjanus timorensis (Quoy & Gaimard, 1824): OShr; [WP] Lutjanus vitta (Quoy & Gaimard, 1824): Cr, Sg, OShr; [IWP]; PMBC Macolor macularis Fowler, 1931: Cr, OShr; [EIO-WP] Macolor niger (Forsskål, 1775): Cr, OShr; [IP]; PMBC Paracaesio xanthurus (Bleeker, 1869): OShr; [INP] Pinjalo lewisi Randall, Allen & Anderson, 1987: Cr; [NIOWP] Pinjalo pinjalo (Bleeker, 1850): OShr; [IWP]; PMBC Pristipomoides flavipinnis Shinohara, 1963: OShr; [WCP] Pristipomoides multidens (Day, 1871): OShr; [IWP]; PMBC Pristipomoides typus Bleeker, 1852: OShr; [IA]; PMBC Symphorichthys spilurus (Günther, 1874): Cr, OShr; [WP] Symphorus nematophorus (Bleeker, 1860): Cr, Sg, OShr; [WP]; PMBC Caesionidae Caesio caerulaurea Lacepède, 1801: Cr, Mg, Sg, OShr; [IP]; PMBC Caesio cuning (Bloch, 1791): Cr, Sg, OShr; [EIO-WP]; PMBC Caesio lunaris Cuvier, 1830: Cr, OShr; [IWP]; PMBC Caesio teres Seale, 1906: Cr, OShr; [IP] Caesio varilineata Carpenter, 1987: Cr, OShr; [IWP] Caesio xanthonota Bleeker, 1853: Cr, OShr; [IO]; PMBC Dipterygonotus balteatus (Valenciennes, 1830): Cr, OShr; [IWP]; PMBC Gymnocaesio gymnoptera (Bleeker, 1856): Cr; [IWP]; PMBC Pterocaesio chrysozona (Cuvier, 1830): Cr, OShr; [IWP]; PMBC Pterocaesio pisang (Bloch, 1853): Cr, OShr; [IWP]; PMBC Pterocaesio randalli Carpenter, 1985: Cr; [EIO-IM] Pterocaesio tessellata Carpenter, 1987: Cr, OShr; [EIOIA]; PMBC 59 The fishes of southwestern Thailand Pterocaesio tile (Cuvier, 1830): Cr, OShr; [IP]; PMBC Lobotidae Lobotes surinamensis (Bloch, 1790): Mg, Sg, OShr; [COS]; PMBC Gerreidae Gerres erythrourus (Bleeker, 1791): Mg, Sg, OShr; [EIOWP]; PMBC Gerres filamentosus (Cuvier, 1829): Cr, Mg, Sg, OShr; [IP]; PMBC Gerres limbatus Cuvier, 1830: Mg, OShr; [EIO-WP]; PMBC Gerres longirostris (Lacepède, 1801): Cr, Mg, OShr; [IWP] Gerres lucidus (Cuvier, 1830): Cr; [EIO-WP] Gerres macracanthus Bleeker, 1854: Mg, Sg, OShr; [IWP]; PMBC Gerres oblongus (Cuvier, 1830): Cr, Mg, Sg, OShr; [IP]; PMBC Gerres oyena (Forsskål, 1775): Cr, Mg, Sg, OShr; [IWP]; PMBC Gerres phaiya Iwatsuki & Heemstra, 2001: Mg, Sg, OShr; [EIO]; PMBC Gerres setifer (Hamilton, 1822): OShr; [EIO] Gerres shima Iwatsuki, Kimura & Yoshino, 2007: OShr; [WP] Pentaprion longimanus (Cantor, 1850): OShr; [EIO-WP]; PMBC Haemulidae Diagramma pictum pictum (Thunberg, 1792): Cr, Sg, OShr; [WP]; PMBC Hapalogenys merguiensis Iwatsuki, Satapoomin & Amaoka, 2000: OShr; [AS]; PMBC Plectorhinchus chaetodonoides Lacepède, 1800: Cr, OShr; [IWP]; PMBC Plectorhinchus gibbosus (Lacepède, 1802): Cr, Mg, Sg, OShr; [IP]; PMBC Plectorhinchus macrospilus Satapoomin & Randall, 2000: Cr; [AS]; PMBC Plectorhinchus vittatus (Linnaeus, 1758): Cr, OShr; [IWP] Pomadasys andamanensis Mckay & Satapoomin, 1994: Cr; [AS]; PMBC Pomadasys argenteus (Forsskål, 1775): OShr; [IWP]; PMBC Pomadasys argyreus (Valenciennes, 1833): Mg, OShr; [NIO-IM]; PMBC Pomadasys auritus (Cuvier, 1830): OShr; [IA] Pomadasys furcatus (Bloch & Schneider, 1801): OShr; [IM]; PMBC Pomadasys kaakan (Cuvier, 1830): Mg, Sg, OShr; [IWP]; PMBC Pomadasys maculatus (Bloch, 1793): OShr; [IWP]; PMBC Pomadasys olivaceus (Day, 1875): OShr; [IO] Pomadasys unimaculatus Tian, 1982: Mg, Sg, OShr; [WP]; PMBC Nemipteridae Nemipterus balinensoides (Popta, 1918): OShr; [WP] Nemipterus bathybius Snyder, 1911: OShr; [IA]; PMBC Nemipterus bipunctatus (Ehrenberg, 1830): Cr, OShr; [IO]; PMBC Nemipterus furcosus (Valenciennes, 1830): Cr, OShr; [EIOWP]; PMBC Nemipterus hexodon (Quoy & Gaimard, 1824): OShr; [WP]; PMBC Nemipterus japonicus (Bloch, 1791): OShr; [IWP]; PMBC Nemipterus marginatus (Valenciennes, 1830): OShr; [WP] Nemipterus mesoprion (Bleeker, 1853): OShr; [IM]; PMBC Nemipterus nematophorus (Bleeker, 1853): OShr; [EIOIM]; PMBC Nemipterus peronii (Valenciennes, 1830): Cr, Mg, OShr; [IWP]; PMBC Nemipterus tambuloides (Bleeker, 1853): OShr; [IM]; PMBC Nemipterus thosaporni Russell, 1991: OShr; [IM]; PMBC Nemipterus zysron (Bleeker, 1856-57): Cr, OShr; [IWP]; PMBC Parascolopsis aspinosa (Rao & Rao, 1981): OShr; [NIO] Parascolopsis eriomma (Jordan & Richardson, 1909): OShr; [IWP]; PMBC Parascolopsis inermis (Schlegel, 1843): OShr; [EIO-WP]; PMBC Scolopsis affinis Peters, 1877: Cr, Sg, OShr; [WP]; PMBC Scolopsis bilineatus (Bloch, 1793): Cr, Sg, OShr; [EIOWP]; PMBC Scolopsis ciliatus (Lacepède, 1802): Cr, Mg, Sg, OShr; [WP]; PMBC Scolopsis lineatus Quoy & Gaimard, 1824: Cr, OShr; [WCP]; PMBC Scolopsis margaritifer (Cuvier, 1830): Cr, OShr; [WP] Scolopsis monogramma (Cuvier, 1830): Cr, Sg, OShr; [WP]; PMBC Scolopsis taeniopterus (Valenciennes, 1830): Mg, Sg, OShr; [IA]; PMBC Scolopsis vosmeri (Bloch, 1792): Cr, OShr; [IWP]; PMBC Scolopsis xenochrous Günther, 1872: Cr, OShr; [EIO-WP] Lethrinidae Gnathodentex aurolineatus (Lacepède, 1802): Cr; [IP] Gymnocranius elongatus Senta, 1973: OShr; [IWP] Gymnocranius grandoculis (Valenciennes, 1830): Cr, OShr; [IP]; PMBC Gymnocranius griseus (Schlegel, 1844): Cr, OShr; [EIOWP]; PMBC Gymnocranius microdon (Bleeker, 1851): Cr, OShr; [WCP] Lethrinus conchyliatus (Smith, 1959): OShr; [IO] Lethrinus crocineus Smith, 1959: Cr, OShr; [IO]; PMBC Lethrinus erythracanthus Cuvier, 1830: Cr, OShr; [IP] Lethrinus erythropterus Valenciennes, 1830: Cr, OShr; [IWP] Lethrinus harak (Forsskål, 1775): Cr, OShr; [IP]; PMBC Lethrinus lentjan (Lacepède, 1802): Cr, Sg, OShr; [IWP]; PMBC Lethrinus mahsena (Forsskål, 1775): Cr; [IO]; PMBC Lethrinus microdon Valenciennes, 1830: Cr, OShr; [IWP] 60 Phuket mar. biol. Cent. Res. Bull. Lethrinus nebulosus (Forsskål, 1775): Cr, OShr; [IP]; PMBC Lethrinus obsoletus (Forsskål, 1775): Cr, OShr; [IP] Lethrinus olivaceus Valenciennes, 1830: Cr, OShr; [IP] Lethrinus ornatus Valenciennes, 1830: Cr, Sg, OShr; [EIOWP]; PMBC Lethrinus rubrioperculatus Sato, 1978: Cr; [IP]; PMBC Lethrinus variegatus Valenciennes, 1830: OShr; [IWP]; PMBC Lethrinus xanthochilus Klunzinger, 1870: Cr, OShr; [IP] Monotaxis grandoculis (Forsskål, 1775): Cr, OShr; [IP]; PMBC Wattsia mossambrica (Smith, 1957): OShr; [IWP]; PMBC Sparidae Acanthopagrus berda (Forsskål, 1775): Mg, OShr; [IWP] Argyrops spinifer (Forsskål, 1775): Cr, OShr; [IWP]; PMBC Polynemidae Eleutheronema tetradactylum (Shaw, 1804): Mg, OShr; [NIO-IA]; PMBC Eleutheronema tridactylum (Bleeker, 1849): OShr; [IM] Filimanus perplexa Feltes, 1991: OShr; [EEIO]; PMBC Filimanus similis Feltes, 1991: OShr; [NIO] Filimanus xanthonema (Valenciennes, 1831): OShr; [EEIO] Leptomelanosoma indicum (Shaw, 1804): Mg, OShr; [NIOIM]; PMBC Polydactylus microstomus (Bleeker, 1851): Mg, OShr; [EIOWP]; PMBC Polydactylus plebeius (Broussonet, 1782): Mg, OShr; [IWP]; PMBC Polydactylus sexfilis (Valenciennes, 1831): OShr; [IP] Polydactylus sextarius (Bloch, 1801): OShr; [EIO-WP] Polydactylus siamensis Motomura, Iwatsuki & Yoshino, 2001: OShr; [IM (presently known from west coast of Thailand and Gulf of Thailand)]; PMBC Sciaenidae Dendrophysa russelli (Cuvier, 1830): Mg, Sg, OShr; [EIOIM]; PMBC Johnius amblycephalus (Bleeker, 1855): OShr; [NIO-IA]; PMBC Johnius belangerii (Cuvier, 1830): Mg, OShr; [NIO-IM] Johnius borneensis (Bleeker, 1851): OShr; [NIO-IA] Johnius carutta Bloch, 1793: OShr; [NIO]; PMBC Johnius coitor (Hamilton, 1822): OShr: [EIO-IM] Johnius dussumieri (Cuvier, 1830): OShr; [NIO] Johnius latifrons Sasaki, 1992: OShr; [IM] Macrospinosa cuja (Hamilton, 1822): OShr; [EIO] Nibea soldado (Lacepède, 1802): Mg, OShr; [EIO-IA] Otolithes cuvieri Trewavas, 1974: Mg, OShr; [IO] Otolithes ruber (Bloch & Schneider, 1801): OShr; [IWP] Otolithoides biauritus (Cantor, 1850): OShr; [EIO-IM] Panna microdon (Bleeker, 1849): OShr; [IM] Pennahia anea (Bloch, 1793): Mg, Sg, OShr; [NIO-IM]; PMBC Protonibea diacanthus (Lacepède, 1802): OShr; [NIO-IA]; PMBC Mullidae Mulloidichthys flavolineatus (Lacepède, 1801): Cr, Sg, OShr; [IP] Mulloidichthys vanicolensis (Valenciennes, 1831): Cr, OShr; [IP]; PMBC Parupeneus barberinus (Lacepède, 1801): Cr, OShr; [IP]; PMBC Parupeneus cyclostomus (Lacepède, 1801): Cr, OShr; [IP]; PMBC Parupeneus heptacanthus (Lacepède, 1801): Cr, Sg, OShr; [IWP]; PMBC Parupeneus indicus (Shaw, 1803): Cr, Sg, OShr; [IWP]; PMBC Parupeneus macronema (Lacepède, 1801): Cr, OShr; [IWP]; PMBC Parupeneus pleurostigma (Bennett, 1830): Cr, OShr; [IP]; PMBC Parupeneus trifasciatus Lacepède, 1801: Cr, OShr; [IO] Upeneus asymmetricus Lachner, 1954: OShr; [IWP]; PMBC Upeneus guttatus (Day, 1868): OShr; [IWP] Upeneus moluccensis (Bleeker, 1855): OShr; [IWP] Upeneus sulphureus Cuvier, 1829: Mg, Sg, OShr; [IWP]; PMBC Upeneus sundaicus (Bleeker, 1855): Mg, OShr; [NIO-IA]; PMBC Upeneus tragula Richardson, 1846: Cr, Mg, Sg, OShr; [IWP]; PMBC Upeneus vittatus (Forsskål, 1775): OShr; [IP]; PMBC Pempheridae Parapriacanthus ransonneti Steindachner, 1870: Cr; [WP]; PMBC Pempheris mangula Cuvier, 1829: Cr; [IWP]; PMBC Pempheris otaitensis Cuvier, 1831: Cr; [IWP] Pempheris vanicolensis Cuvier, 1831: Cr; [IM]; PMBC Monodactylidae Monodactylus argenteus (Linnaeus, 1758): Cr, Mg, OShr; [IWP]; PMBC Toxotidae Toxotes chatareus (Hamiton, 1822): Mg; [EIO-IA]; PMBC Toxotes jaculatrix (Pallas, 1766): Mg; [EIO-WP]; PMBC Kyphosidae Kyphosus cinerascens (Forsskål, 1775): Cr; [IP]; PMBC Kyphosus vaigiensis (Quoy & Gaimard, 1825): Cr; [IP]; PMBC Drepaneidae Drepane longimana (Bloch & Schneider, 1801): Mg, Sg, OShr; [IWP]; PMBC Drepane punctata (Linnaeus, 1758): Mg, Sg, OShr; [IWP]; PMBC Chaetodontidae Chaetodon andamanensis Kuiter & Debelius, 1999: Cr; [EIO]; PMBC Chaetodon auriga Forsskål, 1775: Cr; [IP]; PMBC Chaetodon bennetti Cuvier, 1831: Cr; [IP] Chaetodon citrinellus Cuvier, 1831: Cr; [IP]; PMBC Chaetodon collare Bloch, 1787: Cr; [NIO]; PMBC 61 The fishes of southwestern Thailand Chaetodon decussatus Cuvier, 1831: Cr; [EIO]; PMBC Chaetodon ephippium Cuvier, 1831: Cr; [IP]; PMBC Chaetodon falcula Bloch, 1793: Cr; [IO]; PMBC Chaetodon gardineri Norman, 1939: Cr; [IO]; PMBC Chaetodon guttatissimus Bennett, 1832: Cr; [IO]; PMBC Chaetodon interruptus Ahl, 1922: Cr; [IO] Chaetodon kleinii Bloch, 1790: Cr; [IP]; PMBC Chaetodon lineolatus Cuvier, 1831: Cr; [IP]; PMBC Chaetodon lunula (Lacepède, 1803): Cr; [IP]; PMBC Chaetodon madagaskariensis Ahl, 1923: Cr; [IO]; PMBC Chaetodon melannotus Bloch & Schneider, 1801: Cr; [IP]; PMBC Chaetodon meyeri