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SNAME  Harpalus griseus (Panzer, 1797)
JNAME  keusugomokumushi
Collection ground Matsumae cho, Hokkaido Pref. 1988/9/12
Distribution Hokkaido, Honshu, Shikoku, Kyushu
Ecology flatland (Ishitani 1996), They prefer relatively stable environment in artificial disturbed paddy levee (Lee et al. 2008), cultivated field (Habu and Sadanaga 1965), paddy levee (Inoue 1974c, Inoue 1975a, Inoue 1976a, Inoue 1977a, Inoue 1977b), firm field (Inoue 1979a), grassland, firm field, paddy levee in flatland (Inoue 1977e), crop field (Inoue and Hanada 1979, Inoue 1980a), bank (Togashi 1986, Aono 1988, Aono 1987), apple orchard (Koyama and Yamada 1980), riverbed (Terada 1983, Lee and Ishii 2009, 2010), slope of mound (Sunose 1992, Sunose and Kurosawa 1992), vineyard (Togashi and Ohbata 1995), coppice forest (Hosoda et al. 1996), secondary forest of Alnus sieboldiana (Ishitani 1996), secondary forest of Pinus densiflora (Ishitani 1996), rape field (Ishitani 1996), riverbed (Ishii et al. 1996, Yamazaki et al. 1999), mountain (Hosoda 2000), forest (Hosoda 2000, Siddiquee and Nakamura 2004, Suttiprapan and Nakamura 2007), vegetable field (Siddiquee and Nakamura 2004), field (Salah et al. 2004), paddy field (Yahiro 2007, Kagawa 2008, Lee et al. 2008), urban park (Lee and Ishii 2009), paddy field (Lee and Ishii 2009), Satoyama forest (Lee and Ishii 2009), adult (Suda 1980, Yamazaki et al. 1999), larvae (Inoue 1980a), The eggs were laid during about the middle of September and that of October in our breeding room (Habu 1973d), autumn (Yamazaki et al. 1999), starawberry (Kirchner 1938), the larvae of Callosobruchus chinensis and Listoderes obliquus (Pospielov, 1913), grubs of some curculionid and scarabaeid beetles (larva) (Blunck 1925), collected by light trapping (Inoue 1974c, Inoue 1975a, Inoue 1976a, Inoue 1977a, Inoue 1977b, Inoue and Hanada 1979, Aono 1988, Suttiprapan and Nakamura 2007, Yahiro 2007)
Types 　 　 　
Remarks Carabus griseus Panzer, 1797, Faun. Germ., 38, no. 1 (Europe: Germany) (Habu 1973d) , Carabaus griseus Panzer, Carabus bicolor Marsham, Harpalus ruficornis var. ? griseus Panzer, Harpalus griseus Panzer, Harpalus (Harpalus) Griseus Kugelann, Psedoophonus griseus Panzer, Harpalus (Pseudoaphonus [!]) griseus Panzer, Harpalus Reichei Desbrochers, Psedophnus griseus Panzer, Ophonus (Pseudophonus) griseus Panzer, Harpalus (Pseudophonus) griseus Panzer, Ophonus griseus Panzer, Ophonus (Pseudoophonus) griseus Panzer, Harpalus (Pseudoophonus) griseus Panzer (Habu 1973d)