Bloch & Schneider, 1801: Cr; [IP]; PMBC Chaetodon octofasciatus Bloch, 1787: Cr; [EIO-WP]; PMBC Chaetodon oxycephalus Bleeker, 1853: Cr; [EIO-WP] Chaetodon rafflesi Bennett, 1830: Cr; [EIO-WCP]; PMBC Chaetodon semeion Bleeker, 1855: Cr; [EIO-WCP] Chaetodon triangulum Cuvier, 1831: Cr; [IO]; PMBC Chaetodon trifascialis Quoy & Gaimard, 1825: Cr; [IP]; PMBC Chaetodon trifasciatus Park, 1797: Cr; [IO]; PMBC Chaetodon vagabundus Linnaeus, 1758: Cr; [IP]; PMBC Chaetodon xanthocephalus Bennett, 1832: Cr; [IO]; PMBC Chelmon rostratus (Linnaeus, 1758): Cr; [WP]; PMBC Coradion altivelis Mcculloch, 1916: Cr; [WP] Coradion chrysozonus (Cuvier, 1831): Cr, OShr; [WP]; PMBC Forcipiger flavissimus Jordan & Mcgregor, 1898: Cr; [IP]; PMBC Forcipiger longirostris (Broussonet, 1782): Cr; [IP] Hemitaurichthys zoster (Bennett, 1831): Cr; [IO]; PMBC Heniochus acuminatus (Linnaues, 1758): Cr, Mg, Sg, OShr; [IP]; PMBC Heniochus diphreutes Jordan, 1903: Cr, OShr; [IP] Heniochus pleurotaenia Ahl, 1923: Cr; [EIO]; PMBC Heniochus singularius Smith & Radcliffe, 1911: Cr; [EIOWCP]; PMBC Parachaetodon ocellatus (Cuvier, 1831): Cr, Sg: [EIO-WP]; PMBC Pomacanthidae Apolemichthys trimaculatus (Lacapède, 1831): Cr; [IWP]; PMBC Apolemichthys xanthurus (Bennett, 1832): Cr; [IO]; PMBC Centropyge eibli Klausewitz, 1963: Cr; [EIO]; PMBC Centropyge flavipectoralis Randall & Klausewitz, 1977: Cr; [EIO] Centropyge multispinis (Playfair, 1866): Cr; [IO]; PMBC Chaetodontoplus mesoleucus (Bloch, 1787): Cr; [WP] Genicanthus caudovittatus (Günther, 1860): Cr, OShr; [IO]; PMBC Pomacanthus annularis (Bloch, 1787): Cr, OShr; [IWP]; PMBC Pomacanthus imperator (Bloch, 1787): Cr; [IP]; PMBC Pomacanthus navarchus (Cuvier, 1831): Cr; [IA] Pomacanthus semicirculatus (Cuvier, 1831): Cr; [IP] Pomacanthus sexstriatus (Cuvier, 1831): Cr; [WP] Pomacanthus xanthometopon (Bleeker, 1853): Cr; [EIOWCP] Pygoplites diacanthus (Boddaert, 1772): Cr; [IP]; PMBC Terapontidae Pelates quadrilineatus (Bloch, 1790): Mg, Sg; [IWP]; PMBC Pseudoterapon theraps (Cuvier, 1829): Mg, Sg, OShr; [IWP]; PMBC Terapon jarbua (Forsskål, 1775): Cr, Mg, Sg, OShr; [IWP]; PMBC Terapon puta (Cuvier, 1829): Mg, Sg, OShr; [IWP]; PMBC Kuhliidae Kuhlia mugil (Bloch & Schneider, 1801): Cr; [IP]; PMBC Cirrhitidae Cirrhitichthys falco Randall, 1963: Cr; [EIO-WP] Cirrhitichthys oxycephalus (Bleeker, 1855): Cr; [IP]; PMBC Oxycirrhites typus Bleeker, 1857: Cr; [IP] Paracirrhites arcatus (Cuvier, 1829): Cr; [IP] Paracirrhites fosteri (Schneider, 1801): Cr; [IP]; PMBC Cepolidae Acanthocepola abbreviata (Valenciennes, 1835): OShr; [NIO-IM]; PMBC Acanthocepola indica (Day, 1888): OShr; [IWP] Pomacentridae Abudefduf bengalensis (Bloch, 1787): Cr; [EIO-WP]; PMBC Abudefduf notatus (Day, 1869): Cr; [IWP]; PMBC Abudefduf septemfasciatus (Cuvier, 1830): Cr; [IP]; PMBC Abudefduf sordidus (Forsskål, 1775): Cr; [IP]; PMBC Abudefduf vaigiensis (Quoy & Gaimard, 1825): Cr; [IP]; PMBC Amblyglyphidodon aureus (Cuvier, 1830): Cr; [WCP]; PMBC Amblyglyphidodon indicus Allen & Randall, 2002: Cr; [IO] Amblyglyphidodon leucogaster (Bleeker, 1847): Cr; [WP]; PMBC Amphiprion akallopisos Bleeekr, 1853: Cr; [IO]; PMBC Amphiprion clarkii (Bennett, 1830): Cr, Sg; [IWP]; PMBC Amphiprion ephippium (Bloch, 1790): Cr; [EIO]; PMBC Amphiprion ocellaris Cuvier, 1830: Cr; [WP]; PMBC Amphiprion sebae Bleeekr, 1853: Cr, Sg; [EIO]; PMBC Cheiloprion labiatus (Day, 1877): Cr; [IA]; PMBC Chromis atripectoralis Welander & Schultz, 1951: Cr; [IWP]; PMBC Chromis cinerascens (Cuvier, 1830): Cr; [EIO-IA]; PMBC Chromis delta Randall, 1987: Cr; [EIO-WP] Chromis dimidiata (Kluzinger, 1871): Cr; [IO]; PMBC Chromis elerae Fowler & Bean, 1928: Cr; [EIO-WP] Chromis flavipectoralis Randall, 1990: Cr; [EIO]; PMBC Chromis lepidolepis Bleeker, 1877: Cr; [IP] Chromis opercularis (Günther, 1866): Cr; [IO]; PMBC Chromis ternatensis (Bleeker, 1856): Cr; [IP]; PMBC Chromis viridis (Cuvier, 1830): Cr; [IP]; PMBC 62 Phuket mar. biol. Cent. Res. Bull. Chromis weberi Fowler & Bean, 1928: Cr; [IP]; PMBC Chromis xouthos Allen & Erdmann, 2005: Cr; [AS] Chrysiptera biocellata (Quoy & Gaimard, 1825): Cr; [IP]; PMBC Chrysiptera brownriggii (Bennett, 1828): Cr; [IP]; PMBC Chrysiptera glauca (Cuvier, 1830): Cr; [IP] Chrysiptera rollandi (Whitley, 1961): Cr; [IA]; PMBC Chrysiptera unimaculata (Cuvier, 1830): Cr; [IWP]; PMBC Dascyllus aruanus (Linnaeus, 1758): Cr; [IP]; PMBC Dascyllus carneus Fischer, 1885: Cr; [IO]; PMBC Dascyllus trimaculatus (Rüppell, 1828): Cr; [IP]; PMBC Dischistodus perspicillatus (Cuvier, 1830): Cr; [WP]; PMBC Hemiglyphidodon plagiometopon (Bleeker, 1852): Cr; [IA]; PMBC Lepidozygus tapeinosoma (Bleeker, 1856): Cr; [IP]; PMBC Neoglyphidodon melas (Cuvier, 1830): Cr; [IWP]; PMBC Neoglyphidodon nigroris (Cuvier, 1830): Cr; [WP]; PMBC Neopomacentrus anabatoides (Bleeker, 1847): Cr; [IA]; PMBC Neopomacentrus bankieri (Richardson, 1846): Cr; [WP] Neopomacentrus cyanomos (Bleeker, 1856): Cr; [IWP]; PMBC Neopomacentrus filamentosus (Macleay, 1883): Cr; [IA]; PMBC Neopomacentrus sororius Randall & Allen, 2005: Cr; [IO]; PMBC Neopomacentrus taeniurus (Bleeker, 1856): Cr; [IWP]; PMBC Neopomacentrus violascens (Bleeker, 1848): Cr; [IA] Plectroglyphidodon dickii (Leinard, 1839): Cr; [IP]; PMBC Plectroglyphidodon johnstonianus Fowler & Ball, 1924: Cr; [IP]; PMBC Plectroglyphidodon lacrymatus (Quoy & Gaimard, 1825): Cr; [IP]; PMBC Plectroglyphidodon leucozona (Bleeker, 1859): Cr; [IP]; PMBC Pomacentrus adelus Allen, 1991: Cr; [IA]; PMBC Pomacentrus alleni Burgess, 1981: Cr; [EEIO]; PMBC Pomacentrus amboinensis Bleeker, 1868: Cr; [WCP]; PMBC Pomacentrus azuremaculatus Allen, 1991: Cr; [EEIO]; PMBC Pomacentrus chrysurus Cuvier, 1830: Cr; [EIO-WP]; PMBC Pomacentrus lepidogenys Fowler & Bean, 1928: Cr; [WP]; PMBC Pomacentrus moluccensis Bleeker, 1853: Cr; [WP]; PMBC Pomacentrus nagasakiensis Tanaka, 1917: Cr; [EIO-WP]; PMBC Pomacentrus pavo (Bloch, 1787): Cr; [IP]; PMBC Pomacentrus philippinus Evermann & Seale, 1907: Cr; [EIO-WP]; PMBC Pomacentrus polyspinus Allen, 1991: Cr; [AS]; PMBC Pomacentrus similis Allen, 1991: Cr; [EIO]; PMBC Pomacentrus tripunctatus Cuvier, 1830: Cr; [EIO-WP]; PMBC Pomacentrus xanthosternus Allen, 1991: Cr; [EEIO]; PMBC Premnas biaculeatus (Bloch, 1790): Cr; [IA] Pristotis obtusirostris (Günther, 1862): Cr, OShr; [IWP]; PMBC Stegastes fasciolatus (de Vis, 1884): Cr; [IP]; PMBC Stegastes nigricans (Lacepède, 1803): Cr; [IP]; PMBC Stegastes obreptus (Whitley, 1948): Cr; [EIO-WP]; PMBC Stegastes punctatus (Quoy & Gaimard, 1825): Cr; [IWP]; PMBC Teixeirichthys jordani (Rutter, 1897): OShr; [IWP] Labridae Anampses caeruleopunctatus Rüppell, 1829: Cr; [IP]; PMBC Anampses lineatus Randall, 1972: Cr; [IO]; PMBC Anampses meleagrides Valenciennes, 1840: Cr; [IWP]; PMBC Anampses twistii Bleeker, 1856: Cr; [IP]; PMBC Bodianus axillaris (Bennett, 1832): Cr; [IP] Bodianus diana (Lacepède, 1801): Cr; [IO]; PMBC Bodianus mesothorax (Bloch & Schneider, 1801): Cr; [WP]; PMBC Bodianus neilli (Day, 1867): Cr; [EIO]; PMBC Cheilinus chlorourus (Bloch, 1791): Cr, OShr; [IP]; PMBC Cheilinus fasciatus (Bloch, 1791): Cr; [IP]; PMBC Cheilinus oxycephalus Bleeker, 1853: Cr; [IP] Cheilinus trilobatus Lacepède, 1801: Cr; [IP]; PMBC Cheilinus undulatus Rüppell, 1835: Cr; [IP] Choerodon robustus (Günther, 1862): OShr; [IWP]; PMBC Cirrhilabrus cyanopleura (Bleeker, 1851): Cr; [EEIO]; PMBC Cirrhilabrus exquisitus Smith, 1957: Cr; [IP]; PMBC Cirrhilabrus joanallenae Allen, 2000: Cr; [AS] Coris batuensis (Bleeker, 1856): Cr; [EIO-WCP]; PMBC Coris cuvieri (Bennett, 1831): Cr; [IO]; PMBC Diproctacanthus xanthurus (Bleeker, 1856): Cr; [IA] Epibulus insidiator (Pallas, 1770): Cr; [IP]; PMBC Gomphosus caeruleus Lacepède, 1801: Cr; [IO]; PMBC Halichoeres argus (Bloch, 1791): Cr; [WP]; PMBC Halichoeres bicolor (Bloch & Schneider, 1801): Cr, Mg, Sg; [EIO-IM]; PMBC Halichoeres chloropterus (Bloch, 1791): Cr; [IA]; PMBC Halichoeres cosmetus Randall & Smith, 1982: Cr; [IO] Halichoeres hortulanus (Lacepède, 1801): Cr; [IP]; PMBC Halichoeres kallochroma (Bleeker, 1835): Cr; [AS]; PMBC Halichoeres leucoxanthus Randall & Smith, 1982: Cr; [EIO]; PMBC Halichoeres margaritaceus (Valenciennes, 1839): Cr; [WCP] Halichoeres marginatus Rüppell, 1835: Cr; [IP]; PMBC Halichoeres nebulosus (Valenciennes, 1839): Cr; [IWP]; PMBC Halichoeres nigrescens (Bloch & Schneider, 1801): Cr, Sg; [IWP]; PMBC Halichoeres scapularis (Bennett, 1832): Cr; [IWP]; PMBC 63 The fishes of southwestern Thailand Halichoeres timorensis (Bleeker, 1852): Cr; [EIO-IM]; PMBC Halichoeres vrolikii (Bleeker, 1855): Cr; [EIO]; PMBC Halichoeres zeylonicus (Bennett, 1832): Cr, OShr; [IO]; PMBC Hemigymnus fasciatus (Bloch, 1792): Cr; [IP]; PMBC Hemigymnus melapterus (Bloch, 1791): Cr; [IP]; PMBC Hologymnosus annulatus (Lacepède, 1801): Cr; [IP]; PMBC Hologymnosus doliatus (Lacepède, 1801): Cr; [IP] Iniistius baldwini (Jordan & Evermann, 1903): OShr; [IP]; PMBC Iniistius bimaculatus (Rüppell, 1829): OShr; [IO]; PMBC Iniistius pavo (Valenciennes, 1840): Cr, OShr; [IP]; PMBC Labrichthys unilineatus (Guichenot, 1847): Cr; [IP]; PMBC Labroides bicolor Fowler & Bean, 1928: Cr; [IP] Labroides dimidiatus (Valenciennes, 1839): Cr; [IP]; PMBC Labropsis xanthonota Randall, 1981: Cr; [IWP] Leptojulis chrysotaenia Randall & Ferraris, 1981: Cr; [EIO] Leptojulis cyanopleura (Bleeker, 1853): Cr, OShr; [NIOWP]; PMBC Leptojulis urostigma Randall, 1996: OShr; [IA] Macropharyngodon ornatus Randall, 1978: Cr; [IWP]; PMBC Novaculichthys taeniourus (Lacepède, 1801): Cr; [IP] Oxycheilinus areanatus (Valenciennes, 1840): Cr; [IP] Oxycheilinus bimaculatus (Valenciennes, 1840): Cr; [IP] Oxycheilinus celebicus (Bleeker, 1853): Cr; [WP]; PMBC Oxycheilinus digramma (Lacepède, 1801): Cr, OShr; [IP]; PMBC Oxycheilinus orientalis (Günther, 1862): Cr; [IWP] Paracheilinus mccoskeri Randall & Harmelin-Vivien, 1977: Cr; [IO]; PMBC Pseudocheilinus evanidus Jordan & Evermann, 1903: Cr; [IP] Pseudocheilinus hexataenia (Bleeker, 1857): Cr; [IP]; PMBC Pseudodax moluccanus (Valenciennes, 1840): Cr; [IP]; PMBC Pseudojuloides kaleidos Kuiter & Randall, 1995: Cr; [EIO]; PMBC Pteragogus cryptus Randall, 1981: Cr; [IWP]; PMBC Stethojulis albovittata (Bonnaterre, 1788): Cr; [IO]; PMBC Stethojulis bandanensis (Bleeker, 1851): Cr; [WCP] Stethojulis interrupta (Bleeker, 1851): Cr; [IWP]; PMBC Stethojulis strigiventer (Bennett, 1832): Cr, Sg; [IP] Stethojulis trilineata (Bloch & Schneider, 1801): Cr; [EIOWCP]; PMBC Suezichthys caudovittatus (Steindachner, 1898): OShr; [IO]; PMBC Thalassoma amblycephalum (Bleeker, 1856): Cr; [IP]; PMBC Thalassoma hardwicke (Bennett, 1830): Cr; [IP]; PMBC Thalassoma jansenii (Bleeker, 1856): Cr; [EIO-WP]; PMBC Thalassoma lunare (Linnaeus, 1758): Cr; [IP]; PMBC Thalassoma purpureum (Forsskål, 1775): Cr; [IP] Thalassoma trilobatum (Lacepède, 1801): Cr; [IP]; PMBC Wetmorella nigropinnata (Seale, 1901): Cr; [IP]; PMBC Xiphocheilus quadrimaculatus (Günther, 1880): OShr; [EIO-WP] Scaridae Bolbometopon muricatum (Valenciennes, 1840): Cr; [IP]; PMBC Calotomus carolinus (Valenciennes, 1840): Cr; [IP]; PMBC Cetoscarus bicolor (Rüppell, 1829): Cr; [IP] Chlorurus capistratoides (Bleeker, 1847): Cr; [IO]; PMBC Chlorurus rhakoura Randall & Anderson, 1997: Cr; [EIO]; PMBC Chlorurus sordidus (Forsskål, 1775): Cr; [IP]; PMBC Chlorurus strongylocephalus (Bleeker, 1854): Cr; [IO]; Chlorurus troschelii (Bleeker, 1853): Cr; [EEIO] Hipposcarus harid (Forsskål, 1775): Cr; [IO] Scarus frenatus Lacepède, 1802: Cr; [IP]; PMBC Scarus ghobban Forsskål, 1775: Cr, Sg; [IP]; PMBC Scarus maculipinna Westneat, Satapoomin & Randall, 2007: Cr; [EEIO]; PMBC Scarus niger Forsskål, 1775: Cr; [IP]; PMBC Scarus prasiognathos Valenciennes, 1840: Cr; [EIO-WP]; PMBC Scarus quoyi Valenciennes, 1840: Cr; [EIO-WP]; PMBC Scarus rubroviolaceus Bleeker, 1847: Cr; [IP]; PMBC Scarus russelii Valenciennes, 1840: Cr; [IO]; PMBC Scarus scaber Valenciennes, 1840: Cr; [IO]; PMBC Scarus tricolor Bleeker, 1847: Cr; [IWP]; PMBC Scarus viridifucatus (Smith, 1956): Cr; [IO]; PMBC Champsodontidae Champsodon nudivittis (Ogilby, 1895): OShr; [IWP] Champsodon vorax Günther, 1867: OShr; [EIO-WP] Pinguipedidae Parapercis alboguttata (Günther, 1872): OShr; [NIO-IA]; PMBC Parapercis clathrata Ogilby, 1910: Cr, OShr; [WCP]; PMBC Parapercis cylindrica (Bloch, 1792): Cr: [EIO-WP], PMBC Parapercis hexophthalma (Cuvier, 1829): Cr, OShr; [IWP]; PMBC Parapercis maculata (Bloch & Schneider, 1801): OShr; [IO] Parapercis millepunctata (Günther, 1860): Cr, OShr; [EIOWCP]; PMBC Parapercis punctata (Cuvier, 1829): Mg, OShr; [IO]; PMBC Parapercis quadrispinosa (Weber, 1913): OShr; [EIO]; PMBC Parapercis schauinslandi (Steindachner, 1900): Cr; [IP] Parapercis snyderi Jordan & Starks, 1905: Cr, OShr; [WP] Parapercis tetracantha (Lacepède, 1800): Cr, OShr; [IWP]; PMBC Parapercis xanthozona (Bleeker, 1849): Cr, OShr; [IWP]; PMBC Trichonotidae Trichonotus setiger Bloch & Schneider, 1801: OShr; [WP] 64 Phuket mar. biol. Cent. Res. Bull. Trichonotus sp.1: Cr, OShr Percophidae Bembrops caudimaculata Steindachner, 1876: OShr; [IWP] Bembrops platyrhynchus (Alcock, 1894): OShr; [IWP]; PMBC Ammodytidae Bleekeria kallolepis Günther, 1862: OShr; [EIO] Uranoscopidae Ichthyoscopus lebeck (Schneider, 1801): OShr; [EIO]; PMBC Uranoscopus affinis Cuvier, 1829: OShr; [IWP]; PMBC Uranoscopus bicinctus Temminck & Schlegel, 1843: OShr; [EIO-WP] Uranoscopus cognatus Cantor, 1849: OShr; [EIO-WP] Uranoscopus oligolepis Bleeker, 1878: OShr; [IO] Tripterygiidae Ennaepterygius fasciatus (Weber, 1909): Cr; [IP] Ennaepterygius nanus (Schultz, 1960): Cr; [WP] Ennaepterygius philippinus (Peters, 1868): Cr; [IWP] Enneapterygius sp.1: Enneapterygius tutuilae Jordan & Seale, 1906: Cr; [IP]; PMBC Helcogramma chica Rosenblatt, 1960: Cr; [EIO-WP] Helcogramma lacuna Williams & Howe, 2003: Cr; [AS] Helcogramma obtusirostre Klunzinger, 1871: Cr; [IWP] Helcogramma rosea Holleman, 2006: Cr; [EIO] Helcogramma sp.1: Cr Helcogramma springeri Hansen, 1986: Cr; [WP]; PMBC Helcogramma striata Hansen, 1986: Cr; [EIO-WP]; PMBC Norfolkia brachylepis (Schultz, 1960): Cr; [IP] Ucla xenogrammus Holleman, 1993: Cr; [IP]; PMBC Blenniidae Alticus saliens (Lacepède, 1800): Cr; [IP]; PMBC Andamia reyi Sauvage, 1880: Cr; [IM]; PMBC Aspidontus dussumieri (Valenciennes, 1836): Cr; [IP] Aspidontus tractus Fowler, 1903: Cr; [IO]; PMBC Atrosalarias fuscus (Rüppell, 1835): Cr; [IO]; PMBC Blenniella chrysospilos (Bleeker, 1857): Cr; [IP] Blenniella cyanostigma (Bleeker, 1849): Cr; [IO] Blenniella leopardus (Fowler, 1904): Cr; [AS]; PMBC Blenniella periophthalmus (Valenciennes, 1836): Cr; [IWP] Cirripectes auritus Carlson, 1980: Cr; [IP] Cirripectes castaneus (Valenciennes, 1836): Cr; [IWP] Cirripectes filamentosus (Alleyne & Macleay, 1877): Cr; [IWP]; PMBC Cirripectes stigmaticus Strasburg & Schultz, 1953: Cr; [IWP] Crossosalarias macrospilus Smith-Vaniz & Springer, 1971: Cr; [WP] Ecsenius bicolor (Day, 1888): Cr; [EIO-WP]; PMBC Ecsenius lubbocki Springer, 1988: Cr; [EEIO]; PMBC Ecsenius midas Starck, 1969: Cr; [IP]; PMBC Ecsenius nalolo Smith, 1959: Cr; [IO]; PMBC Ecsenius paroculus Springer, 1988: Cr; [EEIO]; PMBC Enchelyurus kraussi (Klunzinger, 1871): Cr; [IWP]; PMBC Entomacrodus striatus (Quoy & Gaimard, 1836): Cr; [IP]; PMBC Entomacrodus vermiculatus (Valenciennes, 1836): Cr; [IO]; PMBC Exallias brevis (Kner, 1968): Cr; [IP] Glyptoparus delicatulus Smith, 1959: Cr; [IWP] Istiblennius bellus (Günther, 1861): Cr; [IP]; PMBC Istiblennius dussumieri (Valenciennes, 1836): Cr; [IWP]; PMBC Istiblennius edentulus (Schneider, 1801): Cr; [IP]; PMBC Istiblennius lineatus (Valenciennes, 1836): Cr; [EIO-WCP] Laiphognathus multimaculatus Smith, 1955: Cr; [IWP]; PMBC Meiacanthus smithi Klausewitz, 1916: Cr; [EIO]; PMBC Meiacanthus urostigma Smith-Vaniz, Satapoomin & Allen, 2001: Cr; [AS]; PMBC Omobranchus elongatus (Peters, 1855): Cr; [IWP]; PMBC Omobranchus fasciolatus (Valenciennes, 1836): Mg; [IO]; PMBC Omobranchus ferox (Herre, 1927): Mg; [IP]; PMBC Omobranchus obliquus (Garman, 1903): Cr; [WCP]; PMBC Omobranchus punctatus (Valenciennes, 1836): Mg; [IWP]; PMBC Omobranchus smithi (Rao, 1973): Mg; [EIO-IM]; PMBC Omobranchus zebra (Bleeker, 1868): Mg; [EIO-IM] Petroscirtes breviceps (Valenciennes, 1836): Cr, Mg, Sg; [IWP]; PMBC Petroscirtes mitratus Rüppell, 1830: Cr, Sg; [IP]; PMBC Petroscirtes thepassi Bleeker, 1853: Sg; [WP] Petroscirtes variabilis Cantor, 1850: Sg; [EIO-IA]; PMBC Plagiotremus phenax Smith-Vaniz, 1976: Cr; [EIO]; PMBC Plagiotremus rhinorhynchos (Bleeker, 1852): Cr; [IP]; PMBC Plagiotremus tapeinosoma (Bleeker, 1857): Cr; [IP]; PMBC Plagiotremus townsendi (Regan, 1905): Cr; [IO] Praealticus oortii (Bleeker, 1851): Cr; [IM] Praealticus triangulus (Chapman, 1951): Cr; [EEIO] Salarias fasciatus (Bloch, 1786): Cr; [IWP]; PMBC Salarias guttatus Valenciennes, 1836: Cr; [IA]; PMBC Salarias sinuosus Snyder, 1908: Cr; [WP]; PMBC Xiphasia setifer Swainson, 1839: OShr; [IWP] Gobiesocidae Diademichthys lineatus (Sauvage, 1883): Cr; [IWP] Discotrema crinophilum Briggs, 1976: Cr; [WP] Lepadichthys lineatus Briggs, 1966: Cr; [IWP] Callionymidae Bathycallionymus kaianus (Günther, 1880): OShr; [IWP] Calliurichthys japonicus (Houttuyn, 1782): OShr; [WP]; PMBC Dactylopus dactylopus (Valenciennes, 1837): Mg, Sg, OShr; [WP]; PMBC Repomucenus belcheri recurvispinnis (Li, 1966): OShr; [EIO-IM] Repomucenus doryssus (Jordan & Fowler, 1903): OShr; [WP] Repomucenus filamentosus (Valenciennes, 1837): OShr; [IP] 65 The fishes of southwestern Thailand Repomucenus hindsi (Richardson, 1844): Sg, OShr; [NIOIM] Repomucenus octostigmatus (Fricke, 1981): OShr; [IM] Repomucenus sagitta (Pallas, 1770): OShr; [EIO-IM] Repomucenus schaapii (Bleeker, 1852): Mg, Sg, OShr; [EIOIM]; PMBC Paradiplogrammus enneactis (Bleeker, 1879): Mg, Sg; [WP]; PMBC Synchiropus lineolatus (Valenciennes, 1837): Cr; [IWP] Synchiropus stellatus Smith, 1963: Cr; [IO] Eleotridae Bostrychus sinensis (Lacepède, 1801): Mg; [EIO-WP] Butis butis (Hamilton, 1822): Mg, Sg; [IWP]; PMBC Butis humeralis (Valenciennes, 1837): Mg; [EIO-WP]; PMBC Butis koilomatodon (Bleeker, 1849): Mg, Sg; [IWP]; PMBC Calumia godeffroyi (Günther, 1877): Cr; [IP] Eleotris melanosoma Bleeker, 1852): Mg; [IWP]; PMBC Odonteleotris macrodon (Bleeker, 1853): Mg; [EIO-IM]; PMBC Ophiocara aporos (Bleeker, 1854): Mg; [IWP]; PMBC Ophiocara porocephala (Valenciennes, 1837): Mg; [IWP]; PMBC Xenisthmidae Xenisthmus africanus Smith, 1958: Cr; [IO] Xenisthmus polyzonatus (Klunzinger, 1817): Cr; [IP] Gobiidae Acentrogobius audax Smith, 1959: Sg; [IWP]; PMBC Acentrogobius caninus (Valenciennes, 1837): Mg, Sg; [IWP]; PMBC Acentrogobius chlorostigmatoides (Bleeker, 1849): OShr; [WP]; PMBC Acentrogobius cyanomos (Bleeker, 1849): Mg, Sg; [EIOIM]; PMBC Acentrogobius janthinopterus (Bleeker, 1852): Mg; [WP]; PMBC Acentrogobius madraspatensis (Day, 1868): Mg, Sg; [EIOWP]; PMBC Acentrogobius multifasciatus (Herre, 1927): Cr, Sg; [WP]; PMBC Acentrogobius sp.1: Sg Acentrogobius viridipunctatus (Valenciennes, 1837): Mg; [IWP]; PMBC Amblyeleotris aurora (Polunin & Lubbock, 1977): Cr; [IO] Amblyeleotris diagonalis Polunin & Lubbock, 1979: Cr; [IP]; PMBC Amblyeleotris downingi Randall, 1994: Cr; OShr; [NIO]; PMBC Amblyeleotris fontanesii (Bleeker, 1852): Cr, OShr; [WP]; PMBC Amblyeleotris gymnocephala (Bleeker, 1853): Cr; [IA]; PMBC Amblyeleotris latifasciata Polunin & Lubbock, 1979: Cr; [IA]; PMBC Amblyeleotris periophthalma (Bleeker, 1853): Cr; [IWP]; PMBC Amblyeleotris steinitzi (Klausewits, 1974): Cr; [IWP]; PMBC Amblyeleotris triguttatus Randall, 1994: Cr; [IO] Amblyeleotris wheeleri (Polunin & Lubbock, 1977): Cr; [IWP] Amblygobius hectori (Smith, 1957): Cr; [IWP]; PMBC Amblygobius nocturnus (Herre, 1945): Cr; [NIO-WCP]; PMBC Amblygobius semicinctus (Bennett, 1833): Cr; [IO]; PMBC Amblygobius stethophthalmus (Bleeker, 1851): Cr; [IM] Amoya gracilis (Bleeker, 1875): Mg; [IA]; PMBC Amoya moloanus (Herre, 1927): Mg, Sg; [WP]; PMBC Apocryptodon madurensis (Bleeker, 1849): Mg, Sg; [EIOWP]; PMBC Arcygobius baliurus (Valenciennes, 1837): Sg; [IWP]; PMBC Asterropteryx dfh sp.3: Cr Asterropteryx ensiferus (Bleeker, 1874): Cr; [IP]; PMBC Asterropteryx semipunctatus Rüppell, 1828: Cr; [IP]; PMBC Barbuligobius boehlkei Lachner & McKinney, 1974: Cr; [IWP] Bathygobius cocosensis (Bleeker, 1854): Cr; [IP]; PMBC Bathygobius crassiceps (Jordan & Seale, 1906): Cr; [IP]; PMBC Bathygobius cyclopterus (Valenciennes, 1837): Cr; [IWP]; PMBC Bathygobius fuscus (Rüppell, 1830): Cr, Sg; [IWP]; PMBC Bathygobius karachiensis Hoda & Goren, 1990: Cr; [EIO] Bathygobius laddi (Fowler, 1931): Cr; [IP]; PMBC Bathygobius meggitti (Hora & Mukuji, 1936): Cr; [WP] Boleophthalmus boddarti (Pallas, 1770): Mg; [NIO-WP]; PMBC Brachygobius kabiliensis Inger, 1958: Mg; [IM] Bryaninops amplus Larson, 1985: Cr; [IP] Bryaninops erythops (Jordan & Seale, 1906): Cr; [EIOWP] Bryaninops loki Larson, 1985: Cr; [IWP] Bryaninops natans Larson, 1985: Cr; [IWP]; PMBC Bryaninops ridens Smith, 1959: Cr; [IWP]; PMBC Bryaninops youngei (Davis & Cohen, 1969): Cr; [IP] Cabillus tongarevae (Fowler, 1927): Cr; [EIO-WP]; PMBC Callogobius andamanensis Menon & Chatterjee, 1974: Mg; [AS] Callogobius dfh sp.3: Cr Callogobius dfh sp.6: Cr Callogobius dfh sp.11: Cr Callogobius dfh sp.13: Cr Callogobius dfh sp.15: Cr Callogobius dfh sp.16: Cr Callogobius flavobrunneus (Smith, 1958): Cr; [IO]; PMBC Callogobius maculipinnis (Fowler, 1918): Cr; [IWP] Callogobius plumatus (Smith, 1959): Cr; [IO] Callogobius rw-th sp.2: Mg Callogobius sclateri (Steindachner, 1880): Cr; [IP]; PMBC Cristatogobius nonatoae (Ablan, 1940): Mg; [WP]; PMBC 66 Phuket mar. biol. Cent. Res. Bull. Cryptocentrus caeruleomaculatus (Herre, 1933): Cr, Sg; [IP]; PMBC Cryptocentrus cyanotaenia (Bleeker, 1853): Cr, Mg, OShr; [IA]; PMBC Cryptocentrus fasciatus (Playfair & Gunther, 1867): Cr; [IWP]; PMBC Cryptocentrus leucostictus (Günther, 1871): Cr; [WCP] Cryptocentrus maudae Fowler, 1937: Cr; [IA]; PMBC Cryptocentrus melanopus (Bleeker, 1860): Cr, Sg, OShr; [WP]; PMBC Cryptocentrus pavoninoides (Bleeker, 1849): Cr, OShr; [IA]; PMBC Cryptocentrus sp.1 (dfh sp.27): Cr, Sg; [WP]; PMBC Cryptocentrus strigilliceps (Jordan & Seale, 1906): Cr, Sg; [IP]; PMBC Ctenogobiops aurocingulus (Herre, 1935): Cr; [WCP] Ctenogobiops crocineus Smith, 1959: Cr; [IWP]; PMBC Ctenogobiops feroculus Lubbock & Polunin, 1977: Cr; [IWP] Ctenogobiops pomastictus Lubbock & Polunin, 1977: Cr; [WP]; PMBC Drombus globiceps (Hora, 1923): Mg; [EIO-WP]; PMBC Drombus key (Smith, 1947): Cr, Mg, Sg; [IO]; PMBC Drombus ocyurus (Jordan & Seale, 1907): Mg, Sg; [WP]; PMBC Drombus triangularis (Weber, 1911): Mg, Sg; [IWP]; PMBC Eugnathogobius illotus (Larson, 1999): Mg; [IM]; PMBC Eugnathogobius microps Smith, 1931: Mg; [IM (presently known from west coast of Thailand, Gulf of Thailand and southern Vietnam)] Eugnathogobius mindora (Herre, 1945): Mg; WP Eugnathogobius polylepis (Wu & Ni, 1985): Mg; [IA] Eugnathogobius variegatus (Peters, 1869): Mg; [IM]; PMBC Eviota afelei Jordan & Seale, 1906: Cr; [WCP] Eviota albolineata Jewett & Lachner, 1983: Cr; [IP]; PMBC Eviota herrei Jordan & Seale, 1906: Cr; [WCP] Eviota indica Lachner & Karnella, 1980: Cr; [IO]; PMBC Eviota infulata (Smith, 1956): Cr; [IP] Eviota mikiae Allen, 2001: Cr; [EIO] Eviota nigriventris Giltay, 1933: Cr; [IA]; PMBC Eviota pellucida Larson, 1976: Cr; [WCP] Eviota prasina (Klunzinger, 1871): Cr; [IWP]; PMBC Eviota prasites Jordan & Seale, 1906: Cr; [IP] Eviota punctulata Jewett & Lachner, 1983: Cr; [WCP]; PMBC Eviota queenslandica Whitley, 1932: Cr; [WP]; PMBC Eviota rw-th sp.1: Cr Eviota rw-th sp.2: Cr Eviota rw-th sp.4: Cr Eviota sebreei Jordan & Seale, 1906: Cr; [IP]; PMBC Eviota sigillata Jewett & Lachner, 1983: Cr; [IWP]; PMBC Eviota spilota Lachner & Karnella, 1980: Cr; [WP]; PMBC Eviota zebrina Lachner & Karnella, 1978: Cr: [IWP]; PMBC Eviota zonura Jordan & Seale, 1906: Cr; [WCP] Exyrias bellissimus (Smith, 1959): Cr; [IWP]; PMBC Exyrias puntang (Bleeker, 1851): Mg; [EIO-WP] Favonigobius melanobranchus (Fowler, 1934): Cr, Mg, Sg; [IWP]; PMBC Favonigobius reichei (Bleeker, 1853): Cr, Mg, Sg; [IWP]; PMBC Flabelligobius latruncularius (Klausewitz, 1974): Cr; [IO] Flabelligobius russus (Cantor, 1849): OShr; [IM]; PMBC Fusigobius dfh sp.4: Cr Fusigobius duospilus Hoese & Reader, 1985: Cr; [IP]; PMBC Fusigobius humeralis (Randall, 2001): Cr; [IP] Fusigobius inflamaculatus Randall, 1994: Cr; [IWP] Fusigobius maximus (Randall, 2001): Cr; [IWP] Fusigobius melacron (Randall, 2001): Cr; [WP] Fusigobius neophytus (Günther, 1877): Cr; [IP]; PMBC Fusigobius pallidus (Randall, 2001): Cr; [IWP] Fusigobius signipinnis Hoese & Obika, 1988: Cr; [WP] Gladiogobius rex Shibukawa & Allen, 2007: Cr; [IO]; PMBC Glossogobius bicirrhosus (Weber, 1894): Mg; [WP]; PMBC Glossogobius circumspectus (Macleay, 1883): Mg; [WP]; PMBC Glossogobius guiris (Hamilton, 1822): Mg; [IWP] Glossogobius sparsipapillus Akihito & Meguro, 1976: Mg; [WP]; PMBC Gnatholepis anjerensis (Bleeker, 1850): Cr; [IP]; PMBC Gnatholepis cauerensis (Bleeker, 1853): Cr; [IP] Gobiodon citrinus (Rüppell, 1853): Cr; [IWP] Gobiodon dfh sp.11: Cr Gobiodon erythrospilus Bleeker, 1857: Cr; [WP] Gobiodon histrio (Valenciennes, 1837): Cr; [IWP]; PMBC Gobiodon prolixus Winterbottom & Harold, 2005: Cr; [EIO-WP] Gobiodon quinquestrigatus (Cuvier & Valenciennes, 1837): Cr; [WCP] Gobiodon rivulatus (Rüppell, 1828): Cr; [IWP]; PMBC Gobiodon rw-th sp.1: Cr Gobiodon rw-th sp.2: Cr Gobiodon rw-th sp.3: Cr Gobiopsis aporia Lachner & McKinney, 1978: Cr; [EIOWP]; PMBC Gobiopsis macrostoma Steindachner, 1861: Mg; [NIOIM]; PMBC Gobiopsis quinquecincta (Smith, 1931): Cr; [EIO-WP]; PMBC Gobiopterus brachypterus (Bleeker, 1855): Mg; [IA]; PMBC Gobiopterus panayensis (Herre, 1944): Mg; [WP]; PMBC Gobiopterus rw-th sp.1: Mg Hemigobius hoevenii (Bleeker, 1851): Mg; [IA] Hemigobius mingi Herre, 1936: Mg; [IM] Istigobius decoratus (Herre, 1927): Cr; [IWP]; PMBC Istigobius diadema (Steindachner, 1877): Cr, Mg; [EIOIA]; PMBC 67 The fishes of southwestern Thailand Istigobius goldmanni (Bleeker, 1852): Cr; [WP]; PMBC Istigobius ornatus (Rüppell, 1830): Cr; [IP]; PMBC Istigobius rigilius (Herre, 1953): Cr; [WCP] Lophogobius bleekeri Popta, 1921: Mg; [IM]; PMBC Mahidolia mystacina (Valenciennes, 1837): Cr, Sg, OShr; [IWP]; PMBC Mangarinus seshaiyai (Jacop & Rangarajan, 1960): Mg; [EIO] Mangarinus waterousi Herre, 1943: Mg; [WP]; PMBC Mugilogobius chulae (Smith, 1932): Mg; [IM]; PMBC Mugilogobius fasciatus Larson, 2001: Mg; [IM (presently known from west coast of Thailand and Singapore)]; PMBC Mugilogobius mertoni (Weber, 1911): Mg; [IWP] Mugilogobius tigrinus Larson, 2001: Mg; [IM (presently known from west coast of Thailand, Singapore and Gulf of Thailand)]; PMBC Myersina adonis Shibukawa & Satapoomin, 2006; Mg, Sg; [IM (presently known from west coast of Thailand and Singapore]; PMBC Myersina crocatus (Wongratana, 1975): Cr, OShr; [IM]; PMBC Myersina filifer (Cuvier & Valenciennes, 1837): OShr; [IWP]; PMBC Myersina yangii (Chen, 1960): Cr, OShr; [WP]; PMBC Oligolepis acutipennis (Valenciennes, 1837): Mg; [IWP] Oplopomus caninoides (Bleeker, 1852): Cr, OShr; [EIOWP]; PMBC Oplopomus oplopomus (Valenciennes, 1837): Cr, OShr; [IP]; PMBC Oxyurichthys microlepis (Bleeker, 1849): Mg, Sg; [IWP]; PMBC Oxyurichthys ophthalmonema (Bleeker, 1856): Mg, Sg; [EIO-WP] Oxyurichthys papuensis (Valenciennes, 1837): Cr, OShr; [IWP]; PMBC Oxyurichthys tentacularis (Valenciennes, 1837): Mg, Sg; [WP]; PMBC Palutrus scapulopunctatus (De Beaufort, 1912): Cr; [IWP]; PMBC Pandaka lidwilli (McCulloch, 1917): Mg; [WP]; PMBC Pandaka pygmaea Herre, 1927: Mg; [IM] Parachaeturichthys polynema (Bleeker, 1853): Mg; [IWP]; PMBC Paragobiodon modestus (Regan, 1908): Cr; [IP]; PMBC Paratrypauchen microcephalus (Bleeker, 1860): Mg, Sg; [IWP] Periophthalmus argentilineatus Valenciennes, 1837: Mg; [IWP]; PMBC Periophthalmus chrysospilos Bleeker, 1852: Mg; [EIOIM]; PMBC Periophthalmus kalolo Lesson, 1830: Mg; [IWP]; PMBC Periophthalmus minutus Eggert, 1935: Mg; [IA]; PMBC Periophthalmus novemradiatus (Hamilton, 1822): Mg; [EIO-IA]; PMBC Periophthalmus septemradiatus (Hamilton, 1822): Mg; [EIO-IM] Periophthalmus walailakae Darumas & Tantochodok, 2002: Mg; [IM (presently known from west coast of Thailand and Singapore)]; PMBC Pleurosicya bilobata (Koumans, 1941): Sg; [IWP] Pleurosicya coerulea Larson, 1990: Cr; [IP] Pleurosicya fringilla Larson, 1990: Cr; [IWP]; PMBC Pleurosicya labiata (Weber, 1913): Cr; [EIO-IA]; PMBC Pleurosicya micheli Fourmanoir, 1971: Cr; [IP] Pleurosicya mossambica Smith, 1959: Cr; [IP]; PMBC Pleurosicya plicata Larson, 1990: Cr; [IWP] Pleurosicya prognatha Goren, 1984: Cr; [IWP] Priolepis cinctus (Regan, 1908): Cr; [IWP]; PMBC Priolepis inhaca (Smith, 1949): Cr; [IWP] Priolepis nuchifasciatus (Günther, 1873): Cr; [IA] Priolepis semidoliatus (Valenciennes, 1837): Cr; [IP]; PMBC Psammogobius biocellatus (Valenciennes, 1837): Mg, Sg; [IWP]; PMBC Pseudapocryptes lanceolatus (Bloch & Schneider, 1801): Mg; [EIO-IM] Pseudogobius javanicus (Bleeker, 1856): Mg; [IM]; PMBC Redigobius bikolanus Herre, 1927: Mg; [WP]; PMBC Redigobius chrysosoma (Bleeker, 1857): Mg; [IA]; PMBC Scartelaos cantoris (Day, 1871): Mg, Sg; [AS]; PMBC Scartelaos histophorus (Valenciennes, 1837): Mg, Sg; [NIOWP]; PMBC Silhouettea sp.1: Cr, Sg; PMBC Stigmatogobius pleurostigma (Bleeker, 1849): Mg; [IM] Stigmatogobius sandanundio (Hamilton, 1822): Mg; [EIO]; PMBC Stonogobiops nematodes Hoese & Randall, 1982: Cr; [IA] Sueviota lachneri Winterbottom & Hoese, 1988: Cr; [IWP]; PMBC Taenioides cirratus (Blyth, 1860): Mg, Sg, OShr; [IWP]; PMBC Taenioides gracilis (Valenciennes, 1837): OShr; [IWP]; PMBC Trimma emeryi Winterbottom, 1985: Cr; [IO] Trimma flammeum (Smith, 1959): Cr; [IO]; PMBC Trimma fucatum Winterbottom & Southcott, 2007: Cr; [AS]; PMBC Trimma naudei Smith, 1956: Cr; [IWP]; PMBC Trimma okinawae (Aoyaki, 1949): Cr; [WCP]; PMBC Trimma sanguinellus Winterbottom & Southcott, 2007: Cr; [AS]; PMBC Trimma striata (Herre, 1945): Cr; [EIO-WP]; PMBC Trimma talori Lobel, 1979: Cr; [IP] Trimma winterbottomi Randall & Downing, 1994: Cr; [IO]; PMBC Trypauchen vagina (Bloch & Schneider, 1801): OShr; [NIOIM]; PMBC Valenciennea immaculata (Ni, 1981): Sg; [WP] Valenciennea limicola Hoese & Larson, 1994: Cr; [WP]; PMBC Valenciennea mularis (Valenciennes, 1837): Cr, Sg, OShr; [IA] Valenciennea parva Hoese & Larson, 1994: Cr; [IWP] 68 Phuket mar. biol. Cent. Res. Bull. Valenciennea puellaris (Tomiyama, 1956): Cr, OShr; [IP]; PMBC Valenciennea randalli Hoese & Larson, 1994: Cr, OShr; [WP]; PMBC Valenciennea sexguttata (Valenciennes, 1837): Cr, OShr; [IP]; PMBC Valenciennea strigata (Broussonet, 1782): Cr; [IP]; PMBC Valenciennea wardii (Playfair, 1867): Cr, OShr; [IWP]; PMBC Vanderhorstia ambanoro (Fourmanoir, 1957): Cr, OShr; [IWP]; PMBC Vanderhorstia ornatissima Smith, 1959: Cr; [IP] Yongeichthys nebulosus (Forsskål, 1775): Cr, Mg, Sg, OShr; [IWP]; PMBC Microdesmidae Gunnellichthys curiosus Dawson, 1968: Cr; [IP] Gunnellichthys viridescens Dawson, 1968: Cr; [IP] Ptereleotridae Nemateleotris decora Randall & Allen, 1973: Cr; [IWP] Nemateleotris magnifica Fowler, 1928: Cr; [IP] Oxymetopon compressus Chan, 1966: Cr, OShr; [WP]; PMBC Parioglossus formosus (Smith, 1931): Cr; [WP]; PMBC Parioglossus palustris (Herre, 1945): Mg; [WP] Parioglossus philippinus (Herre, 1940): Cr; [IWP]; PMBC Ptereleotris arabica Randall & Hoese, 1985: Cr, OShr; [IO]; PMBC Ptereleotris evides (Jordan & Hubbs, 1925): Cr; [IP] Ptereleotris heteroptera (Bleeker, 1855): Cr; [IP]; PMBC Ptereleotris microlepis (Bleeker, 1856): Cr; [IP]; PMBC Ptereleotris monoptera (Bleeker, 1855): Cr; [IP] Ptereleotris zebra (Fowler, 1938): Cr; [IP] Kurtidae Kurtus indicus Bloch, 1786: OShr; [EIO-IM]; PMBC Ephippidae Ephippus orbis (Bloch, 1787): Mg, Sg, OShr; [IWP]; PMBC Platax boersi Bleeker, 1852: Cr; [IP] Platax orbicularis (Forsskål, 1775): Cr, Mg, Sg, OShr; [IP]; PMBC Platax pinnatus (Linnaeus, 1758): Cr, OShr; [WP]; PMBC Platax teira (Forsskål, 1775): Cr, OShr; [IWP]; PMBC Scatophagidae Scatophagus argus (Linnaeus, 1758): Mg, Sg; [NIO-WP]; PMBC Siganidae Siganus argenteus (Quoy & Gaimard, 1825): Cr; [IP]; PMBC Siganus canaliculatus (Park, 1797): Cr, Mg, Sg, OShr; [NIOIA]; PMBC Siganus corallinus (Valenciennes, 1835): Cr; [IWP]; PMBC Siganus fuscescens (Houttuyn, 1782): Cr; [WP]; PMBC Siganus guttatus (Bloch, 1787): Cr, Mg, Sg, OShr; [IM] Siganus javus (Linnaeus, 1766): Cr, Mg, Sg, OShr; [NIOWP] Siganus lineatus (Valenciennes, 1835): Cr; [EIO-WP] Siganus magnificus (Burgess, 1977): Cr; [EEIO]; PMBC Siganus puelloides Woodland & Randall, 1979: Cr; [EIO]; PMBC Siganus stellatus laqueus Von Bonde, 1934: Cr, OShr; [IO]; PMBC Siganus vermiculatus (Valenciennes, 1835): Cr, Mg, Sg, OShr; [EIO-WP]; PMBC Siganus virgatus (Valenciennes, 1835): Cr, Sg, OShr; [EIOWP]; PMBC Zanclidae Zanclus cornutus (Linnaeus, 1758): Cr; [IP]; PMBC Acanthuridae Acanthurus bariene Lesson, 1830: Cr; [WP-EIO]; PMBC Acanthurus blochii Valenciennes, 1835: Cr; [IP] Acanthurus guttatus Forster, 1801: Cr; [IP] Acanthurus leucocheilus Herre, 1927: Cr; [IWP] Acanthurus leucosternon Bennett, 1832: Cr; [IO]; PMBC Acanthurus lineatus (Linnaeus, 1758): Cr; [IP]; PMBC Acanthurus maculiceps (Ahl, 1923): Cr; [EIO-WCP] Acanthurus mata (Cuvier, 1829): Cr; [IP]; PMBC Acanthurus nigricauda Duncker & Mohr, 1929: Cr; [IP]; PMBC Acanthurus nigrofuscus (Forsskål, 1775): Cr; [IP] Acanthurus olivaceus Forster, 1801: Cr; [WCP] Acanthurus tennenti Günther, 1861: Cr; [IO] Acanthurus thompsoni (Fowler, 1923): Cr; [IP]; PMBC Acanthurus triostegus (Linnaeus, 1758): Cr; [IP]; PMBC Acanthurus tristis Tickell In Day, 1888: Cr; [IO]; PMBC Acanthurus xanthopterus Valenciennes, 1835: Cr; [IP]; PMBC Ctenochaetus binotatus Randall, 1955: Cr; [IP]; PMBC Ctenochaetus striatus (Quoy & Gaimard, 1825): Cr; [IP]; PMBC Ctenochaetus truncatus Randall & Clements, 2001: Cr; [IO]; PMBC Naso brachycentron (Valenciennes, 1835): Cr; [IP] Naso brevirostris (Valenciennes, 1835): Cr; [IP]; PMBC Naso caeruleocauda Randall, 1994: Cr; [IA] Naso elegans (Rüppell, 1829): Cr; [IO]; PMBC Naso hexacanthus (Bleeker, 1855): Cr; [IP]; PMBC Naso lopezi Herre, 1927: Cr; [WP] Naso mcdadei Johnson, 2002: Cr; [IWP] Naso thynnoides (Valenciennes, 1835): Cr; [IP] Naso unicornis (Forsskål, 1775): Cr; [IP]; PMBC Naso vlamingii (Valenciennes, 1835): Cr; [IP]; PMBC Paracanthurus hepatus (Linnaeus, 1766): Cr; [IP] Zebrasoma desjardinii (Bennett, 1835): Cr; [IO]; PMBC Zebrasoma scopas (Cuvier, 1829): Cr; [IP]; PMBC Sphyraenidae Sphyraena baracuda (Walbaum, 1792): Cr, Sg, OShr; [COS]; PMBC Sphyraena forsteri Cuvier, 1829: Cr, OShr; [IP]; PMBC Sphyraena jello Cuvier, 1829: Cr, Mg, Sg, OShr; [IP]; PMBC Sphyraena obtusata Cuvier, 1829: Cr, Sg, OShr; [IWP]; PMBC 69 The fishes of southwestern Thailand Sphyraena pinguis Günther, 1874: Cr; [IWP] Sphyraena putnamae Jordan & Seale, 1905: Mg, Sg, OShr; [IWP] Sphyraena qenie Klunzinger, 1870: Cr; [IP] Gempylidae Gempylus serpens Cuvier, 1829: OShr; [COS] Neoepinnula orientalis (Gilchrist & von Bonde, 1924): OShr; [IWP]; PMBC Nesiarchus nasutus Johnson, 1862: OShr; [COS]; PMBC Promethichthys prometheus (Cuvier, 1832): OShr; [COS] Rexea bengalensis (Alcock, 1884): OShr; [IWP] Thyrsitoides marleyi Fowler, 1929: OShr; [IWP] Trichiuridae Eupleurogrammus muticus (Gray, 1831): OShr; [NIO-IM] Tentoriceps cristatus (Klunzinger, 1884): OShr; [IWP]; PMBC Trichiurus gangeticus Gupta, 1966: OShr; [EIO]; PMBC Trichiurus lepturus Linnaeus, 1758: Mg, Sg, OShr; [COS]; PMBC Scombridae Acanthocybium solandi (Cuvier, 1831): Cr, OShr; [COS] Auxis rochei rochei (Risso, 1810): OShr; [COS]; PMBC Auxis thazard thazard (Lacepède, 1800): OShr; [COS]; PMBC Euthynnus affinis (Cantor, 1849): Cr, OShr; [IP]; PMBC Grammatorcynos bilineatus (Rüppell, 1836): Cr, OShr; [IP]; PMBC Gymnosarda unicolor (Rüppell, 1838): Cr, OShr; [IP]; PMBC Katsuwonus pelamis (Linnaeus, 1758): OShr; [COS]; PMBC Rastrelliger brachysoma (Bleeker, 1851): OShr; [WP]; PMBC Rastrelliger faughni Matsui, 1967: OShr; [EIO-WP]; PMBC Rastrelliger kanagurta (Cuvier, 1817): Cr, Mg, Sg, OShr; [IP]; PMBC Scomberomorus commerson (Lacepède, 1800): Cr, Mg, OShr; [IP]; PMBC Scomberomorus guttatus (Bloch & Schneider, 1801): OShr; [NIO-WP]; PMBC Scomberomorus lineolatus (Cuvier, 1831): OShr; [NIOIM] Thunnus alalunga (Bonnaterre, 1788): OShr; [COS] Thunnus albacares (Bonnaterre, 1788): OShr; [COS]; PMBC Thunnus obesus (Lowe, 1839): OShr; [COS] Thunnus tonggol (Bleeker, 1851): OShr; [IWP]; PMBC Xiphiidae Xiphias gladius Linnaeus, 1758: OShr; [COS] Istiophoridae Istiophorus platypterus (Shaw & Nodder, 1792): OShr; [IP]; PMBC Makaira indica (Cuvier, 1832): OShr; [IP] Tetrapturus audax (Philippi, 1887): OShr; [IP] Centrolophidae Psenopsis obscura Headrich, 1967: OShr; [IM] Nomeidae Cubiceps pauciradiatus Günther, 1872: OShr; [CIR]; PMBC Cubiceps whiteleggii (Waite, 1894): OShr; [IWP] Ariommatidae Ariomma indicum (Day, 1870): OShr; [IWP]; PMBC Stromateidae Pampus argenteus (Euphrasen, 1788): OShr; [NIO-WP]; PMBC Pampus chinensis (Euphrasen, 1788): OShr; [NIO-WP] Caproidae Antigonia rubescens (Günther, 1860): OShr; [WP] Order Pleuronectiformes Psettodidae Psettodes erumei (Bloch & Schneider, 1801): Mg, OShr; [IWP]; PMBC Citharidae Brachypleura novaezeelandiae Günther, 1862: OShr; [NIO-IA]; PMBC Paralichthyidae Pseudorhombus argus Weber, 1913: OShr; [IA] Pseudorhombus arsius (Hamilton, 1822): Mg, Sg, OShr; [IWP]; PMBC Pseudorhombus dupliciocellatus Regan, 1905: OShr; [IA]; PMBC Pseudorhombus elevatus Ogilby, 1912: Mg, OShr; [IWP]; PMBC Pseudorhombus malayanus Bleeker, 1866: Mg, OShr; [NIO-IM] Pseudorhombus quinquocellatus Weber & de Beaufort, 1929: OShr; [EIO-IA]; PMBC Pseudorhombus triocellatus (Schneider, 1801): OShr; [EIOIA]; PMBC Bothidae Arnoglossus aspilos (Bleeker, 1851): Cr; [NIO-IA]; PMBC Arnoglossus macrolophus Alcock, 1889: OShr; [IWP] Arnoglossus tapeinosoma (Bleeker, 1865): OShr; [NIOWP] Arnoglossus tenuis Günther, 1880: OShr; [WP]; PMBC Asterorhombus intermedius (Bleeker, 1866): OShr; [IWP] Bothus pantherinus (Rüppell, 1830): OShr; [IP] Chascanopsetta lugubris Alcock, 1894: OShr; [IWP] Engyprosopon grandisquama (Temminck & Schlegel, 1846): OShr; [IWP]; PMBC Engyprosopon latifrons (Regan, 1908): OShr; [IWP]; PMBC Engyprosopon maldivensis (Regan, 1908): OShr; [EIO-WP] Grammatobothus polyophthalmus (Bleeker, 1866): OShr; [EIO-WP]; PMBC Laeops guentheri Alcock, 1890: OShr; [NIO-IM] Laeops kitaharae (Smith & Pope, 1906): OShr; [IWP] Poecilopsettidae Poecilopsetta colorata Günther, 1880: OShr; [EIO-IM] Samaridae Samaris cristatus Gray, 1831: OShr; [IWP]; PMBC 70 Phuket mar. biol. Cent. Res. Bull. Soleidae Aesopia cornuta Kaup, 1858: OShr; [IWP]; PMBC Aseraggodes dubius Weber, 1913: OShr; [WP] Aseraggodes satapoomini Randall & Desoutter-Meniger, 2007: Cr; [AS] Brachirus pan (Hamilton, 1822): OShr; [EIO-IM] Heteromycteris oculus (Alcock, 1889): OShr; [IWP]; PMBC Liachirus melanospilos (Bleeker, 1854): OShr; [EIO-WP] Pardachirus pavoninus (Lacepède, 1802): Cr, Sg, OShr; [EIO-WP]; PMBC Solea ovata Richardson, 1846: Sg, OShr; [EIO-IM]; PMBC Synaptura commersonii (Lacepède, 1802): Sg, OShr; [NIOIM]; PMBC Zebrias altipinnis (Alcock, 1890): OShr; [EIO] Zebrias quagga Kaup, 1858: Mg, Sg, OShr; [IWP]; PMBC Zebrias synapturoides (Jenkins, 1910): OShr; [NIO-WP] Cynoglossidae Cynoglossus arel (Schneider, 1801): Mg, OShr; [NIO-IM] Cynoglossus bilineatus (Lacepède, 1802): Mg, OShr; [NIOWP]; PMBC Cynoglossus cynoglossus (Hamilton, 1822): OShr; [NIOIM]; PMBC Cynoglossus kopsi (Bleeker, 1851): OShr; [NIO-WP] Cynoglossus lida (Bleeker, 1851): Mg, Sg, OShr; [IWP]; PMBC Cynoglossus lingua Hamilton, 1822: Mg, OShr; [IWP] Cynoglossus macrolepidotus (Bleeker, 1851): OShr; [WP]; PMBC Cynoglossus monopus (Bleeker, 1849): OShr; [EIO-IM] Cynoglossus oligolepis (Bleeker, 1854): OShr; [WP]; PMBC Cynoglossus puncticeps (Richardson, 1846): Sg, OShr; [NIO-IA]; PMBC Cynoglossus semifasciatus Day 1877: Mg, Sg, OShr; [EIO]; PMBC Cynoglossus sibogae Weber, 1913: OShr; [IM]; PMBC Paraplagusia bilineata (Bloch, 1787): OShr; [IWP] Paraplagusia blochii (Bleeker, 1851): OShr; [NIO-WP] Symphurus septemstriatus (Alcock, 1891): OShr; [EIOIM] Order Tetraodontiformes Triacanthodidae Halimochirurgus alcocki Weber, 1913: OShr; [IWP] Tydemania navigatoris Weber, 1913: OShr; [IWP]; PMBC Triacanthidae Pseudotriacanthus strigilifer (Cantor, 1849): OShr; [NIOIA]; PMBC Triacanthus biaculeatus (Bloch, 1786): Mg, Sg, OShr; [NIO-WP]; PMBC Triacanthus nieuhofi Bleeker, 1852: Sg, OShr; [EIO-IA] Trixiphichthys weberi (Chaudhuri, 1910): OShr; [EIO-IA]; PMBC Balistidae Abalistes stellatus (Anonymous, 1798): Cr, OShr; [IWP]; PMBC Balistapus undulatus (Park, 1797): Cr, OShr; [IP]; PMBC Balistoides conspicillum (Bloch & Schneider, 1801): Cr, OShr; [IP]; PMBC Balistoides viridescens (Bloch & Schneider, 1801): Cr, OShr; [IP]; PMBC Canthidermis maculatus (Bloch, 1786): OShr; [COS]; PMBC Canthidermis rotundatus (Marion de Procé, 1822): OShr; [IWP]; PMBC Melichthys indicus Randall & Klausewitz, 1973: Cr, OShr; [IO]; PMBC Melichthys vidua (Solander, 1844): Cr; [IP] Odonus niger (Rüppell, 1837): Cr, OShr; [IP]; PMBC Pseudobalistes flavimarginatus (Rüppell, 1829): Cr, OShr; [IP] Pseudobalistes fuscus (Bloch & Schneider, 1801): Cr, OShr; [IP]; PMBC Rhinecanthus aculeatus (Linnaeus, 1758): Cr, OShr; [IP] Rhinecanthus rectangulus (Bloch & Schneider, 1801): Cr, OShr; [IP] Rhinecanthus verrucosus (Linnaeus, 1758): Cr, OShr; [IWP] Sufflamen bursa (Bloch & Schneider, 1801): Cr; [IP] Sufflamen chrysopterum (Bloch & Schneider, 1801): Cr, OShr; [IWP]; PMBC Sufflamen fraenatum (Latreille, 1804): Cr, OShr; [IP]; PMBC Xanthichthys auromarginatus (Bennett, 1831): Cr, OShr; [IP] Monacanthidae Acreichthys tomentosus (Linnaeus, 1758): Sg; [IA] Aluterus monoceros (Linnaeus, 1758): OShr; [CIR]; PMBC Aluterus scriptus (Osbeck, 1765): Cr, OShr; [CIR]; PMBC Amanses scopas (Cuvier, 1829): Cr; [IP]; PMBC Anacanthus barbatus Gray, 1831: Sg, OShr; [NIO-IA] Cantherhines dumerilli (Hollard, 1854): Cr, OShr; [IP] Cantherhines multilineatus (Tanaka, 1918): OShr; [IWP; PMBC Cantherhines pardalis (Rüppell, 1837): Cr, OShr; [IP]; PMBC Monacanthus chinensis Osbeck, 1765: Cr, Mg, Sg, OShr; [WP]; PMBC Oxymonacanthus longirostris (Bloch & Schneider, 1801): Cr; [IP]; PMBC Paraluteres sp.1: Cr; [AS] Paramonacanthus japonicus (Tilesius, 1810): OShr; [EIOWP] Paramonacanthus tricuspis (Hollard, 1854): Mg, Sg, OShr; [EIO]; PMBC Pseudomonacanthus macrurus (Bleeker, 1857): Mg, Sg, OShr; [IWP]; PMBC Pseudomonacanthus peroni (Hollard, 1854): OShr; [IA] Ostraciidae Lactoria cornuta (Linnaeus, 1758): Cr, Mg, Sg, OShr; [IP]; PMBC Lactoria diaphana (Bloch & Schneider, 1801): Mg, OShr; [IP] 71 The fishes of southwestern Thailand Ostracion cubicus Linnaeus, 1758: Cr, Mg, Sg, OShr; [IP]; PMBC Ostracion meleagris Shaw, 1796: Cr; [IP]; PMBC Ostracion nasus Bloch, 1785: OShr; [EIO-WP]; PMBC Ostracion rhinorhynchus Bleeker, 1852: Cr, OShr; [IWP] Ostracion solorensis Bleeker, 1853: Cr; [IWP] Tetrosomus gibbosus (Linnaeus, 1758): Cr, OShr; [IWP]; PMBC Tetrosomus reipublicae (Whitley, 1930): OShr; [IWP] Tetraodontidae Amblyrhynchotes honckenii (Bloch, 1795): OShr; [IWP] Arothron caeruleopunctatus Matsuura, 1994: Cr, Mg, Sg; [EIO-WP] Arothron hispidus (Linnaeus, 1758): Cr, Mg, Sg, OShr; [IP]; PMBC Arothron immaculatus (Bloch & Schneider, 1801): Cr, OShr; [IWP]; PMBC Arothron manilensis (Marion de Porcé, 1822): Cr, Sg, OShr; [WCP] Arothron mappa (Lesson, 1830): Cr, OShr; [IP]; PMBC Arothron nigropunctatus (Bloch & Schneider, 1801): Cr, OShr; [IP]; PMBC Arothron reticularis (Bloch & Schneider, 1801): Mg, OShr; [IWP]; PMBC Arothron stellatus (Bloch & Schneider, 1801): Cr, OShr; [IP]; PMBC Canthigaster smithae Allen & Randall, 1977: Cr; [IO] Canthigaster solandri (Richardson, 1844): Cr, OShr; [IP]; PMBC Chelonodon patoca (Hamilton, 1822): Mg, Sg, OShr; [IWP]; PMBC Lagocephalus gloveri Abe & Tabeta, 1983: OShr; [IWP] Lagocephalus inermis (Temminck & Schlegel, 1844): OShr; [IWP]; PMBC Lagocephalus lunaris (Bloch & Schneider, 1801): Mg, Sg, OShr; [IWP]; PMBC Lagocephalus scleratus (Gmelin, 1789): OShr; [IWP]; PMBC Lagocephalus spadiceus (Richardson, 1844): OShr; [IWP] Takifugu oblongus (Bloch, 1786): Mg, Sg, OShr; [IWP]; PMBC Tetraodon fluviatilis Hamilton, 1822: OShr; [EIO-IM]; PMBC Tetraodon nigroviridis Procé, 1822: Mg, Sg, OShr; [EIOWP]; PMBC Torquigener hypselogeneion (Bleeker, 1852): Cr; [IWP]; PMBC Diodontidae Cyclichthys orbicularis (Bloch, 1785): Mg, Sg, OShr; [IWP]; PMBC Diodon eydouxii Brisout de Barneville, 1846: OShr; [IWP] Diodon holocanthus Linnaeus, 1758: OShr; [CIR]; PMBC Diodon hystrix Linnaeus, 1758: Cr, OShr; [CIR]; PMBC Diodon liturosus Shaw, 1804: Cr, Sg, OShr; [IP]; PMBC Molidae Masturus lanceolatus (Lienard, 1840): OShr; [COS] Mola mola (Linnaeus, 1758): OShr; [COS]; PMBC 72 Phuket mar. biol. Cent. Res. Bull. Appendix B Selected taxonomic literature pertaining to the records of fishes from southwestern Thailand, the Andaman Sea Class Myxini Wongratana, T. 1983. Eptatretus indrambaryai, a new species of hagfish (Myxinidae) from the Andaman Sea. Nat Hist. Bull. Siam Soc. 31(2): 139–150. Class Chondrichthyes Compagno, L.J.V. and P.R. Last. 2008. A new species of wedgefish, Rhynchobatus palpebratus sp. nov. (Rhynchobatoidei: Rhynchobatidae), from the Indo-West Pacific. In: P.R. Last, W.T. White and J.J. Pogonoski (eds.). Descriptions of New Australian Chondrichthyans. CISRO Marine and Atmospheric Research Paper No. 022. pp. 227–240. Krajangdara, T., S. Puteeka, R. Phoonsawat, S. Vibunpant, R. Sojittosakul and R. Chayakul. 2006. Sharks found in Thai waters and adjacent areas. Marine Fisheries Research and Development Bureau, Department of Fisheries. Technical Paper No. 17/2006. 99 p. (in Thai) Monkolprasit, S. 1984. The cartilaginous fishes (Class Elasmobranchii) found in Thai waters and adjacent areas. Department of Fishery Biology, Faculty of Fisheries, Kasetsart University, Bangkok, Thailand. 175 p. Class Actinopterygii Order Albuliformes Hidaka, K., Y. Iwatsuki and J.E. Randall. 2008. A review of the Indo-Pacific bonefishes of the Albula argentea complex, with a description of a new species. Ichthyol. Res. 55(1): 53–64. Order Anguilliformes McCosker, J.E. 1972. Two new genera and two new species of western Pacific snake-eels (Apodes: Ophichthidae). Proc. Cal. Acad. Sci. 39(10): 111–119. McCosker, J.E. and J.E. Randall. 2001. Revision of the snake-eel genus Brachysomophis (Anguilliformes: Ophichthidae), with descriptions of two new species and comments on the species of Mystriophis. Indo-Pacific Fishes 33: 1– 32, 2 pls. Böhlke, E.B. 2000. Notes on the identity of small, brown, unpatterned Indo-Pacific moray eels, with descriptions of three new species (Anguilliformes: Muraenidae). Pac. Sci. 54(4): 395–416. Böhlke, E.B. and J.E. Randall. 2000. A review of the moray eels (Anguilliformes; Muraenidae) of the Hawaiian Islands, with descriptions of two new species. Proc. Acad. nat. Sci. Philad. 150: 203–278, 9 pls. Order Clupeiformes Whitehead, P.J.P. 1985. FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies, and wolf-herrings. Part 1 – Chirocentridae, Clupeidae and Pristigasteridae. FAO Species Synopsis No. 125, Vol. 7, Pt. 1: 1–303. Whitehead, P.J.P., G.J. Nelson and T. Wongratana. 1988. FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies, and wolf-herrings. Part 2 – Engraulididae. FAO Species Synopsis No. 125, Vol. 7, Pt. 2: 305– 579. Order Ophidiiformes Markle, D.F. and J.E. Olney. 1990. Systematics of the pearlfishes (Pisces: Carapidae). Bull. Mar. Sci. 47(2): 269–410. Møller, P.R. and W. Schwarzhans. 2008. Review of the Dinematichthyini (Teleostei: Bythitidae) of the Indo-west Pacific. Part IV. Dinematichthys and two new genera with descriptions of nine new species. The Beagle, Records of the Museum of Art Galleries of the Northern Territory 24: 87–146. Nielsen, J.G. and Y. Machida. 1988. Revision of the Indo-Pacific bathyal fish genus Glyptophidium (Ophidiiformes, Ophidiidae). Jap. J. Ichthyol. 35(3): 289–319. Order Lophiiformes Bradbury, M.G. 1988. Rare fishes of the deep-sea genus Halieutopsis: a review with descriptions of four new species (Lophiiformes: Ogcocephalidae). Fieldiana, Zool. 44: 1–22. Order Atheriniformes Kimura, S., D. Golani, Y. Iwatsuki, M. Tabuchi and T. Yoshino. 2007. Redescriptions of the Indo-Pacific atherinid fishes Atherinomorus forskalii, Atherinomorus lacunosus, and Atherinomorus pinguis. Ichthyol. Res. 54(2): 145– 159. Parenti, L.R. 1989. A phylogenetic revision of the phallostethid fishes (Atherinomorpha, Phallostethidae). Proc. Cal. Acad. Sci. 46(11): 243–277. Order Beloniformes 73 The fishes of southwestern Thailand Parenti, L.R. 2008. A phylogenetic analysis and taxonomic revision of ricefishes, Oryzias and relatives (Beloniformes, Andrianichthyidae). Zool. J. Linn. Soc. 154: 494–610. Order Beryciformes Randall, J.E. 1998. Revision of the Indo-Pacific squirrelfishes (Beryciformes: Holocentridae: Holocentrinae) of the genus Sargocentron, with descriptions of four new species. Indo-Pacific Fishes 27: 1–105, 11 pls. Order Gasterosteiformes Dawson, C.E. 1985. Indo-Pacific Pipefishes (Red Sea to the Americas). The Gulf Coast Research Laboratory, Ocean Springs, Mississippi. 230 p. Order Scorpaeniformes Imamura, H. 1996. Phylogeny of the family Platycephalidae and related taxa (Pisces: Scorpaeniformes). Species Diversity 1(2): 123–233. Knapp, L.W. and T. Wongratana. 1987. Sorsogona melanoptera, a new flathead fish from the northern Indian Ocean (Teleostei: Platycephalidae). Proc. Biol. Soc. Wash. 100(2): 381–385. Poss, S.G. and W.N. Eschmeyer. 1980. Xenaploactis, a new genus for Prosopodasys asperrimus Günther (Pisces: Aploactinidae), with descriptions of two new species. Proc. Cal. Acad. Sci. 42(8): 287–293. Randall, J.E. and W.N. Eschmeyer. 2001. Revision of the Indo-Pacific scorpionfish genus Scorpaenopsis, with descriptions of eight new species. Indo-Pacific Fishes 34: 1–79, 12 pls. Wongratana, T. 1975. A record of a flathead fish, Thysanophrys papillolabium Schultz, in the Andaman Sea, with a key to fourteen species of the Platycephalidae from Thai waters (Pisces: Platycephalidae). Phuket mar. biol. Cent. Res. Bull. 7: 1–9. Order Perciformes Family Acropomatidae Okamoto, M. and H. Ida. 2002. Acropoma argentistigma, a new species from the Andaman Sea, off southern Thailand (Perciformes: Acropomatidae). Ichthyol. Res. 49(3): 281–285. Family Serranidae Randall, J.E. and C.C. Baldwin. 1997. Revision of the serranid fishes of the subtribe Pseudogrammina, with descriptions of five new species. Indo-Pacific Fishes 26: 1–56, 1 pl. Randall, J.E. and P.C. Heemstra. 1991. Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pacific Fishes 20: 1–332, 41 pls. Randall, J.E. and P.C. Heemstra. 2007. Meganthias filiferus, a new species of anthiine fish (Perciformes: Serranidae), from the Andaman Sea off southwestern Thailand. Phuket mar. biol. Cent. Res. Bull. 68: 5–9. Randall, J.E. and R. Lubbock. 1981. A revision of the serranid fishes of the subgenus Mirolabrichthys (Anthiinae: Anthias), with descriptions of five new species. Contr. Sci. 333: 1–27. Randall, J.E. and U. Satapoomin. 2000. Cephalopholis polyspila, a new species of grouper (Perciformes: Serranidae: Epinephelinae) from southwestern Thailand and Sumatra. Phuket mar. biol. Cent. Res. Bull. 63: 1–7. Family Pseudochromidae Gill, A.C. 2004. Revision of the Indo-Pacific dottyback fish subfamily Pseudochrominae (Perciformes: Pseudochromidae). Smithiana Monograph 1. 213 p, 12 pls. Winterbottom, R. and J.E. Randall. 1994. Two new species of congrogadins (Teleostei: Pseudochromidae), with range extensions for four other species. Can. J. Zool. 72: 750–756. Family Plesiopidae Mooi, R.D. 1995. Revision, phylogeny, and discussion of biology and biogeography of the fish genus Plesiops (Perciformes: Plesiopidae). Royal Ontario Museum Life Sciences Contributions No. 159. iv + 107 p. Family Opistognathidae Smith-Vaniz, W.F. 2009. Three new species of Indo-Pacific jawfishes (Opistognathus: Opistognathidae), with the posterior end of the upper jaw produced as a thin flexible lamina. Aqua, Int. J. Ichthyol. 15(2): 69–108. Family Apogonidae Gon, O. 1996. Revision of the cardinalfish subgenus Jaydia (Perciformes, Apogonidae, Apogon). Trans. Roy. Soc. S. Afr. 51: 147–194. Gon, O. 2000. The taxonomic status of the cardinalfish species Apogon niger, A. nigripinnis, A. pharaonis, A. sialis, and related species (Perciformes: Apogonidae). J.L.B. Smith Inst. Ichthyol. Spec. Publ. 65: 1–20. Gon, O. and J.E. Randall. 2003. Revision of the Indo-Pacific cardinalfish genus Archamia (Perciformes: Apogonidae), with description of a new species. Indo-Pacific Fishes 35: 1–49, 3 pls. Fraser, T.H. 2005. A review of the species in the Apogon fasciatus group with a description of a new species of cardinalfish from the Indo-west Pacific (Perciformes: Apogonidae). Zootaxa 924: 1–30. Fraser, T.H. 2008. Cardinafishes of the genus Nectamia (Apogonidae, Perciformes) from the Indo-Pacific region with descriptions of four new species. Zootaxa 1691: 1–52. 74 Phuket mar. biol. Cent. Res. Bull. Fraser, T.H. and E.A. Lachner. 1985. A revision of the cardinalfish subgenera Pristiapogon and Zoramia (Genus Apogon) of the Indo-Pacific region (Teleostei: Apogonidae). Smithson. Contr. Zool. 42. 47 p. Randall, J.E., T.H. Fraser and E.A. Lachner. 1990. On the validity of the Indo-Pacific cardinalfishes Apogon aureus (Lacepède) and A. fleurieu (Lacepède), with description of a related new species from the Red Sea. Proc. Biol. Soc. Wash. 103(1): 39–62. Family Sillaginidae McKay, R.J. 1985. A revision of the fishes of the family Sillaginidae. Mem. Qd. Mus. 22(1): 1–73. Family Leiognathidae Kimura, S., T. Ito, T. Peristiwady, Y. Iwatsuki and T. Yoshino. 2005. The Leiognathus splendens complex (Perciformes: Leiognathidae) with the description of a new species, Leiognathus kupanensis Kimura and Peristiwady. Ichthyol. Res. 52(3): 275–291. Kimura, S., R. Kimura and K. Ikejima. 2008. Revision of the genus Nuchequula with descriptions of three new species (Perciformes: Leiognathidae). Ichthyol. Res. 55(1): 22–42. Kimura, S., T. Yamashita and Y. Iwatsuki. 2000. A new species, Gazza rhombea, from the Indo-west Pacific, with a redescription of G. achlamys Jordan & Starks, 1917 (Perciformes: Leiognathidae). Ichthyol. Res. 47(1): 1–12. Mochizuki, K. and M. Hayashi. 1989. Revision of the leiognathid fishes of the genus Secutor, with two new species. Sci. Rept. Yokosuka City Mus. 37: 83–95. Yamashita, T., S. Kimura and Y. Iwatsuki. 1998. Validity of the leiognathid fish, Gazza dentex (Valenciennes in Cuvier & Valenciennes, 1835), with designation of a lectotype, and redescription of G. minuta (Bloch, 1795). Ichthyol. Res. 45(3): 271–280. Family Lutjanidae Allen, G.R. and F.H. Talbot. 1985. Review of the snappers of the genus Lutjanus (Pisces: Lutjanidae) from the IndoPacific, with description of a new species. Indo-Pacific Fishes 11: 1–87, 10 pls. Family Caesionidae Carpenter, K.E. 1987. Revision of the Indo-Pacific fish family Caesionidae (Lutjanoidea), with descriptions of five new species. Indo-Pacific Fishes 15: 1–56, 7 pls. Family Gerreidae Iwatsuki, Y. and P.C. Heemstra. 2001. Gerres phaiya: a new species of gerreid fish (Teleostei: Perciformes: Gerreidae) from India, with comments on Gerres poieti and the Gerres erythrourus complex. Copeia 2001(4): 1043– 1049. Iwatsuki, Y., S. Kimura, H. Kishimoto and T. Yoshino. 1996. Validity of the gerreid fish, Gerres macracanthus Bleeker, 1854, with designation of lectotype, and designation of a neotype for G. filamentosus Cuvier, 1829. Ichthyol. Res. 43(4): 417–429. Iwatsuki, Y., S. Kimura and T. Yoshino. 1999a. Description of Gerres chrysops sp. nov. from Thailand and redescription of Gerres setifer (Hamilton, 1822) and G. decacanthus (Bleeker, 1865) (Perciformes: Gerreidae). Ichthyol. Res. 46(1): 27–41. Iwatsuki, Y., S. Kimura and T. Yoshino. 1999b. Redescription of Gerres baconensis (Evermann & Seale, 1907), G. equulus Temminck & Schlegel, 1844 and G. oyena (Forsskål, 1775), included in the “G. oyena complex”, with notes on other related species (Perciformes: Gerreidae). Ichthyol. Res. 46(4): 377–395. Iwatsuki, Y., S. Kimura and T. Yoshino. 2001. Redescription of Gerres longirostris (Lacepède, 1801) and Gerres oblongus Cuvier in Cuvier and Valenciennes, 1830, included in the Gerres longirostris complex (Perciformes: Gerreidae). Copeia 2001(4): 954–965. Iwatsuki, Y., S. Kimura and T. Yoshino. 2007. A review of the Gerres subfasciatus complex from the Indo-West Pacific, with three new species (Perciformes: Gerreidae). Ichthyol. Res. 54(2): 168–185. Family Haemulidae Johnson, J.W., J.E. Randall and S.F. Chenoweth. 2001. Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review. Mem. Qd. Mus. 46(2): 657–676. Iwatsuki, Y. and B.C. Russell. 2006. Revision of the genus Hapalogenys (Teleostei: Perciformes) with two new species from Indo-West Pacific. Mem. Mus. Vict. 63(1): 29–46. Iwatsuki, Y., U. Satapoomin and K. Amaoka. 2000. New species: Hapalogenys merguiensis (Teleostei; Perciformes) from Andaman Sea. Copeia 2000(1): 129–139. McKay, R.J. and U. Satapoomin. 1994. Pomadasys andamanensis, a new species of haemulid fish from Thailand. Phuket mar. biol. Cent. Res. Bull. 59: 1–4. Satapoomin, U and J.E. Randall. 2000. Plectorhinchus macrospilus, a new species of sweetlip (Perciformes: Haemulidae) from the southwestern Thailand, Andaman Sea. Phuket mar. biol. Cent. Res. Bull. 63: 9–16. Family Nemipteridae 75 The fishes of southwestern Thailand Russell, B.C. 1986. Review of the western Indian Ocean species of Nemipterus Swainson 1839, with description of a new species. Senckenberg. biol. 67(1/3): 19–35. Russell, B.C. 1991. On the validity of Nemipterus furcosus (Valenciennes) (Nemipteridae). Cybium 15(1): 35–41. Russell, B.C. and D. Golani. 1993. A review of the fish genus Parascolopsis (Nemipteridae) of the western Indian Ocean, with description of a new species from the northern Red Sea. Israel J. Zool. 39: 337–347. Family Polynemidae Feltes, R.M. 1991. Revision of the polynemid fish genus Filimanus, with description of two new species. Copeia 1991(2): 302–322. Motomura, H. 2002. Revision of the Indo-Pacific threadfin genus Polydactylus (Perciformes, Polynemidae) with a key to the species. Bull. Natl. Sci. Mus., Ser. A. 28(3): 171–194. Motomura, H., Y. Iwatsuki, S. Kimura and T. Yoshino. 2001. A new species, Polydactylus siamensis, from Thailand and redescription of P. plebeius (Broussonet, 1782) with designation of a neotype (Perciformes: Polynemidae). Ichthyol. Res. 48(2): 117–126. Motomura, H., Y. Iwatsuki and T. Yoshino. 2002. Revision of the Indo-Pacific polynemid fish genus Eleutheronema (Teleostei: Perciformes). Ichthyol. Res. 49(1): 47–61. Motomura, H., U. Satapoomin and Y. Iwatsuki. 2000. A new record of the threadfin, Filimanus perplexa Feltes, 1991 (Perciformes: Polynemidae), from the Andaman Sea, Thailand. Phuket mar. biol. Cent. Res. Bull. 63: 17–20. Family Sciaenidae Sasaki, K. 1992. Two new and two resurrected species of sciaenid genus Johnius (Johnius) from the west Pacific. Jap. J. Ichthyol. 39(3): 191–199. Sasaki, K. 1996. Sciaenid fishes of the Indian Ocean (Teleostei, Perciformes). Mem. Fac. Sci. Kochi Univ. Ser. D (Biol.) 16/17: 83–95. Family Mullidae Randall, J.E. 2004. Revision of the goatfish genus Parupeneus (Perciformes: Mullidae), with descriptions of two new species. Indo-Pacific Fishes 36: 1–64, 16 pls. Family Kuhliidae Randall, J.E. and H.A. Randall. 2001. Review of the fishes of the genus Kuhlia (Perciformes: Kuhliidae) of the central Pacific. Pac. Sci. 55(3): 227–256. Family Pomacentridae Allen, G.R. 1991. Damselfishes of the World. Mergus Publishers, Melle, Germany. 271 p. Allen, G.R. and A.R. Emery. 1985. A review of the pomacentrid fishes of the genus Stegastes from the Indo-Pacific, with descriptions of two new species. Indo-Pacific Fishes 3: 1–31, 3 pls. Allen, G.R. and M.V. Erdmann. 2005. Chromis xouthos, a new species of damselfish (Pomacentridae) from the east Andaman Sea and central Indian Ocean. Aqua, J. Ichthyol. Aquat. Biol. 10(3): 89–94. Allen, G.R. and J.E. Randall. 2002. A review of the leucogaster species complex of the Indo-Pacific pomacentrid genus Amblyglyphidodon, with descriptions of two new species. Aqua, J. Ichthyol. Aquat. Biol. 5(4): 139–152. Randall, J.E. and G.R. Allen. 2005. Neopomacentrus sororius, a new species of damselfish from the Indian Ocean, with description of a neotype for its sister species, N. azysron (Bleeker). Aqua, J. Ichthyol. Aquat. Biol. 10(2): 73– 80. Family Labridae Gomon, M.F. 2006. A revision of the labrid fish genus Bodianus with descriptions of eight new species. Rec. Aust. Mus. Suppl. 30: 1–133. Randall, J.E. 1996. Second revision of the labrid fish genus Leptojulis, with descriptions of two new species. IndoPacific Fishes. 24: 1–20, 3 pls. Randall, J.E. 1999. Revision of the Indo-Pacific labrid fishes of the genus Pseudocheilinus, with descriptions of three new species. Indo-Pacific Fishes. 28: 1–34, 2 pls. Randall, J.E. 1999. Revision of the Indo-Pacific labrid fishes of the genus Coris, with descriptions of five new species. Indo-Pacific Fishes. 29: 1–74, 22 pls. Randall, J.E. 2000. Revision of the Indo-Pacific labrid fishes of the genus Stethojulis, with descriptions of two new species. Indo-Pacific Fishes. 31: 1–42, 6 pls. Randall, J.E. and Lubbock, R. 1981. Labrid fishes of the genus Paracheilinus, with descriptions of three new species from the Philippines. Jap. J. Ichthyol. 28(1): 19–30, 2 pls. Randall, J.E. and M.M. Smith. 1982. A review of the labrid fishes of the genus Halichoeres of the western Indian Ocean, with descriptions of six new species. Ichthyol. Bull. J.L.B. Smith Inst. Ichthyol. 45: 1–26, 6 pls. Family Scaridae Satapoomin, U., R.H. Kuiter and J.E. Randall. 1994. First Record of the parrotfish Scarus viridifucatus from Thailand (the Andaman Sea) and Indonesia. Phuket mar. biol. Cent. Res. Bull. 59: 5–9. 76 Phuket mar. biol. Cent. Res. Bull. Westneat, M., U. Satapoomin and J.E. Randall. 2007. Scarus maculipinna, a new species of parrotfish (Perciformes, Scaridae) from eastern Indian Ocean. Zootaxa. 1628: 59–68. Family Ammodytidae Ida, H., P. Sirimontaporn and S. Monkolprasit. 1994. Comparative morphology of the fishes of the family Ammodytidae, with description of two new genera and two new species. Zool. Stud. 33(4): 251–277. Family Tripterygiidae Holleman, W. 2005. A review of the triple fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the western Indain Ocean, with descriptions of four new species. Smithiana Bulletin. 5: 1–25, 2 pls. Holleman, W. 2006. Fishes of the Helcogramma steinitzi species group (Blennioidei: Tripterygiidae) from the Indain Ocean, with descriptions of two new species. Aqua, J. Ichthyol. Aquat. Biol. 11(3): 89–104. Williams, J.T. and J.C. Howe. 2003. Seven new species of the triplefin genus Helcogramma (Tripterygiidae) from the Indo-Pacific. Aqua, J. Ichthyol. Aquat. Biol. 7(4): 151–176. Family Blenniidae Bath, H. 1992. Revision der gattung Praealticus Schultz & Chapman 1960 (Pisces: Blenniidae). Senckenberg. biol. 72(4/ 6): 237–316. Carlson, B.A. 1980. A new Indo-Pacific fish of the genus Cirripectes (Blenniidae, Salariini). Pac. Sci. 34(4): 407–414. Smith-Vaniz, W.F. 1976. The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae). The Academy of Natural Sciences of Philadelphia. Monograph 19. vii + 196 p. Smith-Vaniz, W.F., U. Satapoomin and G.R. Allen. 2001. Meiacanthus urostigma, a new fangblenny from the northeastern Indian Ocean, with discussion and examples of mimicry in species of Meiacanthus (Teleostei: Blenniidae: Nemophini). Aqua, J. Ichthyol. Aquat. Biol. 5(1): 25–43. Springer, V.G. 1988. The Indo-Pacific blenniid fish genus Ecsenius. Smithson. Contr. Zool. 465. 134 p., 14 pls. Springer, V.G. and M.F. Gomon. 1975. Revision of the blenniid fish genus Omobranchus with descriptions of three new species and notes on other species of the tribe Omobranchini. Smithson. Contr. Zool. 177. 135 p. Springer, V.G. and J.E. Randall. 1999. Ecsenius polystictus, new species of blenniid fish from Mentawai Islands, Indonesia, with notes on other species of Ecsenius. Revue fr. Aquariol. 26(1–2): 39–48. Springer, V.G. and J.T. Williams. 1994. The Indo-west Pacific blenniid fish genus Istiblennius reappraised: A revision of Istiblennius, Blenniella, and Paralticus, new genus. Smithson. Contr. Zool. 565. 193 p. Williams, J.T. 1988. Revision and phylogenetic relationships of the blenniid fish genus Cirripectes. Indo-Pacific Fishes. 17: 1–78, 7 pls. Family Callionymidae Fricke, R. 1983. Revision of the Indo-Pacific genera and species of the dragonet family Callionymidae. Theses Zoologicae Vol. 3. 774 p. Family Gobiidae Akihito and Meguro, K. 2000. Review of the Gobiid genus Cristatogobius found in Japan with description of a new species. Ichthyol. Res. 47(3): 249–261. Darumas, U. and P. Tantichodok. 2002. A new species of mudskipper (Gobiidae: Oxucercinae) from southern Thailand. Phuket mar. biol. Cent. Res. Bull. 64: 101–107. Hoese, D.F. and H.K. Larson. 1994. Revision of the Indo-Pacific gobiid fish genus Valenciennea, with descriptions of seven new species. Indo-Pacific Fishes. 23: 1–71, 6 pls. Lachner, E.A. and S.J. Karnella. 1980. Fishes of the Indo-Pacific fish genus Eviota with descriptions of eight new species (Teleostei: Gobiidae). Smithson. Contr. Zool. 315. 127 p. Lachner, E.A. and J.F. McKinney. 1978. A revision of the Indo-Pacific fish genus Gobiopsis with descriptions of four new species (Pisces: Gobiidae). Smithson. Contr. Zool. 262. 52 p. Larson, H.K. 1999. A review of the mangrove goby genus Hemigobius (Gobioidei, Gobiidae, Gobionellinae). The Beagle, Records of the Museums and Art Galleries of the Northern Territory. 15: 23–42. Larson, H.K. 1999. Allocation to Calamiana and redescription of the fish species Apocryptes variegates and Vaimosa mindora (Gobioidei: Gobiidae: Gobionellinae), with description of a new species. Raffles Bull. Zool. 47(1): 257–281. Larson, H.K. 2001. A revision of the gobiid fish genus Mugilogobius (Teleostei: Gobioidei). Rec. West. Aust. Mus. Suppl. No. 62: 1–233. Larson, H.K. 2005. A revision of the gobiid genus Stigmatogobius (Teleostei: Gobiidae), with descriptions of two new species. Ichthyol. Explor. Freshwaters. 16(4): 347–370. Larson, H.K. 2009. Review of the gobiid fish genera Eugnathogobius and Pseudogobiopsis (Gobioidei: Gobiidae: Gobionellinae), with descriptions of three new species. Raffles Bull. Zool. 57(1): 127–181. Murdy, E.O. 1985. A review of the gobiid fish genera Exyrias and Macrodontogobius, with description of a new species of Exyrias. Indo-Pacific Fishes. 10: 1–14, 2 pls. 77 The fishes of southwestern Thailand Murdy, E.O. 1989. A taxonomic revision and cladistic analysis of the oxudercine gobies (Gobiidae: Oxudercinae). Rec. Aust. Mus. Suppl. 11: 1–93. Murdy, E.O. and D.F. Hoese. 1985. Revision of the gobiid fish genus Istigobius. Indo-Pacific Fishes 4: 1–41, 3 pls. Polunin, N.V.C. and R. Lubbock. 1979. Five new prawn-associated gobies (Teleostei: Gobiidae) of the genus Amblyeleotris. Bull. Br. Mus. Nat. Hist. (Zool.). 36(4): 239–249. Randall, J.E. 2001. Five new Indo-Pacific gobiid fishes of the genus Coryphopterus. Zool. Stud. 40(3): 206–225. Randall, J.E. and D.W. Greenfield. 2001. A preliminary review of the Indo-Pacific gobiid fishes of the genus Gnatholepis. Ichthyol. Bull. J.L.B. Smith Inst. Ichthyol. 69: 1–17, 2 pls. Randall, J.E. and D.F. Hoese. 1985. Revision of the Indo-Pacific dartfishes, genus Ptereleotris (Perciformes: Goioidei) Indo-Pacific Fishes. 7: 1–36, 4 pls. Shibukawa, K. and G.R. Allen. 2007. Review of the cheek-spine goby genus Gladiogobius (Actinopterigii, Perciformes, Gobiidae), with descriptions of two new species from the Indo-Pacific. Bull. Natl. Mus. Nat. Sci., Ser. A, 33(4): 193–206. Shibukawa, K. and U. Satapoomin. 2006. Myersina adonis, a new species of shrimp-associated goby (Pisces: Perciformes: Gobiidae) from the Andaman Sea. Bull. Natl. Sci. Mus., Ser. A, 32(1): 29–37. Shibukawa, K., T. Suzuki, H. Senou and K. Yano. 2005. Records of three shrimp-goby species (Teleostei, Perciformes, Gobiidae) from the Ryukyu Archipelago, Japan. Bull. Natl. Sci. Mus., Ser. A, 31(4): 191–204. Satapoomin, U. and R. Winterbottom. 2002. Redescription of the gobioid fish Cryptocentrus pavoninoides (Bleeker, 1849), with notes on sexual dichromatism in shrimp gobies. Aqua, J. Ichthyol. Aquat. Biol. 5(2): 53–64. Winterbottom, R. 2002. A redescription of Cryptocentrus crocatus Wongratana, a redefinition of Myersina Herre (Acanthopterygii; Gobiidae), a key to the species, and comments on relationships. Ichthyol. Res. 49(1): 69– 75. Winterbottom, R. and M. Burridge. 1993. Revision of the species of Priolepis possessing a reduced transverse pattern of cheek papillae and no predorsal scales (Teleostei: Gobiidae). Can. J. Zool. 71: 494–514. Winterbottom, R. and A.S. Harold. 2005. Gobiodon prolixus, a new species of gobiid fish (Teleostei: Perciformes: Gobiidae) from the Indo-west Pacific. Proc. Biol. Soc. Wash. 118(3): 582–589. Winterbottom, R. and L. Southcott. 2007. Two new species of the genus Trimma (Percomorpha: Gobiidae) from western Thailand. Aqua, Int. J. Ichthyol. 13(2): 69–76. Family Siganidae Woodland, D.J. 1990. Revision of the fish family Siganidae with descriptions of two new species and comments on distribution and biology. Indo-Pacific Fishes. 19: 1–136, 11 pls. Family Acanthuridae Randall, J.E. 1993. Acanthurus tristis, a valid Indian Ocean surgeonfish (Perciformes: Acanthuridae). J.L.B. Smith Inst. Ichthyol. Spec. Publ. 54: 1–8. Randall, J.E. and K.D. Clements. 2001. Second revision of the surgeonfish genus Ctenochaetus (Perciformes: Acanthuridae), with descriptions of two new species. Indo-Pacific Fishes. 32: 1–33, 6 pls. Family Sphyraenidae Doiuchi, R. and T. Nakabo. 2005. The Sphyraena obtusata group (Perciformes: Sphyraenidae) with a description of a new species from southern Japan. Ichthyol. Res. 52(2): 132–151 Order Pleuronectiformes Menon, A.G.K. 1977. A systematic monograph of the tongue soles of the genus Cynoglossus Hamilton-Buchanan (Pisces: Cynoglossidae). Smithson. Contr. Zool. 238. 129 p. Randall, J.E. and M. Desoutter-Meniger. 2007. Review of the soles of the genus Aseraggodes (Pleuronectiformes: Soleidae) from the Indo-Malayan region, with descriptions of nine new species. Cybium. 31(3): 301–331. Randall, J.E. and J.W. Johnson. 2007. Revision of the soleid fish genus Pardachirus. Indo-Pacific Fishes. 39: 1–22, 4 pls. Order Tetraodontiformes Hutchins, J.B. 1997. Review of the monacanthid fish genus Paramonacanthus, with descriptions of three new species. Rec. West. Aust. Mus. Suppl. No. 54. 57 p. Leis, J.M. 2006. Nomenclature and distribution of the species of the porcupinefish family Diodontidae. Mem. Mus. Vict. 63(1): 77–90. Randall, J.E. and W. Klausewitz. 1973. A review of the triggerfish genus Melichthys, with description of a new species from the Indian Ocean. Senckenberg. biol. 54(1/3): 57–69. 78 Phuket mar. biol. Cent. Res. Bull.